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    Figs 65-68 in Revision of Dadagulella gen. nov., the "Gulella radius group" (Gastropoda: Streptaxidae) of the eastern Afrotropics, including six new species and three new subspecies

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    Figs 65-68. Radulae of Dadagulella gen. nov. species. 65. D. radius radius, Amboni. 66. D. pembensis sp. nov., Ngezi, radula in situ on odontophore, ventral view. 67. D. pembensis sp. nov., Ngezi. 68. D. pembensis sp. nov., Ngezi, teeth from ventral end of radular ribbon. Abbreviations: c = central tooth; ii, iii, and iv = examples of bi-, tri- and quadricuspid teeth respectively. All scalebars = 10 μm.Published as part of Rowson, Ben & Tattersfield, Peter, 2013, Revision of Dadagulella gen. nov., the "Gulella radius group" (Gastropoda: Streptaxidae) of the eastern Afrotropics, including six new species and three new subspecies, pp. 1-46 in European Journal of Taxonomy 37 on page 37, DOI: 10.5852/ejt.2013.37, http://zenodo.org/record/380677

    Fig. 4 in Revision of the carnivorous snail genus Indoartemon Forcart, 1946 and a new genus Carinartemis from Thailand (Pulmonata: Streptaxidae)

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    Fig. 4. Genitalia of Indoartemon medius, new species, paratype CUMZ 5017: A, reproductive system; B, attachment of vas deferens into distal end of the penial sheath; C, details of albumin gland and talon.Published as part of Siriboon, Thanit, Sutcharit, Chirasak, Naggs, Fred, Rowson, Ben & Panha, Somsak, 2014, Revision of the carnivorous snail genus Indoartemon Forcart, 1946 and a new genus Carinartemis from Thailand (Pulmonata: Streptaxidae), pp. 161-174 in Raffles Bulletin of Zoology 62 on page 166, DOI: 10.5281/zenodo.450407

    Figs 6-26 in Revision of Dadagulella gen. nov., the "Gulella radius group" (Gastropoda: Streptaxidae) of the eastern Afrotropics, including six new species and three new subspecies

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    Figs 6-26. Adult shells of Dadagulella radius (Preston, 1910) comb. nov., D. browni (van Bruggen, 1969) comb. nov. and their respective subspecies. 6-20. D. radius comb. nov. s.l. 6-8. D. radius calva (Connolly, 1922): 6. Lectotype, Taru Desert. 7. Taru Desert near Malindi. 8. Near Mombasa. 9-20. D. radius radius: 11. Lectotype, Shimba Hills. 12. Gazi. 13. also Gazi. 14. Diani Beach. 15. Amboni. 16. Kimboza. 17. Mkungwe. 18. Pugu. 19. Mbudya I. 20. Jozani. 21-26. D. browni comb. nov. s.l.: 21. D. browni mafiensis subsp. nov., holotype, Mlula. 22. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 23-26. D. browni browni: 23. Mwanihana. 24. Mzelezi. 25. Pomene Bay. 26. Holotype, Lake Sibayi (after van Bruggen 1969).Published as part of Rowson, Ben & Tattersfield, Peter, 2013, Revision of Dadagulella gen. nov., the "Gulella radius group" (Gastropoda: Streptaxidae) of the eastern Afrotropics, including six new species and three new subspecies, pp. 1-46 in European Journal of Taxonomy 37 on page 10, DOI: 10.5852/ejt.2013.37, http://zenodo.org/record/380677

    Fig. 7 in Revision of the carnivorous snail genus Indoartemon Forcart, 1946 and a new genus Carinartemis from Thailand (Pulmonata: Streptaxidae)

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    Fig. 7. Genitalia of Carinartemis spp: A, B, Carinartemis vesperus, new species, paratype CUMZ 6201, (A) reproductive system, and (B) insertion of vas deferens into penial sheath; C, D, Carinartemis striatus, new species, paratype CUMZ 6205, (C) reproductive system, and (D) attachment of vas deferens into distal end of the penial sheath.Published as part of Siriboon, Thanit, Sutcharit, Chirasak, Naggs, Fred, Rowson, Ben & Panha, Somsak, 2014, Revision of the carnivorous snail genus Indoartemon Forcart, 1946 and a new genus Carinartemis from Thailand (Pulmonata: Streptaxidae), pp. 161-174 in Raffles Bulletin of Zoology 62 on page 169, DOI: 10.5281/zenodo.450407

    Fig. 5 in Revision of the carnivorous snail genus Indoartemon Forcart, 1946 and a new genus Carinartemis from Thailand (Pulmonata: Streptaxidae)

