323,540 research outputs found

    Dadagulella pembensis Rowson & Tattersfield 2013, sp. nov.

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    Dadagulella pembensis sp. nov. urn:lsid:zoobank.org:act: 4442210F-A097-400D-B34F-96 A 4 B 7677CE7 Figs 32, 46, 60, 66-68, 70, 75, 79-80, 84; Table 1 ““ Gulella ” radius (Preston, 1910) ” – Rowson et al. 2010 b: 9-10, 28-29, figs 60-61. Etymology From Pemba Island, with the final ‘a’ elided for euphony. Type material examined TANZANIA: holotype NMW.Z.2009.013.00227: 1 ad., Ngezi Forest Reserve (4.94°S, 39.69°E), Pemba I., Zanzibar, dry forest on dark, sandy soil on coral rag on Tondooni peninsula within reserve, leg. BR, B. H. Warren & CFN, 7 Feb. 2009; fig. 60 in Rowson et al. (2010 b). Paratypes NMW. Z.2009.013.00228: 2 ads, data as holotype. Paratypes MRAC. MT.803796-7: 2 ads, data as holotype. Paratypes NHMUK.20120261: 2 ads, data as holotype. Paratypes NMK: 2 ads, data as holotype. Paratypes NMT: 2 ads, data as holotype. Paratypes NMSA. L 8694/ T 3063: 2 ads, data as holotype. Paratypes RMNH. MOL.288090: 2 ads, data as holotype. Other material examined TANZANIA: NMW. Z.2009.013.00229: 15 juvs, data as holotype; one figured as fig. 61 in Rowson et al. (2010 b). NMW. Z.2009.013.00230: 3 ads, near Chwaka (5.39° S, 39.77° E), Pemba I., Zanzibar, clove and fruit tree woodland on dark, sandy soil, leg. BR, B. H. Warren & CFN, 15 Feb. 2009. NMW. Z.2009.013.00231: 3 ads, Msitu Mkuu Forest Reserve (5.00° S, 39.83° E), Pemba I., Zanzibar, moist forest on dark (not sandy) soil on coral rag in high forest in north of reserve, leg. BR, B. H. Warren & CFN, 10 Feb. 2009. NMW. Z.2009.013.00232: 3 ads, Ngezi Forest Reserve, Pemba I., Zanzibar, dry forest and thicket on dark, sandy soil on coral rag on coast of Tondooni peninsula within reserve, leg. BR, B. H. Warren & CFN, 8 Feb. 2009. NMW. Z.2009.013: Many additional ads and juvs from the type locality and other sites on Pemba I., Feb. 2009; see table 2 of Rowson et al. (2010 b). Description SHELL (Figs 32, 46, 60). Large (4.80 - 5.50 mm high x 2.00 - 2.50 mm wide), of 6.5 - 7.5 whorls. Ovateacuminate, spire narrowly to broadly acuminate (spire angle 51 - 65°). Apex sharply pointed. Embryonic whorls smoothly granulate. Later whorls with relatively coarse ribs (12 - 15 per mm on penultimate whorl). Sutures shallow, shell appearing relatively straight-sided. Umbilicus closed or nearly so. Peristome complete, or incomplete parietally. Outer palatal surface of aperture with a depression corresponding to the palatal tooth and often another corresponding to the basal tooth. Dentition 6-fold, consisting of: one lamella-like parietal tooth; one slab-like palatal tooth, forming a parieto-palatal sinus; one basal denticle; a deep-set columellar baffle, sometimes folded, sub-bifid or sub-trifid; and two shallower, broad columellar denticles. Juvenile shells (Fig. 46) with 3-fold dentition, similar to some forms of D. r. radius comb. nov.: one parietal lamella; one basal tooth; and one columellar thickening. Earlier basal teeth are retained in juveniles. CEPHALOPODIUM. Pale yellow, with apricot to orange tentacle retractors. SALIVARY GLANDS (Fig. 70). United, soft, not tumid, elongate, Y-shaped; each duct leaving at the apex of the lobe and evenly thick throughout. RADULA (Figs 66-68). With a unicuspid central tooth and around 15 laterals in each half-row, gradually diminishing in size laterally. All laterals bicuspid, tricuspid, or quadricuspid, with outer cusps much smaller than inner cusps. Teeth delicate, short and flake-like at the ventral end of the radular ribbon. GENITALIA (Figs 75, 79, 80). Vas deferens appearing thickened prior to insertion on penis, but actually with a short, broad parallel diverticulum. Penial sheath absent but with a thin sheath-like layer contiguous with wall of lower penis. Interior of penis with weak radial pilasters and small rhombic pads. Apical, muscular part of penis with a single large hook, associated with a spatulate “scoop” with microscopically serrated tip. Elsewhere in penis, one or two longitudinal rows of short, simple hooks mounted on rhombic pads. “Spermatophore” (see Discussion) present in penis of two Ngezi specimens; partially digested remains in bursa of another. Range and habitat Pemba Island, Tanzania, where widespread in forest and other vegetated habitats. Remarks This species was discussed by Rowson et al. (2010 b), who suggested a thorough revision of the group to establish whether the Pemba populations were part of a variable D. radius comb. nov. or a subspecies or species endemic to Pemba. This uncertainty was incorporated into their conclusions on endemism on the island (Rowson et al. 2010 b: 28-29, 32). We find here that there is little or no overlap with D. radius comb. nov. s.l. or other species, so conclude that this is an island endemic, which we here name D. pembensis sp. nov. Like specimens of D. r. radius comb. nov. on the opposite mainland at Amboni (e.g. Figs 15, 53), D. pembensis sp. nov. is large and has a complex pattern of 6-fold dentition. However it is distinguishable from them by its even larger size, shallower sutures, and having on average an extra 0.5 whorls. It differs from D. minuscula mahorana subsp. nov. which has similar dentition, in its larger size, shallower sutures, and stronger and more widely spaced ribs.Published as part of Rowson, Ben & Tattersfield, Peter, 2013, Revision of Dadagulella gen. nov., the " Gulella radius group " (Gastropoda: Streptaxidae) of the eastern Afrotropics, including six new species and three new subspecies, pp. 1-46 in European Journal of Taxonomy 37 on pages 20-22, DOI: 10.5852/ejt.2013.37, http://zenodo.org/record/380677

