101,022 research outputs found
Lebinthus estrellae Robillard 2015
No full textLebinthus estrellae Robillard, 2015 (Fig. 1) Lebinthus estrellae Robillard, 2015, in Robillard & Yap, 2015: 63 (original description); Baroga et al., 2016: 94 (taxonomic key). Type material. Holotype male. Philippines. [Visayas] Leyte: Burauen, Barangay Villa Corazon, forêt secondaire sur pente [secondary forest on slope] (GPS Bar2), 10°57’52.1”N 124°46’39.8”E, 345 m, (TR29), iii.2013, coll. T. Robillard (UPLBMNH) [examined]. Paratypes (2♂, 15♀): same information as holotype (MNHN) [examined]. Type locality. Leyte: Philippines. Burauen, Barangay Villa Corazon, secondary forest on slope Distribution. Philippines: Leyte, Barauen Diagnosis. This species can be recognized from other species of the sanchezi group by its dark brown colouration, short rounded lophi in male genitalia, presence of vertical whitish bands below eyes, clypeus black with yellow edge; male genitalia with short rounded lophi, and very short female FWs (reaching posterior margin of first tergite). Habitat. Found in bushes and leaf litter.Published as part of Baroga-Barbecho, Jessica B., Tan, Ming Kai, Yap, Sheryl A. & Robillard, Tony, 2020, Taxonomic study of Lebinthus Stål, 1877 (Orthoptera: Gryllidae: Eneopterinae) with description of six new species in the Philippines, pp. 401-438 in Zootaxa 4816 (4) on page 415, DOI: 10.11646/zootaxa.4816.4.1, http://zenodo.org/record/395452
Cardiodactylus singapura Robillard, sp. nov.
No full textCardiodactylus singapura Robillard, sp. nov. (Figs 1–5) Type material. Holotype male: Singapore. Central Catchment Nature Reserve, 01° 22 ' 58.3 ''N 103 ° 48 ' 50.3 ''E, 37 m, 4.VII. 2009, nuit, adulte en élevage, enregistrement appel (F0-male 4), T. Robillard (ZRC). Allotype female: Singapore. Bukit Timah, 01° 21 '01.0''N 103 ° 46 ' 44.3 ''E, 95 m, VII. 2009, adulte en élevage, T. Robillard (ZRC). Paratypes: Singapore. MacRitchie Reservoir, 01° 21 ' 10.2 ''N 103 ° 49 ' 32.2 ''E, 36 m, 2.VII. 2009, nuit, adultes en élevage, T. Robillard: 23, enregistrement appel (F0-males 1-2) (MNHN-ENSIF 2754-2755); 1 Ƥ (MNHN- ENSIF 2756); 1 Ƥ (ZRC). Central Catchment Nature Reserve, 01° 22 ' 58.3 ''N 103 ° 48 ' 50.3 ''E, 37 m, 4.VII. 2009, nuit, adultes en élevage, T. Robillard: 13, enregistrement appel (F0-male 3) (MNHN-ENSIF 2757); 1 Ƥ (MNHN- ENSIF 2758), 13, 1Ƥ (ZRC). Bukit Timah, 01° 21 '08.4''N 103 ° 46 ' 43.8 ''E, 116 m, VII. 2009, nuit (8: 30 PM), T. Robillard: 13, adulte en élevage, enregistrement appel (F0-male 6) (MNHN-ENSIF 2759); 13 (TR 39), sur plante de sous-bois (feuille, h= 1.5 m), 2 Ƥ, adultes en élevage, (ZRC). Singapore [no precision], 13, Coll. Baker, 14618 (MNHN-ENSIF 2753). Singapore [no precision], VII- 2009, T. Robillard, nuit, adulte en élevage, 13, enregistrement appel (F0-male 5) (MNHN-ENSIF 2760) Type locality. Singapore, Central Catchment Nature Reserve, 01° 22 ' 58.3 ''N 103 ° 48 ' 50.3 ''E, 37 m. Etymology. Named after the type locality. Other material examined. Singapore. Bukit Timah, 01° 21 '08.4''N 103 ° 46 ' 43.8 ''E, 116 m, 1.VII. 2009, nuit, T. Robillard: 1 juvenile (MNHN). Central Catchment Nature Reserve, 01° 22 ' 58.3 ''N 103 ° 48 ' 50.3 ''E, 37 m, 4.VII. 2009, nuit, T. Robillard: 1 juvenile (MNHN). Singapore [no precision], reared specimens (F 1 generation, TR- 2010): 53, 2Ƥ (MNHN). Distribution. Singapore island. Diagnosis. Species small with contrasted coloration, close to C. pelagus Otte, 2007 from Borneo, from which it differs by the shape of the pseudepiphallus in male genitalia and lighter colour pattern. Description. Size small for the species group. General coloration contrasted, light brown with dark brown patterns. Head dorsum yellow brown with 4 wide dark brown bands, more or less punctuated, bordered by dark brown triangles posterior to eyes and a yellow line more externally (Figs 1 A, 2 A). Fastigium mottled with dark brown. Scapes light brown with a transverse dark brown line anteriorly (Figs 1 A, 2 B). Antennae heterogeneously brown. Face yellowish to light brown, diversely mottled with brown, with 2 parallel lines on the fastigium, 4 dark spots on face, and 2 dark brown spots ventral to eyes. Mouthparts yellow brown. Maxillary palpi yellowish, dark brown apically. Head lateral side with a dark brown area posterior to eyes, progressively lighter ventrally. Pronotum: Dorsal disk trapezoidal, posterior margin straight or