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    Fig. 5. Internal sculpture of genitalia of Indoartemon medius, new species, paratype CUMZ 5017: A, details of atrial pore on the atrium surface; B, low magnification shows arrangement of penial hooks; C, high magnification of penial hooks in proximal area; D, top view of penial hook, hooks shrunken in tubed penial papillae; E, low magnification of penial hook in distal area; F, lateral view of penial hook in distal area; G, high magnification of penial hooks in distal area; H, arrangement of vaginal folds; I, radula morphology.Published as part of Siriboon, Thanit, Sutcharit, Chirasak, Naggs, Fred, Rowson, Ben & Panha, Somsak, 2014, Revision of the carnivorous snail genus Indoartemon Forcart, 1946 and a new genus Carinartemis from Thailand (Pulmonata: Streptaxidae), pp. 161-174 in Raffles Bulletin of Zoology 62 on page 167, DOI: 10.5281/zenodo.450407

    Dadagulella delgada Rowson & Tattersfield 2013, comb. nov.

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    Dadagulella delgada (Muratov, 2010) comb. nov. Figs 27, 84; Table 1 Gulella delgada Muratov, 2010: 274, 276-277, figs 37, 39-45. Type material examined None. Type locality “ Mozambique: Cabo Delgado: 1.1 km WNW of lighthouse, 19 km NE of Palma, 10.68883° S, 40.62806° E, alt. 11 m, 24 Nov. 2009, I. V. Muratov.” (Muratov 2010). Other material examined None. Description SHELL (Fig. 27). Medium-sized (3.80 - 4.00 mm high x 1.80 - 1.80 mm wide), of 6.5 - 7.5 whorls. Elongate ovate-acuminate, spire narrowly acuminate (spire angle 43 - 49°). Apex sharply pointed (with axis slightly deviated in one syntype). Embryonic whorls smooth. Later whorls with very few, widely-spaced, flaring, subtriangular, lamella-like ribs (about 7 per mm on penultimate whorl). Sutures very deep. Umbilicus narrowly open. Peristome incomplete parietally. Outer palatal surface of aperture with a depression corresponding to the palatal tooth. Dentition 5-fold, consisting of: one lamella-like parietal tooth; one slab-like palatal tooth, forming a parieto-palatal sinus; one basal denticle; a deep-set columellar baffle and one shallower columellar denticle. Juvenile shells with 3-fold dentition: one parietal lamella; one basal tooth and one columellar thickening. Muratov (2010) found no internal dentition in the upper whorls. Anatomy unknown. Range and habitat At the type locality, vegetation on coral rag (Muratov 2010: 284). Remarks The few, flaring, lamella-like ribs and elongate, narrowly acuminate spire of D. delgada comb. nov. allow it to be separated from other species, including some D. radius comb. nov. s.l. which it resembles in size and dentition.Published as part of Rowson, Ben & Tattersfield, Peter, 2013, Revision of Dadagulella gen. nov., the " Gulella radius group " (Gastropoda: Streptaxidae) of the eastern Afrotropics, including six new species and three new subspecies, pp. 1-46 in European Journal of Taxonomy 37 on page 15, DOI: 10.5852/ejt.2013.37, http://zenodo.org/record/380677

    Dadagulella meredithae Rowson & Tattersfield 2013, comb. nov.