    Dadagulella rondoensis Rowson & Tattersfield 2013, comb. nov.

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    Dadagulella rondoensis (Verdcourt, 1994) comb. nov. Figs 41, 84; Table 1 Gulella rondoensis Verdcourt, 1994: 137-139, fig. 1. Gulella rondoensis – Verdcourt 2006: 48. Type material examined TANZANIA: holotype SMF.310150: 1 ad., Rondo Forest Reserve, Rondo Plateau, Lindi District (10.12°S, 39.22°E), evergreen forest with Milicia Sim, Albizia Durazz., Dialium L., etc. in small gully and amphitheatre around a well on escarpment, 650 m alt., leg. Bidgood, Abdallah & Vollesen, 10 Feb. 1991 (examined digital photograph only). Other material examined None. Description SHELL (Fig. 41). Large (4.10 mm high x 2.20 mm wide), of 6.25 whorls. Subconical (maximum width being in bottom third of the shell, at body whorl). Spire narrowly acuminate, almost cyrtoconoid (convex) rather than coeloconoid (spire angle 52°). Apex sharply pointed. Embryonic whorls punctate or malleate, rather than merely granulate. Later whorls with relatively fine ribs (about 14 per mm on penultimate whorl). Sutures of intermediate depth. Umbilicus narrowly open. Peristome incomplete parietally. Dentition weak, 3-fold (although could be interpreted as 2-fold), consisting of: one lamellalike parietal tooth, with a swelling above it that recalls the one in D. conoidea; one weak palatal tooth, not forming a parieto-palatal sinus; and one very weak, shallow columellar swelling. Further minute swellings just perceptible in the holotype (Fig. 41) were not noted by Verdcourt (1994) who interpreted the dentition as 2-fold. Shells and anatomy of juveniles unknown. Range and habitat Forest at the type locality, southeastern Tanzania. Remarks This species differs from D. conoidea comb. nov. in its weaker dentition, smaller size, and in having punctate apical whorls. Along with D. conoidea comb. nov. and D. delta sp. nov. it has weaker dentition than other Dadagulella gen. nov. However, both D. rondoensis comb. nov. and D. conoidea comb. nov. are distinctively more conical, i.e. less ovate than other Dadagulella gen. nov. These two are attributed to the genus somewhat speculatively, on the basis of their acuminate spire and pointed apex. These features mean they do not obviously fit into Gulella or any of its named subgenera, or indeed other plausible streptaxid genera. No anatomical or juvenile shell data are available for either species, both being known from single specimens. The punctuate apex of D. rondoensis comb. nov., unique in Dadagulella gen. nov., may not be significant in this respect (see Rowson 2007b for a discussion on the value of apical sculpture in distinguishing African streptaxid genera). The apex of D. conoidea comb. nov. is less obviously punctuate, although it may have been worn smooth (Verdcourt 1996). In the description of D. rondoensis comb. nov., Verdcourt (1994) discussed a resemblance only to Gulella galactochila (Crosse, 1885), a much larger and more broadly acuminate Tanzanian species that we consider to lack the characteristic features of Dadagulella gen. nov. Although G. galactochila has not been dissected, the anatomy of another species very similar to it (G. udzungwensis van Bruggen, 2003) lacks the anatomical features of Dadagulella gen. nov. (Rowson unpublished). In his discussion of D. conoidea comb. nov., Verdcourt (1996) made no reference to either D. rondoensis comb. nov. or G. galactochila, but only to two species that we