slightly bisinuated; light brown mottled with greyish brown and dark brown spots; anterior corners yellowish. Lateral lobes dark brown, ventral edges yellow. Legs light brown to yellowish brown, femora mottled with dark brown; tibiae more or less banded; tarsomeres with dark brown ends. Tarsomeres III- 1 with 2–3 spines on dorso-external edges (m= 2.2, n= 10). Hind wings longer than FWs, the dark brown tail exceeding the forewings twice longer than the pronotum. Cerci light brown with black rings. Tergites light brown mottled with black. Male: FW coloration (Figs 1 B, 2 C): Dorsal field cells dark brown, veins mostly orange brown, except for whitish areas in the chords, part of anal veins and in the region posterior to mirror. Base of chords with a brown sclerotization. Harp posterior angle with a whitish sclerotized area. Lateral field dark brown; veins MA, MP, R orange brown, transverse veins and projections of R whitish. FW venation: 1 A slightly bisinuated; stridulatory vein (Figs 2 C, 2 D) with 156–171 teeth (Fig. 2 E) (m= 164, n= 4), distributed on both transverse (123–143 teeth, m= 133) and longitudinal (15–20 teeth, m= 19) parts of 1 A, plus a group of 9–16 teeth near base of 1 A (m= 13, n= 4). CuP missing. Harp longer than wide, with 2 w-shaped harp veins; posterior margin raised along diagonal vein. Mirror area variable, c 1 long and narrow, c 2 quite large; mirror (d 1) longer than wide, not rounded, generally separated in two parts by a transverse vein, the posterior part triangular; e 1 cell very elongated, along the mirror. Apical field long and triangular, with 3 wide cell alignments posterior to mirror and a narrow apical alignment. Lateral field: R with 6–7 projections (m= 6, n= 5). Subgenital plate yellow brown, the median area dark brown. Male genitalia (Fig. 3): Pseudepiphallus very sclerotized, little setose, slightly narrowed at mid-length. Dorsal ridges parallel, forming a wide gutter, flattened anteriorly, their external edges carenated. Pseudepiphallic sclerite with latero-anterior expansions, but without a membranous sac as in C. novaeguineae; membrane between the expansions with short strong setae. Apex of pseudepiphallus spoon-like, not rounded, and slightly indented. Rami narrow, with wide preapical plates. Pseudepiphallic parameres trilobate, the posterior lobe wide. Ectophallic arc complete, with a short posterior expansion. Ectophallic apodemes thin, divergent. Apex of ectophallic fold almost hidden by pseudepiphallic parameres, trilobate, membranous. Base of ectophallic apodemes with short sclerotized expansions oriented posteriorly. Endophallic sclerite small. Endophallic apodeme with both lateral lamellas and a short medio-dorsal crest. Membrane of endophallic cavity smooth. Female: FW coloration: cells dark brown, veins orange brown, with whitish sclerotization on distal half of CuA and MP, including cells between CuA and MA. MA and R orange brown, R bifurcations whitish. Anterior part of FW with variable number of whitish transverse veins. FW venation (Fig. 4 A): 9–11 (m= 10, n= 5) strong longitudinal veins on dorsal field; lateral field with 9–11 (m= 9.5, n= 5) longitudinal veins including 5–6 R bifurcations and 3–4 ventral veins. Ovipositor: Shorter than hind femora; apex with both dorsal and ventral edges denticulate (Fig. 4 E). Female genitalia (Fig. 4 B–D): Copulatory papilla triangular with baso-lateral sclerotizations; apex rounded and folded ventrally. Juvenile: First instar after hatching observed in laboratory, similar in coloration to that of C. novaeguineae, yellow with brown longitudinal bands. Later instars with more contrasted coloration, body black, legs and head yellow brown (Fig. 1 C– 2 C). Black coloration reverses in sub-adults, characterized by a more homogeneous brown coloration (Fig. 1 C–D). Measurements. See Table 1. PronL PronW FWL FWW HWT FIIIL FIIIW TIIIL Male holotype 2.4 4 14.1 4.3 4.2 14 3.7 11.5 Males (n= 5) 2.2–2.4 3.7–4.2 13.3–14.1 4.1–4.3 4.2–5.4 13.4–15 3.6–4.1 11.5–13.1 (Mean) (2.4) (4) (13.7) (4.2) (4.9) (14.3) (3.9) (12.5) Female allotype 2.5 4.3 14.5 4.2 5.5 15.5 3.9 13.4 Females (n= 5) 2.2–2.7 4.1–4.5 13.5–15.2 3.9–4.4 4.5–5.5 14.1–15.5 3.9–4.4 12.5–13.5 (Mean) (2.4 (4.3) (14.3) (4.1) (5.1) (14.9) (4.1) (13) continued. TIIIs TaIIIs ST OL Ias Ibs Oas Obs Tt Lt Bt Male holotype 8 6 11 7 3 ??? - Males (n= 5) 8–10 6 11 – 13 7 2–3 - (Mean) (8.8) (6) (11.8) (7) 2.2 - Female allotype 6 6 11 7 3 - - - 10.9 Females (n= 5) 6–10 6–7 11 – 13 5–7 2–3 - - - 9.7–10.9 (Mean) (8) (6.4) (11.8) (6.4) (2.2) - - - (10.2) Habitat and life history traits. Cardiodactylus singapura is a nocturnal species living in the forested areas. Males produce their discreet calling songs at night, from low branches and leaves (Fig. 1). Juveniles are found day and night in the leaf litter and on leaves of low plants. Behavior. Calling song (Fig. 5): High speed video observations at 1250 frames per second confirm that each calling song corresponds to only one FW closure, the gaps within the syllable being made by the jerking movement of the FWs. At 26 °C, this mono-syllabic calling song (TR-male 1 -F0male: MNHN-ENSIF 2754) lasts for 87.0 ± 11.8 ms with a period of 8.2 ± 2.2 s and a duty cycle of 1.06 %. The dominant frequency is 19.0 ± 0.7 kHz and corresponds to the third frequency peak of the song.Published as part of Robillard, Tony, 2011, New Cardiodactylus species from unsuspected places in Southeast Asia (Orthoptera, Grylloidea, Eneopterinae), pp. 14-26 in Zootaxa 2909 on pages 16-22, DOI: 10.5281/zenodo.20145
Paranisitra leytensis Robillard & Yap 2015
No full textParanisitra leytensis Robillard, 2015 (Fig. 1) Paranisitra leytensis Robillard & Yap, 2015: 85; Baroga et al. 2016: 94 (taxonomic key). Type material. Holotype male. Philippines. [Visayas]: Leyte [Island], Burauen, Barangay Villa Corazon, zone herbacée [herbaceous area] (GPS Bar1), 10°57’50.5”N 124°46’ 35.3”E, 282 m (TR14), iii.2013, coll. T. Robillard (UPLBMNH). Allotype female. Philippines. [Visayas]: Leyte [Island]: Buo, zone secondaire près de la route [secondary area near road], (TR34), iii.2013, coll. T. Robillard (UPLBMNH). Paratypes (1 male, 5 females). Philippines. same information as HT and AT (MNHN-EO-ENSIF-3162–3166). [examined] Other material examined. Philippines. [Mindanao]: [Surigao del Norte]: Siargao Island, Del Carmen, Brgy. Esperanza, 1♂ (JBB139), 01–05.x.2016, coll. S.A. Yap (UPLBMNH), right middle leg for molecular analysis (N40); Del Carmen, Brgy. Katipunan, 2♀, 07.iv.2018, rainy, coll. J.B. Baroga-Barbecho, S.A. Yap, M.K. Tan, & H. Yeo (UPLBMNH, ZRC); 2♀ (Siargao18_44, 84), 14–17.x.2018, coll. M.K. Tan, J.B. Baroga-Barbecho & S.A. Yap (ZRC). [Mindanao Island: Agusan del Sur]: Casawangan Lake, 1♂ (JBB363), 22.vii.2015, coll. N.M. Barbecho (NMP 13579), left middle leg for molecular analysis (N39). Type locality. Philippines. Visayas, Leyte Island, Burauen, Barangay Villa Corazon, 10°57’50.5”N 124°46’ 35.3”E, 282 m. Distribution. Philippines. Visayas: Leyte Island; Mindanao: Siargao Island (new record); Mindanao Island: Agusan del Sur (new record). Diagnosis. Species close to P. longipes in terms of face coloration with a wide yellow band; differing by its smaller size and male genitalia with smaller and triangular pseudepiphallic lophi, shorter endophallic sclerite, and narrower endophallic apodemes. Remarks. The male specimens from Siargao Island and Mindanao Island (Agusan del Sur) were identified as P. leytensis based on the features of the body, face, size and male genitalia. They slightly differ from the specimens from Leyte by the following characters: in Siargao specimens, male genitalia wider than long, short and less setaceous lophi, and short rami; in Agusan specimen, thin and long male genitalia, ectophallic fold longer and narrowing basally. Male specimen from Mindanao Island is the smallest among examined specimens of the species.Published as part of Baroga-Barbecho, Jessica B., Yap, Sheryl A., Tan, Ming Kai & Robillard, Tony, 2019, Taxonomic review of the genus Paranisitra Chopard (Orthoptera: Gryllidae: Eneopterinae: Nisitrini) with description of a new species from Mindanao, pp. 81-96 in Zootaxa 4568 (1) on page 86, DOI: 10.11646/zootaxa.4568.1.5, http://zenodo.org/record/259926
Lebinthus puyos Robillard 2013