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    Dadagulella meredithae (van Bruggen, 2000) comb. nov. Figs 37, 84; Table 1 Gulella meredithae van Bruggen, 2000: 226-232, figs 1-7. “…an as yet unidentified species from Malawi ” – van Bruggen & Meredith 1984: 165. Gulella meredithae – Rowson & Lange 2007: 31. Type material examined MALAWI: holotype RMNH.59399: 1 ad., Nyika National Park (approx. 10.6°S, 33.8°E), Rumphi District, Juniper Forest, approx. 2100 m alt., leg. H.M. Meredith, 16 Sep. 1983. Other material examined None. Description SHELL (Fig. 37). Small (2.30 - 3.10 mm high x 1.40 - 1.60 mm wide), of 5.5 - “<6” whorls. Ovate-acuminate, although spire (spire angle 58 - 77°) less acuminate than in other Dadagulella gen. nov species. Apex rounded (in holotype) to weakly pointed. Embryonic whorls smoothly granulate. Later whorls with very fine, very numerous ribs (about 27 per mm on penultimate whorl). Sutures of intermediate depth. Umbilicus closed or nearly so. Peristome incomplete parietally. Outer palatal surface of aperture with a depression corresponding to the palatal tooth. Dentition 3-fold to 4-fold, consisting of at least: one lamella-like parietal tooth; one slab-like palatal tooth, not forming a parieto-palatal sinus; and a mammillate columellar baffle. Additional teeth limited to a shallow, weak columellar swelling. Juvenile shells with 2-fold to 3-fold dentition: one parietal tooth; one columellar tooth; and usually one basal tooth (termed labral by van Bruggen 2000). Van Bruggen (2000) showed that earlier sets of dentition are visible through the shells of some transparent juveniles, even in the preceding whorls, suggesting slow or no resorbtion. Anatomy unknown. Range and habitat Montane forest (above 1500 m to around 2450 m) in northern and central Malawi, and adjacent part of Zambia (Chowo Forest). Van Bruggen (2000) suspected it to range into parts of Tanzania adjoining northern Malawi. Remarks This species is distinctive in its small size, very fine, numerous ribs, and dentition. It differs from D. radius comb. nov. s.l. and D. browni comb. nov. s.l. in the lack of a basal tooth or denticle. The apex is rounded in the holotype but more conical in paratypes figured by van Bruggen (2000).Published as part of Rowson, Ben & Tattersfield, Peter, 2013, Revision of Dadagulella gen. nov., the " Gulella radius group " (Gastropoda: Streptaxidae) of the eastern Afrotropics, including six new species and three new subspecies, pp. 1-46 in European Journal of Taxonomy 37 on pages 26-27, DOI: 10.5852/ejt.2013.37, http://zenodo.org/record/380677

    Dadagulella cresswelli Rowson & Tattersfield 2013, sp. nov.

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    Dadagulella cresswelli sp. nov. urn:lsid:zoobank.org:act: B 3362D6 F- 2118 -4D1 D- 8 B 25-098 E 25 B 0 C 3 B 2 Figs 29, 57, 84; Table 1 Etymology After Pete Cresswell, who collected the specimen. Type material examined TANZANIA: holotype NMW. Z.2012.042.00001: 1 ad., Ngorongoro Crater, Arusha Region, crater rim on southeastern side, heavy rainforest leaf litter, leg. P. L. Cresswell, 2 Jun.1996. Other material examined None. Description SHELL (Figs 29, 57). Medium-sized (3.70 mm high x 1.80 mm wide), of 7.0 whorls. Ovate-acuminate, spire coeloconoid (spire angle 52°). Apex sharply pointed. Embryonic whorls smoothly granulate. Later whorls with relatively fine ribs (13 per mm on penultimate whorl). Sutures shallow. Umbilicus narrowly open. Peristome complete. Outer palatal surface of aperture with a very deep, long, furrow-like depression corresponding to the palatal tooth. Dentition 5-fold, consisting of: one V-shaped parietal tooth; one bifid slab-like palatal tooth, forming a clear parieto-palatal sinus, with the upper cusp projecting into the sinus; a deep-set columellar baffle and two shallower columellar denticles. Shells and anatomy of juveniles unknown. Range and habitat Forest at the type locality in northern Tanzania. The vegetation is presumably of a montane type, since the crater floor is above 1700 m while the rim rises to over 2400 m or higher. Remarks This species is distinctive in its deep, long furrow on the outer palatal surface in combination with the coeloconoid spire and dentition. D. minerata sp. nov. shares these features, but differs in having weaker ribs and less complex dentition. It is the only Dadagulella gen. nov. species thus far collected in the volcanic (as opposed to block-faulted) highlands of Tanzania or Kenya.Published as part of Rowson, Ben & Tattersfield, Peter, 2013, Revision of Dadagulella gen. nov., the " Gulella radius group " (Gastropoda: Streptaxidae) of the eastern Afrotropics, including six new species and three new subspecies, pp. 1-46 in European Journal of Taxonomy 37 on page 18, DOI: 10.5852/ejt.2013.37, http://zenodo.org/record/380677

    Dadagulella frontierarum Rowson & Tattersfield 2013, sp. nov.