here treat in Dadagulella gen. nov. (D. r. radius comb. nov. and D. cuspidata comb. nov.). We contend firstly that D. rondoensis comb. nov. and D. conoidea comb. nov. are more similar to one another than either is to G. galactochila, and secondly that the resemblance between D. conoidea comb. nov., D. r. radius comb. nov. and D. cuspidata comb. nov. extends also to D. rondoensis comb. nov.. Our attribution of them to Dadagulella gen. nov. reflects this point of view.Published as part of Rowson, Ben & Tattersfield, Peter, 2013, Revision of Dadagulella gen. nov., the " Gulella radius group " (Gastropoda: Streptaxidae) of the eastern Afrotropics, including six new species and three new subspecies, pp. 1-46 in European Journal of Taxonomy 37 on pages 30-31, DOI: 10.5852/ejt.2013.37, http://zenodo.org/record/380677

    Dadagulella meredithae Rowson & Tattersfield 2013, comb. nov.

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    Dadagulella meredithae (van Bruggen, 2000) comb. nov. Figs 37, 84; Table 1 Gulella meredithae van Bruggen, 2000: 226-232, figs 1-7. “…an as yet unidentified species from Malawi ” – van Bruggen & Meredith 1984: 165. Gulella meredithae – Rowson & Lange 2007: 31. Type material examined MALAWI: holotype RMNH.59399: 1 ad., Nyika National Park (approx. 10.6°S, 33.8°E), Rumphi District, Juniper Forest, approx. 2100 m alt., leg. H.M. Meredith, 16 Sep. 1983. Other material examined None. Description SHELL (Fig. 37). Small (2.30 - 3.10 mm high x 1.40 - 1.60 mm wide), of 5.5 - “<6” whorls. Ovate-acuminate, although spire (spire angle 58 - 77°) less acuminate than in other Dadagulella gen. nov species. Apex rounded (in holotype) to weakly pointed. Embryonic whorls smoothly granulate. Later whorls with very fine, very numerous ribs (about 27 per mm on penultimate whorl). Sutures of intermediate depth. Umbilicus closed or nearly so. Peristome incomplete parietally. Outer palatal surface of aperture with a depression corresponding to the palatal tooth. Dentition 3-fold to 4-fold, consisting of at least: one lamella-like parietal tooth; one slab-like palatal tooth, not forming a parieto-palatal sinus; and a mammillate columellar baffle. Additional teeth limited to a shallow, weak columellar swelling. Juvenile shells with 2-fold to 3-fold dentition: one parietal tooth; one columellar tooth; and usually one basal tooth (termed labral by van Bruggen 2000). Van Bruggen (2000) showed that earlier sets of dentition are visible through the shells of some transparent juveniles, even in the preceding whorls, suggesting slow or no resorbtion. Anatomy unknown. Range and habitat Montane forest (above 1500 m to around 2450 m) in northern and central Malawi, and adjacent part of Zambia (Chowo Forest). Van Bruggen (2000) suspected it to range into parts of Tanzania adjoining northern Malawi. Remarks This species is distinctive in its small size, very fine, numerous ribs, and dentition. It differs from D. radius comb. nov. s.l. and D. browni comb. nov. s.l. in the lack of a basal tooth or denticle. The apex is rounded in the holotype but more conical in paratypes figured by van Bruggen (2000).Published as part of Rowson, Ben & Tattersfield, Peter, 2013, Revision of Dadagulella gen. nov., the " Gulella radius group " (Gastropoda: Streptaxidae) of the eastern Afrotropics, including six new species and three new subspecies, pp. 1-46 in European Journal of Taxonomy 37 on pages 26-27, DOI: 10.5852/ejt.2013.37, http://zenodo.org/record/380677