No full textLebinthus puyos Robillard, 2013 (Figs. 1, 6, 7) Lebinthus puyos Robillard, 2013: 49 (original description); Baroga et al. 2016: 94 (taxonomic key). Type material. Holotype male. Philippines. Luzon: Laguna, Los Baños, Mount Makiling, base, secondary forest on campus, 14°09’12.9”N 121°14’05.0”E, 168 m (GPS Maki1), 27.vi.2011, night, leaf litter (TR652), coll. T. Robillard (UPLBMNH) [examined]; Allotype female: same information as holotype, TR653 (UPLBMNH) [examined]; Paratypes (5♂): same information as holotype (MNHN) [examined]. Other material examined. Philippines. [Luzon]: Laguna: College, 50 m, 1♂ (JBB318), 20.ii.1958, coll. E.S. Novero (UPLBMNH ORT-01161); UPLB, Mt. Makiling, 1♀ (JBB106), 06.viii.2014, coll. J.B. Baroga (UPLBMNH); Los Baños, 300 m, 1♀ (JBB349), 21.iv.1960, coll. A. Djamin (UPLBMNH ORT-01192); [Quezon Province], Mt. Banahaw, 1♂ (JBB297), 2.vii.1994, coll. O. Reyes (UPLBMNH ORT-01140); same information as holotype, 6♂, lab colony, call recording (F0-male1, F0-male1-3, F1-male1-2, F5-male1) (MNHN). Type locality. Philippines. Luzon: Laguna, Los Baños, Mount Makiling, base, secondary forest on campus. Distribution. Philippines: Laguna: Mt. Makiling and Los Baños; Quezon: Mt. Banahaw (new record). Diagnosis. This species differs from L. sanchezi by its whitish face, a fastigium that is not orange apically, without clear longitudinal bands in the vertex (Robillard et al., 2013), and male genitalia with long and rounded pseudepiphallic lophi and C-shaped pseudepiphallic parameres. Calling song. The song of L. puyos consists of long trills (duration = 3.1± 0.8 s [1.7– 4.0 s]; period = 25.6± 2.8 s [22.4– 33.1 s]) made up of more than 200 syllables of increasing intensity (Figs. 7 A–C). Syllable duration = 12.6±1.2 ms (10–15 ms). The spectrogram analysis reveals that the frequency spectrum is completely ultrasonic. The syllables show a clear dominant peak at 23.12±1.06 kHz typically corresponding to the first peak of a harmonic series with an intermediary peak at ca. 25 to 30 kHz (Figs. 7D, 7E). Habitat. The species is found in secondary habitats in the type locality, i.e., foot of Mt. Makiling; specimens were found in leaf litter and on top of leaves of small plants (Fig. 6A).Published as part of Baroga-Barbecho, Jessica B., Tan, Ming Kai, Yap, Sheryl A. & Robillard, Tony, 2020, Taxonomic study of Lebinthus Stål, 1877 (Orthoptera: Gryllidae: Eneopterinae) with description of six new species in the Philippines, pp. 401-438 in Zootaxa 4816 (4) on page 412, DOI: 10.11646/zootaxa.4816.4.1, http://zenodo.org/record/395452
Cardiodactylus borneoe Robillard & Gorochov 2014
No full textCardiodactylus borneoe Robillard & Gorochov, 2014 (Figs. 11, 12) Robillard et al., 2014: 12; Dong et al., 2018: 6 (molecular phylogeny); Cigliano et al., 2019 (Orthoptera Species File). Material examined: Malaysia, Sabah, Sandakan, Rainforest Discovery Centre, 1 female (SDK.19.6), N5.87410, E117.94366, 50.2± 5.1 m.a.s.l., 8 January 2019, 2102 hours, on foliage, coll. M. K. Tan & S. T. Toh (ZRC); 1 female (SDK.19.21, Fig. 11A) N5.87542, E117.94231, 31.1± 4.8 m.a.s.l., 9 January 2019, 1930 hours, on foliage, coll. M. K. Tan, R. Japir, Momin Binti & J. Y. Lee (FRC); 1 male (SDK.19.36, Figs. 9 B–9D), N5.87547, E117.94110, 34.1± 5.4 m.a.s.l., 9 January 2019, 2046 hours, under foliage calling, coll. M. K. Tan, R. Japir, Momin Binti & J. Y. Lee (ZRC). Calling song (Fig. 12): At 28.4°C and RH 74%, the calling song of C. borneoe consists of doublets (or rarely triplets) of long syllables including strong amplitude modulation forming pulse-like structures. Echemes (doublet) last 660 ± 75 ms with a period of 3.5 ± 1.2 s. Within doublets, syllables last 196 ± 43 ms, with a period of 372 ± 47 ms. Dominant frequency value is 17.6 ± 0.4 kHz with no clear harmonic structure. Habitat and life history traits. C. borneoe inhabits the dipterocarp forest in Sandakan and is typically found on or under the foliage of tree sapling or shrub at night. Remark. The calling song of C. borneoe differs from that of Cardiodactylus admirabilis Tan & Robillard, 2014 from Singapore by consisting of doublets with longer echeme duration (instead of triplets with shorter echeme duration of 205.6 ms) and higher dominant frequency (instead of 14.3 kHz). This is despite of the similarities in the morphology and genitalia of these two species. Another morphologically similar species is Cardiodactylus thailandia Robillard, 2011 from Thailand but its song is unknown.Published as part of Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C. & Robillard, Tony, 2019, Crickets of the subfamily Eneopterinae (Orthoptera: Grylloidea) from Sandakan, Sabah: one new species and calling songs of a sympatric species, pp. 347-363 in Zootaxa 4619 (2) on pages 355-362, DOI: 10.11646/zootaxa.4619.2.9, http://zenodo.org/record/324967
Cardiodactylus lombrinjani Robillard & Gorochov & Poulain & Suhardjono 2014, n. sp.