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    Dadagulella frontierarum sp. nov. urn:lsid:zoobank.org:act: 9D028 B 17-9FCB-47 E 8-96 B 1-5901 B 8 E 3 EA 4 E Figs 34, 61, 84; Table 1 Etymology After Frontier Tanzania, the organisation which collected the specimens; given the ending ‘- arum ’ for a feminine noun in the genitive plural. Type material examined TANZANIA: holotype NMW. Z.2003.074.00001: 1 ad., Mtai Forest Reserve (4.87° S, 38.77° E), East Usambara Mts, Muheza District (plot 69/3), submontane forest at 970 m alt., leg. Frontier Tanzania, 8 Mar. 1997. Paratypes NMW. Z.2003.074.00002: 2 ads, data as holotype. Paratype NMW. Z.2003.074.00003: 1 ad., data as holotype. Paratype NMW. Z.2003.074.00004: 1 ad., data as holotype but 7 Mar. 1997. Paratypes NMT: 2 ads, data as holotype. Other material examined None. Description SHELL (Figs 34, 61). Medium-sized (3.15 - 3.40 mm high x 1.80 - 2.05 mm wide), of 4.5 - 5.0 whorls. Ovateacuminate, spire narrowly acuminate (spire angle 68 - 77°). Apex sharply pointed. Embryonic whorls smoothly granulate. Later whorls with few, widely-spaced, coarse ribs (5 - 7 per mm on penultimate whorl). Sutures very deep. Umbilicus narrowly open. Peristome complete, almost detached. Outer palatal surface of aperture with a depression corresponding to the lower palatal teeth. Dentition 6-fold, consisting of: one complex, V-shaped and flaring parietal tooth; three palatal teeth, the lower two larger and set low down on the palatal surface, not forming a parieto-palatal sinus; one basal, in-running denticle; and one shallow but strong, in-running columellar tooth. Shells and anatomy of juveniles unknown. Range and habitat Submontane forest (970 m elevation) in the East Usambara Mountains, northeastern Tanzania. Remarks The complex dentition and detached peristome of D. frontierarum sp. nov. are unlike that of any other Dadagulella gen. nov. species. It is also characteristic for its few whorls, few ribs and sharply pointed apex. Biogeographically a close relationship with other species in and around the East Usambaras would seem likely but there is no strong resemblance.Published as part of Rowson, Ben & Tattersfield, Peter, 2013, Revision of Dadagulella gen. nov., the " Gulella radius group " (Gastropoda: Streptaxidae) of the eastern Afrotropics, including six new species and three new subspecies, pp. 1-46 in European Journal of Taxonomy 37 on page 23, DOI: 10.5852/ejt.2013.37, http://zenodo.org/record/380677

    Dadagulella browni subsp. mafiensis Rowson & Tattersfield 2013, subsp. nov.

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    Dadagulella browni mafiensis subsp. nov. Figs 21, 54, 84; Table 1 Etymology From Mafia Island, with the final ‘a’ elided for euphony. Type material examined NHMUK.20110174: 1 ad., Mlula, Mafia I. (approx. 7.87° S, 39.76° E), evergreen coastal thicket on clay loam coral rag, leg. Frontier Tanzania, Oct.- Nov. 1990, labelled “ Gulella radius aggreg.” by B. Verdcourt. Other material examined None. Description SHELL (Figs 21, 54). Small (3.30 mm high x 1.80 mm wide), of 6.0 whorls. Ovate-acuminate, spire broadly acuminate (spire angle 65°). Apex conical. Embryonic whorls smoothly granulate. Later whorls with relatively coarse ribs (12 per mm on penultimate whorl). Sutures relatively shallow. Umbilicus narrowly open. Peristome incomplete parietally. Outer palatal surface of aperture with a depression corresponding to the palatal tooth. Dentition 5-fold, consisting of: one lamella-like parietal tooth and one additional parietal denticle; one slab-like palatal tooth, forming a narrow parieto-palatal sinus; one basal denticle; and a deep-set, folded columellar baffle, but no shallower columellar denticles. Shells and anatomy of juveniles unknown. Range and habitat Evergreen coastal thicket at the type locality. Remarks This taxon is in some respects intermediate between the type specimens of D. r. radius comb. nov. and D. b. browni comb. nov., and thus distinct from either, yet difficult to place. It is larger and not as squat as other D. browni comb. nov. s.l., and lacks the additional basal denticle and shallower columellar dentition. Conversely, it has a parietal denticle not seen in D. radius comb. nov. s.l, is smaller and squatter than most D. radius comb. nov. s.l., and has a longer and narrower parieto-palatal sinus than any D. radius comb. nov. s.l. Mafia Island, from which no other Dadagulella gen. nov. are yet known, lies at almost the same latitude (7.8° S) as the apparent northernmost limit of D. b. browni comb. nov. in mainland Tanzania. Given the morphology of this specimen, its latitude and its isolation as an island population, we treat it as a subspecies of D. browni comb. nov.Published as part of Rowson, Ben & Tattersfield, Peter, 2013, Revision of Dadagulella gen. nov., the " Gulella radius group " (Gastropoda: Streptaxidae) of the eastern Afrotropics, including six new species and three new subspecies, pp. 1-46 in European Journal of Taxonomy 37 on pages 13-14, DOI: 10.5852/ejt.2013.37, http://zenodo.org/record/380677
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