    Dadagulella nictitans Rowson & Tattersfield 2013, comb. nov.

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    &lt;p&gt; &lt;i&gt;Dadagulella nictitans&lt;/i&gt; (Rowson &amp; Lange, 2007) comb. nov.&lt;/p&gt; &lt;p&gt;Figs 33, 84; Table 1&lt;/p&gt; &lt;p&gt; &lt;i&gt;Gulella nictitans&lt;/i&gt; Rowson &amp; Lange, 2007: 27-31, figs 4-7, 14-19.&lt;/p&gt; Type material examined &lt;p&gt;KENYA: holotype NMK: 1 ad., Taita Hills, Macha Forest Reserve (3.45&deg;S, 38.37&deg;E), leaf litter in indigenous forest (site IB), 1550 m alt., leg. C.N. Lange, 27 Nov. 1998. Paratypes NMW. Z.2007.024.00007-00009: 3 ads, data as holotype. Paratype NMW. Z.2007.024.00010: 1 juv., data as holotype.&lt;/p&gt; Other material examined &lt;p&gt;None.&lt;/p&gt; Description &lt;p&gt; SHELL (Fig. 33). Medium-sized (3.60 &lt;i&gt;-&lt;/i&gt; 3.90 mm high x 2.00 &lt;i&gt;-&lt;/i&gt; 2.20 mm wide), of 5.5 whorls. Ovateacuminate, although spire (spire angle 64 &lt;i&gt;-&lt;/i&gt; 69&deg;) less acuminate than in other &lt;i&gt;Dadagulella&lt;/i&gt; gen. nov. Apex rounded. Embryonic whorls smoothly granulate. Later whorls with relatively coarse ribs (13 &lt;i&gt;-&lt;/i&gt; 15 per mm on penultimate whorl). Sutures of intermediate depth. Umbilicus closed or nearly so. Peristome complete. Outer palatal surface of aperture with a long, furrow-like depression corresponding to the palatal tooth. Dentition 5-fold or 6-fold, consisting of: one V-shaped parietal tooth; one slab-like palatal tooth, forming a conspicuous and narrow parieto-palatal sinus; and two to three shallow columellar teeth, the lower the largest. A deep set-columellar baffle is partly or completely hidden by the constricted aperture, while a basal denticle is completely hidden behind the palatal tooth. However, the baffle and basal denticle are probably present in all specimens. One broken juvenile shell is known; it appears to lack teeth.&lt;/p&gt; &lt;p&gt;CEPHALOPODIUM. Pale cream, with pale tentacle retractors.&lt;/p&gt; &lt;p&gt;SALIVARY GLANDS. United, soft, not tumid, elongate, Y-shaped; each duct leaving at the apex of the lobe and evenly thick throughout.&lt;/p&gt; &lt;p&gt;RADULA. Not successfully prepared.&lt;/p&gt; &lt;p&gt;GENITALIA. Figured in Rowson &amp; Lange (2007).Vas deferens thickened prior to insertion on penis but apparently without diverticulum. Penial sheath absent. Interior of penis with weak radial pilasters, a single longitudinal pilaster, and small rhombic pads. Apical, muscular part of penis with two large hooks, associated with a spatulate &ldquo;scoop&rdquo; with microscopically serrated tip. Elsewhere in penis a single longitudinal row of short, simple hooks mounted on rhombic pads.&lt;/p&gt; Range and habitat &lt;p&gt;Montane forest (1400 - 1900 m) at the type locality and three other Forest Reserves in the Taita Hills, southeastern Kenya (Rowson &amp; Lange 2007).&lt;/p&gt; Remarks &lt;p&gt; This species is included in &lt;i&gt;Dadagulella&lt;/i&gt; gen. nov. on the basis of the apical penial scoop and hooks, and the Y-shaped salivary gland. Its shell features are consistent with inclusion in &lt;i&gt;Dadagulella&lt;/i&gt; gen. nov. although the apex is more rounded than in other species. A slightly or strongly rounded apex occurs in &lt;i&gt;D. selene&lt;/i&gt; comb. nov. and &lt;i&gt;D. meredithae&lt;/i&gt; comb. nov. The parieto-palatal sinus, hidden basal denticle, and columellar dentition of &lt;i&gt;D. nictitans&lt;/i&gt; comb. nov. allow it to be separated from other species.&lt;/p&gt;Published as part of &lt;i&gt;Rowson, Ben &amp; Tattersfield, Peter, 2013, Revision of Dadagulella gen. nov., the " Gulella radius group " (Gastropoda: Streptaxidae) of the eastern Afrotropics, including six new species and three new subspecies, pp. 1-46 in European Journal of Taxonomy 37&lt;/i&gt; on pages 22-23, DOI: 10.5852/ejt.2013.37, &lt;a href="http://zenodo.org/record/3806770"&gt;http://zenodo.org/record/3806770&lt;/a&gt