No full textCardiodactylus lombrinjani Robillard, n. sp. (Figs 4, 6A, 10A, 11G, 13K, 15A, 38, 39, 40) Type material. Male holotype: Indonesia. West Nusa Tenggara Province, Lombok, Joben district, Rinjani National Park, forêt de Joben, 08°31'04.7"S 116°23'33.4"E, 870 m, 29.VI.2010, TR393, jour, litière, T. Robillard (MZB). Female allotype: Lombok, Tetebatu district, Rinjani National Park, 29.VI.2010, morte en élevage, T. Robillard (MZB). Paratypes (5♂, 1♀): Lombok, Tetebatu district, Rinjani National Park, forêt de Jeruk Manis, 08°30'39.7"S 116°25'23"E, 848 m, 29.VI.2010, T. Robillard: 1♂ (TR403), jour (5PM), sur plante, enregistrement appel, photo TR (MNHN-EO-ENSIF3673); 1♂ (TR404), nuit, sur plante en lisière (h = 80 cm), enregistrement appel (MNHN-EO-ENSIF3672). Lombok, Tetebatu district, Rinjani National Park, 08°30'S 116°25'E, 29.VI.2010, morts en élevage, T. Robillard: 1♂ (TR446) (ZIN), 2♂ (TR447,?) (MZB), 1♀ (MNHN-EO-ENSIF3674). Type locality. Indonesia, Lombok, Rinjani National Park. Other material examined. Indonesia. West Nusa Tenggara Province, Lombok, Tetebatu district, Rinjani National Park, 08°30'S 116°25'E, 29.VI.2010, 2 juveniles, morts en élevage, T. Robillard (MNHN, MZB). Lombok, Pusuk forest, 550 m, 9–13.X.1991, 1♀, Krikken, Huijbregts & de Vries (RMNH). Sumbawa, Batu Dulang, forêt au S du village, sommet 1 au-dessus du village, 08°36'56.4’’S 117°15'58.4’’E, 26–27.VI.2010 (GPS2), zone ouverte herbacée, 26–27.VI.2010, T. Robillard: 1♂ (TR375), nuit, sur plante (h = 1.3 m), enregistrement appel (MNHN-EO-ENSIF3675); 1♂ (TR376), nuit, sur plante (h = 80 cm), enregistrement appel (MNHN-EO-ENSIF3677). Batu Dulang, forêt au S du village, 08°37'S 117°15'E, 26–27.VI.2010, mort en élevage, T. Robillard, morts en élevage: 2♂ (TR390-TR391) (MZB); 1♂ (Male 1-F0), enregistrement appel en captivité (MNHN-EO-ENSIF3676); 2♂ (Male 2-F0, Male 3-F0), enregistrement appel en captivité (MNHN-EO-ENSIF3678- 3679); 1♀ (MZB); 3 juveniles (MZB); 2 juveniles (MNHN). Etymology. Named after the contraction of the type locality and island (Lombock and Rinjani National Park). Distribution. Indonesia, Lombok and Sumbawa Islands. Diagnosis. Species of average size, general coloration brown, little contrasted. Characterized by male pseudepiphallic dorsal ridges showing wide expansions folded innerly, close to that of C. vella Otte, 2007a and C. riga. Description. Average size for the species group. General coloration brown, little contrasted. Head dorsum yellow brown with 4 brown bands including 2 wide lateral ones, fused to the dark brown fastigium, and 2 median punctuated bands; area posterior to eyes with dark brown lines (Fig. 6A). Scapes yellow brown, with a transverse dark brown band. Antennae orange brown. Front part of fastigium yellow, sometimes with 2 dark brown spots. Lateral side of head mostly light brown. Face yellow brown, with a faint whitish transverse band ventral to eyes. Mouthparts whitish; maxillary palpi yellow brown. Pronotum: Dorsal disk mostly yellow brown mottled with dark brown; posterior area dark brown. Lateral lobes dark brown dorsally, ventral part yellow brown to whitish. Legs I–II orange brown, femora with faint dark brown spots and tibiae with rings. FIII homogeneously yellow to orange brown, knees dark brown; TIII dark brown, with faint yellow brown rings. Tarsomeres III-1 dark brown. Hind wing tail dark brown. Cerci yellow brown, faintly mottled with dark brown near base, ringed at apex. Abdomen mostly yellow brown dorsally; lateral and dorsal sides dark brown with discontinuous yellow lines on posterior tergites. Subgenital plate yellow brown. Male: FWs brown with yellow and dark brown patterns (Fig. 10A). Yellow brown to whitish areas including bases of anal veins and of CuA, harp veins, part of chords and a wide band posterior to mirror. Black spot near base of 1A and CuA. Bases of M, R and Sc yellow brown, then dark orange brown, including veins and areas between them; apex of Sc whitish. Projections of Sc and more ventral veins of lateral field yellow brown, their bases brown. FW venation (Fig. 10A): 1A clearly bisinuated. CuP absent. Stridulatory file with about 240 teeth on the transverse and curved parts of 1A, and 21 more teeth on top of a bump near 1A base (Table 9). Harp with 2 w-shaped veins. Mirror area: mirror (d1) longer than wide, its posterior region variably crossed by longitudinal and transverse accessory veins; d2 as wide as mirror, variable. Cell e1 crossed at mid-length by an accessory vein. Apical field with 3–4 cell alignments posterior to mirror (m = 4, n = 4). Lateral field with 8–10 projections of Sc (m = 9; n = 4) and 4–5 ventral veins (m = 4; n = 4). Male genitalia (Fig. 38E–H): Pseudepiphallus shaped as a wide gutter at the level of the dorsal ridges, almost closed dorsally by wide inner expansions of the ridges touching each other posteriorly (Fig. 38H). Pseudepiphallic sclerite with large triangular anterior expansions, the membrane between them setose. Posterior pseudepiphallic apex rounded, with a long oval translucent area. Rami with short convergent apical stems. Ectophallic arc complete, without posterior expansion. Ectophallic fold with weak lateral sclerites; apex trilobate, membranous, median lobe oval. Endophallic sclerite long, with a rectangular posterior expansion and short lateral arms. Endophallic apodeme with a dorsal crest and wide lateral lamellas. Membrane of endophallic cavity finely plicate. Female: FW coloration dark brown (cells and veins), their bases yellow, and with a whitish area on external edge at mid-length. CuA/M area whitish on basal part. R orange brown. Sc dark brown, apex whitish, Sc projections and more ventral veins yellow, the cells between them dark brown. FW venation (Fig. 11G): dorsal field with 12 (n = 2) strong longitudinal veins; lateral field with 10–11 (n = 2) longitudinal veins including 