    La lettre, l’esprit et l’image : Tristram Shandy de Martin Rowson entre texte(s), contexte(s) et hors-texte(s)

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    Martin Rowson’s 1996 comic book version of Tristram Shandy attempts a visual rendering of Sterne’s novel while retaining its «Shandean spirit». Along what lines is the notion of «spirit» to be defined ? In French terminology, «hors-texte» encompasses not only images as a means to reach beyond the text, but also the tenets of the epistemological and aesthetic context(s) which provide both Sterne and Rowson with backdrops for their narratives. This article argues that there can be no «hors-texte» without fully apprehending the context(s).La bande dessinée de Martin Rowson (1996) relève le défi de la représentation visuelle de Tristram Shandy. Bien plus que la lettre du roman, c’est son esprit qui transparaît dans les planches du caricaturiste. Cet article vise à cerner ce qui constitue cet esprit, et comment Sterne l’auteur se trouve lié à Rowson le graphiste. La connaissance des contextes épistémologiques et esthétiques permet à Rowson de jeter des ponts entre passé et présent, suggérant qu’il ne peut y avoir de hors-texte(s) sans contexte(s).Friant-Kessler Brigitte, Lasne Sandra. La lettre, l’esprit et l’image : Tristram Shandy de Martin Rowson entre texte(s), contexte(s) et hors-texte(s) . In: XVII-XVIII. Revue de la société d'études anglo-américaines des XVIIe et XVIIIe siècles. N°62, 2006. Âges de la vie et rites de passage. pp. 241-260

    Dadagulella cuspidata Rowson & Tattersfield 2013, comb. nov.

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    Dadagulella cuspidata (Verdcourt, 1962) comb. nov. Figs 31, 59, 84; Table 1 Gulella cuspidata Verdcourt, 1962: 3, 27-28, pl. 3, fig. 2. “ Gulella sp. nov. ” – Verdcourt 1958: 94, 100, fig. 10. Gulella cuspidata – van Bruggen 1969: 71. — Verdcourt 1983: 232. — Richardson 1988: 72. — Verdcourt 1996: 136. — Tattersfield 1998 b: 37. — Tattersfield et al. 1998: 135. — van Bruggen 2000: 233. — Verdcourt 2006: 46. — Rowson & Lange 2007: 31. — Muratov 2010: 277. Type material examined TANZANIA: holotype MRAC.792352: 1 ad., Worlds View, Shume, West Usambara Mts (4.70°S, 38.20°E), in rather dry evergreen forest, leg. B. & L. Verdcourt, Dec. 1956. Other material examined None. Description SHELL (Figs 31, 59). Large (4.30 - 4.80 mm high x 1.80 x 2.50 mm wide), of 7.75 whorls. Ovate-acuminate, spire coeloconoid (spire angle 58°). Apex sharply pointed. Embryonic whorls appear smooth but with fine regular radial striae. Later whorls with relatively few, coarse ribs (about 8 per mm on penultimate whorl). Sutures of intermediate depth. Umbilicus narrowly open. Peristome incomplete parietally. Outer palatal surface of aperture with a furrow-like depression corresponding to the upper palatal tooth. Dentition 6-fold (alternatively recognisable as 5-fold), consisting of: one lamella-like parietal tooth; two palatal teeth, the upper much larger and forming a parieto-palatal sinus; one basal, in-running tooth; and a strong, squarish columellar tooth, running in to form a columellar baffle which is connected to it. Juvenile shells and anatomy unknown. Range and habitat Elevation not stated but probably between 1500-2000 m at the type locality in northeastern Tanzania. Remarks This species differs from D. minareta sp. nov. in having two palatal teeth, in having a basal tooth, in the shape of the parietal tooth, and in size.Published as part of Rowson, Ben & Tattersfield, Peter, 2013, Revision of Dadagulella gen. nov., the " Gulella radius group " (Gastropoda: Streptaxidae) of the eastern Afrotropics, including six new species and three new subspecies, pp. 1-46 in European Journal of Taxonomy 37 on page 20, DOI: 10.5852/ejt.2013.37, http://zenodo.org/record/380677