7 projections of Sc and 3–4 more ventral veins. Ovipositor of average length, apex with both dorsal and ventral edges denticulate (Fig. 13K). Female genitalia (Fig. 15A): Copulatory papilla triangular with baso-lateral sclerotizations; apex rounded, slightly sclerotized and folded ventrally. Juvenile: Brown, little contrasted, head and legs with same pattern of coloration as adults (Fig. 39D). Variation. One female specimen from Pusuk forest may belong to another species according to its larger size (Table 9). The specimens from Sumbawa Island are slightly smaller than that from Lombok (type series), but they differ only by details of male genitalia. In particular the region posterior to the pseudepiphallic dorsal ridges seems smaller and the inner triangular expansions of the dorsal ridges are slightly stockier, and the endophallic sclerite tends to be smaller. However, these differences are not stable among the specimens of each island, so only one species is distinguished here. Measurements. See Table 9. Habitat and life history traits. Cardiodactylus lombrinjani is a nocturnal species living both in forested areas and in/or near secondary habitats such as bushes and hedges. Males produce calling songs at night, from tree branches and bushes (Fig. 39). During the day, couples have been observed mating under the bark of trees and several specimens have been found in blocks of aerial litter. Juveniles are found day and night on low vegetation. Behavior. Calling song (Fig. 40): Cardiodactylus lombrinjani has a mono-syllabic calling song. At 23°C, the calling songs show a very indented amplitude profile. Syllables have the following characteristics: syllable duration = 193 ± 12 ms; syllable period = 18.2 ± 11 s; syllable duty cycle = 1.6 %. The power spectrum consists of a rather broad band dominating at 13.94 ± 0.37 kHz which corresponds to the third frequency peak of the song despite harmonic pattern is unclear.Published as part of Robillard, Tony, Gorochov, Andrej V., Poulain, Simon & Suhardjono, Yayuk R., 2014, Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species, pp. 1-104 in Zootaxa 3854 (1) on pages 63-67, DOI: 10.11646/zootaxa.3854.1.1, http://zenodo.org/record/492933
Agnotecous minoris Robillard, n. sp.
No full textAgnotecous minoris Robillard n. sp. (Fig. 4) Type material. Holotype male: New Caledonia. Grande Terre, Base du Mont Mou, forêt sur pente 22 °04’ 30 ’’S 165 ° 19 ’ 52 ’’E, 390 m (Pt 29), (TR 266), 16.V. 2008, jour, litière, T. Robillard & F. Muller (MNHN-ENSIF 1389). Type locality. New Caledonia, Grande Terre, Base du Mont Mou. Etymology. Named after the small size of the species. Distribution. New Caledonia, Grande Terre, Province Sud, Mont Mou. Diagnosis. Species small, close to A. brachypterus from which it differs by its smaller size, details of colouration and by the shape of the pseudepiphallus in male genitalia. Description. Size very small for the genus. Colouration contrasted, quite dark with lighter areas on dorsum. Head dorsum with 6 brown longitudinal bands without yellow brown background; dorsal eye margins yellow. Fastigium yellow brown with median black spots. Scapes yellow brown with dark brown patterns anteriorly, antennae light brown basally then dark brown with yellow rings. Cheeks black dorsally, dark brown ventrally, with a whitish spot bellow eyes. Face homogeneously dark brown, the dorsal limit with fastigium black. Mouthparts yellow mottled with black, brown and whitish. Palpi black with a yellow brown ring on the 5 th segment. Pronotum: Dorsal disk cream except posterior margin dark brown. Lateral lobes black. FI, FII black with few yellow brown spots; knees yellow brown. TI, TII black with a faint yellowish ring; tarsomeres yellow brown, their distal part black. Legs III almost homogeneously brown. TIII serrulation: 5 inner and 9 outer spines above spurs; 5 inner and 7 outer spines between spurs. Tarsomeres III- 1: 1 spine on dorsal inner edges, 3 spines on dorsal outer edges; 1 outer lateral spine. Abdomen homogeneously dark brown dorsally; lateral parts of tergites mottled with yellow brown; sternites homogeneously brown. Cerci yellow brown with dark brown spots. Male: FW not reaching abdomen mid length. Dorsal field shorter than lateral field. FW colouration: Cells and veins homogeneously brown, a yellow brown sclerotized area around 1 A and posterior to the plectrum; lateral field with orange longitudinal veins and dark brown transverse veins; few whitish cells between R and its bifurcations. FW venation (Fig. 4 A): CuP absent, harp triangular, flat, without complete transverse vein; c 1 cell wide, not crossed by a transverse vein; D alignment and mirror (d 1) well differentiated, but mirror not rounded. Apical field restricted to a few cells in E alignment (e 1 -e 3) and one cell in F alignment. Lateral field: latero-dorsal angle made by MP; distal part of MP weak, fused to MA; R bifurcated twice, strong until distal third, then faint and fused apically with MA; ventral part of lateral field with 6 longitudinal veins. Male genitalia (Figs 4 B, C): Close to that of A. brachypterus Pseudepiphallus not widened preapically, apex with a short, flat median process. Pseudepiphallic lophi forming lateral crests ventrally. Female: unkwown. Juvenile: unkwown. Measurements. See Table 4. Habitat and life history traits. A. minoris lives in forested areas in the leaf litter. Behaviour. Mating behaviour not observed. Calling song. unknown. PronL PronW FWL FWW FIIIL FIIIW TIIIL OL Male holotype 2.7 3.9 3.3 2.9 10.4 3.6 8.9 -Published as part of Robillard, Tony, Nattier, Romain & Desutter-Grandcolas, Laure, 2010, New species of the New Caledonian endemic genus Agnotecous (Orthoptera, Grylloidea, Eneopterinae, Lebinthini), pp. 17-35 in Zootaxa 2559 on pages 27-28, DOI: 10.5281/zenodo.19699
Lebinthus villemantae Robillard, 2010, n. sp.