    Dadagulella ecclesiola Rowson & Tattersfield 2013, sp. nov.

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    Dadagulella ecclesiola sp. nov. urn:lsid:zoobank.org:act: CCE 56537- B 647-4144- AB 07- FE 322 A 139087 Figs 28, 49, 56, 84; Table 1 Etymology From Latin ‘ ecclesiola ’, feminine, diminutive of ‘church’ (or ‘the church’); used arbitrarily to distinguish the species from D. minareta sp. nov. with which it occurs. Type material examined TANZANIA: holotype NMW. Z.2003.001.00015: 1 ad., Kimboza Forest Reserve (7.01° S, 37.78° E), Uluguru Mts, Morogoro District, lowland forest on limestone, approx. 300 m alt., leg. BR, PT, MBS, & CFN, 5 Feb. 2003. Paratypes NMW. Z.2003.001.00016: 8 ads, data as holotype. Paratype MRAC. MT.803794: 1 ad., data as holotype. Paratype NHMUK.20120259: 1 ad., data as holotype. Paratype NMK: 1 ad., data as holotype. Paratype NMT: 1 ad., data as holotype. Paratype NMSA. L 8692/ T 3061: 1 ad., data as holotype. Paratype RMNH. MOL.288089: 1 ad., data as holotype. Other material examined TANZANIA: NMW. Z.2003.001.00017: 1 ad., data as holotype, sequenced by Rowson et al. (2010 a) as “ Gulella cf. browni Uluguru ”. NMW. Z.2003.001.00018: 3 ads in poor condition, 3 juvs, data as holotype. Description SHELL (Figs 28, 49, 56). Small to medium-sized (3.20 - 3.70 mm high x 1.80 - 1.90 mm wide), of 5.5 - 7.0 whorls. Ovate-acuminate, spire narrowly accuminate (spire angle 53 - 65°). Apex pointed. Embryonic whorls smoothly granulate. Later whorls with relatively coarse, often sinuous ribs (8 - 14 per mm on penultimate whorl). Sutures deep. Umbilicus closed. Peristome complete. Outer palatal surface of aperture with a depression, often furrow-like, corresponding to the palatal tooth. Dentition 4-fold, consisting of: one strongly oblique parietal tooth, V-shaped when shell turned to the left; one large slablike palatal tooth, forming a narrow, horizontal parieto-palatal sinus; and a deep-set columellar baffle. A basal denticle is also present, presumably in all specimens, but is partly or completely hidden by the palatal tooth which occludes much of the aperture. The denticle is visible when the shell is turned to the right (Fig. 56). Juvenile shells not known with certainty: an individual from Kimboza (Fig. 49), with dentition like that of a juvenile D. r. radius comb. nov., might belong to this species. Anatomy unknown. Range and habitat In forest at the type locality in the eastern Tanzanian lowlands. Remarks This species has much simpler dentition than D. minareta sp. nov. (which also occurs at Kimboza), and is usually smaller, with straight rather than sinuous ribs. Its dentition is more like that of D. r. radius comb. nov. (which again also occurs at Kimboza; Fig. 16) and D. minuscula minuscula (Morelet, 1877) comb. nov. but D. ecclesiola sp. nov. lacks the shallow columellar tooth, has a hidden or partly hidden basal denticle, and has a much narrower parieto-palatal sinus. It further differs from D. minuscula minuscula comb. nov. in having stronger ribs.Published as part of Rowson, Ben & Tattersfield, Peter, 2013, Revision of Dadagulella gen. nov., the " Gulella radius group " (Gastropoda: Streptaxidae) of the eastern Afrotropics, including six new species and three new subspecies, pp. 1-46 in European Journal of Taxonomy 37 on page 16, DOI: 10.5852/ejt.2013.37, http://zenodo.org/record/380677

    Dadagulella delgada Rowson & Tattersfield 2013, comb. nov.

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    Dadagulella delgada (Muratov, 2010) comb. nov. Figs 27, 84; Table 1 Gulella delgada Muratov, 2010: 274, 276-277, figs 37, 39-45. Type material examined None. Type locality “ Mozambique: Cabo Delgado: 1.1 km WNW of lighthouse, 19 km NE of Palma, 10.68883° S, 40.62806° E, alt. 11 m, 24 Nov. 2009, I. V. Muratov.” (Muratov 2010). Other material examined None. Description SHELL (Fig. 27). Medium-sized (3.80 - 4.00 mm high x 1.80 - 1.80 mm wide), of 6.5 - 7.5 whorls. Elongate ovate-acuminate, spire narrowly acuminate (spire angle 43 - 49°). Apex sharply pointed (with axis slightly deviated in one syntype). Embryonic whorls smooth. Later whorls with very few, widely-spaced, flaring, subtriangular, lamella-like ribs (about 7 per mm on penultimate whorl). Sutures very deep. Umbilicus narrowly open. Peristome incomplete parietally. Outer palatal surface of aperture with a depression corresponding to the palatal tooth. Dentition 5-fold, consisting of: one lamella-like parietal tooth; one slab-like palatal tooth, forming a parieto-palatal sinus; one basal denticle; a deep-set columellar baffle and one shallower columellar denticle. Juvenile shells with 3-fold dentition: one parietal lamella; one basal tooth and one columellar thickening. Muratov (2010) found no internal dentition in the upper whorls. Anatomy unknown. Range and habitat At the type locality, vegetation on coral rag (Muratov 2010: 284). Remarks The few, flaring, lamella-like ribs and elongate, narrowly acuminate spire of D. delgada comb. nov. allow it to be separated from other species, including some D. radius comb. nov. s.l. which it resembles in size and dentition.Published as part of Rowson, Ben & Tattersfield, Peter, 2013, Revision of Dadagulella gen. nov., the " Gulella radius group " (Gastropoda: Streptaxidae) of the eastern Afrotropics, including six new species and three new subspecies, pp. 1-46 in European Journal of Taxonomy 37 on page 15, DOI: 10.5852/ejt.2013.37, http://zenodo.org/record/380677

    Dadagulella frontierarum Rowson & Tattersfield 2013, sp. nov.

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    Dadagulella frontierarum sp. nov. urn:lsid:zoobank.org:act: 9D028 B 17-9FCB-47 E 8-96 B 1-5901 B 8 E 3 EA 4 E Figs 34, 61, 84; Table 1 Etymology After Frontier Tanzania, the organisation which collected the specimens; given the ending ‘- arum ’ for a feminine noun in the genitive plural. Type material examined TANZANIA: holotype NMW. Z.2003.074.00001: 1 ad., Mtai Forest Reserve (4.87° S, 38.77° E), East Usambara Mts, Muheza District (plot 69/3), submontane forest at 970 m alt., leg. Frontier Tanzania, 8 Mar. 1997. Paratypes NMW. Z.2003.074.00002: 2 ads, data as holotype. Paratype NMW. Z.2003.074.00003: 1 ad., data as holotype. Paratype NMW. Z.2003.074.00004: 1 ad., data as holotype but 7 Mar. 1997. Paratypes NMT: 2 ads, data as holotype. Other material examined None. Description SHELL (Figs 34, 61). Medium-sized (3.15 - 3.40 mm high x 1.80 - 2.05 mm wide), of 4.5 - 5.0 whorls. Ovateacuminate, spire narrowly acuminate (spire angle 68 - 77°). Apex sharply pointed. Embryonic whorls smoothly granulate. Later whorls with few, widely-spaced, coarse ribs (5 - 7 per mm on penultimate whorl). Sutures very deep. Umbilicus narrowly open. Peristome complete, almost detached. Outer palatal surface of aperture with a depression corresponding to the lower palatal teeth. Dentition 6-fold, consisting of: one complex, V-shaped and flaring parietal tooth; three palatal teeth, the lower two larger and set low down on the palatal surface, not forming a parieto-palatal sinus; one basal, in-running denticle; and one shallow but strong, in-running columellar tooth. Shells and anatomy of juveniles unknown. Range and habitat Submontane forest (970 m elevation) in the East Usambara Mountains, northeastern Tanzania. Remarks The complex dentition and detached peristome of D. frontierarum sp. nov. are unlike that of any other Dadagulella gen. nov. species. It is also characteristic for its few whorls, few ribs and sharply pointed apex. Biogeographically a close relationship with other species in and around the East Usambaras would seem likely but there is no strong resemblance.Published as part of Rowson, Ben & Tattersfield, Peter, 2013, Revision of Dadagulella gen. nov., the " Gulella radius group " (Gastropoda: Streptaxidae) of the eastern Afrotropics, including six new species and three new subspecies, pp. 1-46 in European Journal of Taxonomy 37 on page 23, DOI: 10.5852/ejt.2013.37, http://zenodo.org/record/380677

    Dadagulella browni subsp. mafiensis Rowson & Tattersfield 2013, subsp. nov.

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    Dadagulella browni mafiensis subsp. nov. Figs 21, 54, 84; Table 1 Etymology From Mafia Island, with the final ‘a’ elided for euphony. Type material examined NHMUK.20110174: 1 ad., Mlula, Mafia I. (approx. 7.87° S, 39.76° E), evergreen coastal thicket on clay loam coral rag, leg. Frontier Tanzania, Oct.- Nov. 1990, labelled “ Gulella radius aggreg.” by B. Verdcourt. Other material examined None. Description SHELL (Figs 21, 54). Small (3.30 mm high x 1.80 mm wide), of 6.0 whorls. Ovate-acuminate, spire broadly acuminate (spire angle 65°). Apex conical. Embryonic whorls smoothly granulate. Later whorls with relatively coarse ribs (12 per mm on penultimate whorl). Sutures relatively shallow. Umbilicus narrowly open. Peristome incomplete parietally. Outer palatal surface of aperture with a depression corresponding to the palatal tooth. Dentition 5-fold, consisting of: one lamella-like parietal tooth and one additional parietal denticle; one slab-like palatal tooth, forming a narrow parieto-palatal sinus; one basal denticle; and a deep-set, folded columellar baffle, but no shallower columellar denticles. Shells and anatomy of juveniles unknown. Range and habitat Evergreen coastal thicket at the type locality. Remarks This taxon is in some respects intermediate between the type specimens of D. r. radius comb. nov. and D. b. browni comb. nov., and thus distinct from either, yet difficult to place. It is larger and not as squat as other D. browni comb. nov. s.l., and lacks the additional basal denticle and shallower columellar dentition. Conversely, it has a parietal denticle not seen in D. radius comb. nov. s.l, is smaller and squatter than most D. radius comb. nov. s.l., and has a longer and narrower parieto-palatal sinus than any D. radius comb. nov. s.l. Mafia Island, from which no other Dadagulella gen. nov. are yet known, lies at almost the same latitude (7.8° S) as the apparent northernmost limit of D. b. browni comb. nov. in mainland Tanzania. Given the morphology of this specimen, its latitude and its isolation as an island population, we treat it as a subspecies of D. browni comb. nov.Published as part of Rowson, Ben & Tattersfield, Peter, 2013, Revision of Dadagulella gen. nov., the " Gulella radius group " (Gastropoda: Streptaxidae) of the eastern Afrotropics, including six new species and three new subspecies, pp. 1-46 in European Journal of Taxonomy 37 on pages 13-14, DOI: 10.5852/ejt.2013.37, http://zenodo.org/record/380677
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