No full textLebinthus villemantae n. sp. (Figs 1 G, 3 G, 4 G, 6 D, 7 D, 8 J–L, 10 D–F, 11, 12, 13) Type material. Holotype male: Indonesia: Province Sulawesi Selatan: Sulawesi Selatan, Mont Bulu Saraung, Pos 5, forêt de pente, espace dégagé en bord de piste, litière, 20 -VIII- 2007, jour (T. Robillard), adulte en élevage (MZB Orth). Allotype female: same locality as HT, 31 -VIII- 2007, jour (C. Villemant), adulte en élevage (MZB). Paratypes (183, 5Ƥ): Indonesia: Province Sulawesi Selatan: same locality, date and collector as AT, 73, 2Ƥ, adultes en élevage (MZB); 93 (recorded), 2 Ƥ, adultes en élevage (MNHN-ENSIF 2572-2582). Sulawesi Selatan, région de Malawa, forêt humide au-dessus de Boto Siri, sentier longeant ruisseau, litière, 25 -VIII- 2007, jour (T. Robillard), 13, 1Ƥ (MZB); 13 (MNHN-ENSIF 2583). Type locality. Indonesia, Sulawesi Selatan, Mount Bulu Saraung. Etymology. Species dedicated to the French hymenopterologist Dr. Claire Villemant (MNHN). Other material examined. Indonesia: Province Sulawesi Selatan: Sulawesi Selatan, Mont Bulu Saraung, Pos 5, forêt de pente, espace dégagé en bord de piste, litière, 20 -VIII- 2007, jour (T. Robillard), 12 juveniles (7 in alcohol) (MNHN-ENSIF 2591-2596); 30 juveniles (MZB); 31 -VIII- 2007, jour (C. Villemant), 83, 4Ƥ (MNHN-ENSIF 2567 -2571, 2584- 2590), 6 juveniles (MZB). Distribution. Indonesia, South Sulawesi. Diagnosis. Species of small size, slightly smaller and darker than Lebinthus truncatipennis, and differing from L. truncatipennis and L. makilingus by male genitalia and details of colouration. Description. Size small. Colouration variable, most often dark brown with more or less contrasted colour patterns (Fig. 11). Head dorsum with 4–6 wide dark brown longitudinal bands more or less distinct, with a yellow band posterior to eyes prolonging their colouration and underlined by black pattern. Eyes well protruding, black with a yellow longitudinal band dorsally. Fastigium wider than long, setose, brown with a dark spot behind median ocellus, yellow. Scapes yellow; antennae yellow at base then progressively dark brown. Upper part of front head yellow. Black mask including lower part of front head, region below antennae and eyes, base of mandibles and upper part of clypeus; epistomal suture yellowish brown; yellow to whitish colouration around the mask, including parts of jaws and lower parts of mandibles and clypeus. Palpi whitish with dark brown rings. Pronotum: Dorsal disk slightly trapezoidal, straight posteriorly; generally dark brown, lighter laterally. Lateral lobes dorsally black, ventral margin yellow with a black line (Fig. 3 G). Legs: Fore and mid legs light brown, femora with dark brown spots, tibiae with dark rings. Hind femora brown, with striated dark patterns on outer faces, 3–5 black spots on each ventral edge, knees dark brown; hind tibiae with black rings, distal half of tarsomeres III- 1 black. Hind tibiae with 6–12 inner (m = 7.9, n = 10) and 10–13 outer (m = 11.3, n = 10) spines above spurs and 4–5 inner (m = 4.3, n = 10) and 4–7 outer (m = 5.4, n = 10) spines between spurs. Tarsomeres III- 1 with 3–4 spines on dorsal outer edges (m = 3.4, n = 10) and 0–2 (m = 1.3, n= 8) on outer faces. Abdomen homogeneously brown dorsally, yellowish brown ventrally with sometimes a median dark brown band. Cerci yellowish brown basally, with black rings near apex. Male: FW not reaching abdomen midlength. FW colouration: Cells and veins brown, not translucent; intermedian area whitish, except for dark brown veins; lateral field brown, its dorsal margin (MA/R area) dark brown. FW venation (Fig. 1 G): 1 A angle wide (> 100 °); stridulatory file with 198–213 teeth (m = 203, n = 4), located on transverse part of 1 A only. CuP visible at FW basis only. Harp wide, including a false mirror, i.e. a distinctive rounded area delimited by the strong harp vein, polyfurcated anteriorly, and fused with both diagonal vein and CuA posteriorly. Distal part of CuA weak and curved inwards, surrounding the median fold. Distal part of diagonal vein very weak. Longitudinal veins very strong at apex, transverse veins very weak. Mirror (d 1) not differentiated. Apical field absent, with no bifurcation of CuA posterior to diagonal vein. Lateral field with 5 strong longitudinal veins including MA, R and 3 more ventral veins; latero-dorsal angle made by MP; R without strong bifurcating veins. Subgenital plate elongate, clog-shaped (Fig. 4 G). Male genitalia (Fig. 10 D–F): Pseudepiphallic sclerite triangular, convex dorsally; posterior apex with short individualized lophi, setose, parallel and separated by a V-shaped indentation; anterior margin slightly indented, its lateral margins slightly curved dorsally. Rami short. Pseudepiphallic parameres trilobate, the posterior lobe dorsal, the two other lobes ventral, the median one rectangular. Ectophallic arc complete, narrow, very close to basis of pseudepiphallic parameres. Ectophallic fold short and wide, with a wide preapical sclerotization including a rounded part and two anterior expansions laterally. Ectophallic apodemes parallel and very long, exceeding anterior margin of pseudepiphallus. Endophallic sclerite very long, exceeding anterior margin of pseudepiphallus, dorsally convex, its posterior apex with a median triangular expansion and with short lateral arms; endophallic apodeme made of a narrow median crest. Female: FW short (Fig. 6 D), exceeding posterior margin of second tergite, not overlapping but close together; dorsal field with 5 strong longitudinal veins, brown. Lateral field darker than dorsal field, with 3–4 strong dark brown longitudinal veins, intermedian area whitish as in male. Ovipositor shorter than hind femora; apex lanceolate, denticulate on dorsal edge (Fig. 7 D). Female genitalia: Copulatory papilla (Fig. 8 J–L) with small basal sclerotized ring; apical part narrowed and long, slightly plicate ventrally and sclerotized at apex. Juvenile: Similar to adults in colouration, dark brown (Fig. 12). Measurements. see Table 1. Habitat and life history traits. L. villemantae lives in the leaf litter of forested areas (Fig. 12). It is found in dense populations in wet leaf litter, generally on or under large dead leaves. Behaviour. Only 3 adult specimens were observed in the field, by day in leaf litter in Malawa region, South Sulawesi. Singing activity was recorded in laboratory mostly during night period, but sexual activity seems to occur during both day and night. Mating behaviour has been observed in laboratory; multiple copulations have been recorded and will be described in a forthcoming paper (Robillard et al. in prep.). Calling song (Fig. 13; Table 2): Five males recorded in laboratory at 18–26 °C are analysed here. The calling song consists of a succession of short echemes of 1.8 ± 0.1 s emitted every 47.1 ± 19 s, with a duty cycle of 3.8 %. At 20 °C each echeme is made of 99.5 ± 10.6 syllables, with the following characteristics: syllable rate = 58.7 ± 3.8 /s; syllable duration = 7.2 ± 1.9 ms; syllable period = 17.5 ± 4.1 ms; syllable duty cycle = 41.1 %; the dominant frequency is 20.1 ± 2.3 kHz and corresponds to the fundamental frequency of the song. Courtship song: Less intense and shorter than calling song. Aggressive song: Not observed despite many observations of male interactions. This song is probably absent from the species repertoire. Specimens T°C Echeme Syllable The field work in South Sulawesi was organised by Louis Deharveng (MNHN), Anne Bedos (MNHN) and Yayuk Suhardjono (LIPI), with a grant from the PPF "État et structure phylogénétique de la biodiversité actuelle et fossile", MNHN (P. Janvier). I thank Claire Villemant (MNHN) for helping collecting crickets in Sulawesi. I also thank Simon Poulain (CNRS) for preparation of the specimens, Constance Boitard for specimen measurements, Gilbert Hodebert (MNHN) for habitus drawings and Laure Desutter-Grandcolas for helpful comments on the manuscript. I also thank: Judith Marshall and George Beccaloni (BMNH), for their help during the study of Eneopterinae crickets in the Natural History Museum, London, funded by the SYNTHESYS European program (GB-TAF- 531); Rob de Vries and Caroline Pepermans (RMNH) for their help during the study in Leiden collections, funded by a grant from the PPF "État et structure phylogénétique de la biodiversité actuelle et fossile", MNHN (P. Janvier).Published as part of Robillard, Tony, 2010, New species of the genus Lebinthus (Orthoptera, Grylloidea, Eneopterinae, Lebinthini) from Indonesia and the Solomon Islands, pp. 25-48 in Zootaxa 2386 on pages 43-47, DOI: 10.5281/zenodo.19375
Xenogryllus mozambicus Jaiswara & Dong & Ma & Yin & Robillard 2019, n. sp.
No full textXenogryllus mozambicus Robillard n. sp. (Figs 2 A–D, 3K–L, 4H, 5F, 7J–K, 8F, 9D, 11H–J, 12C, 15, 16) Xenogryllus eniopteroides (wrong spelling of X. eneopteroides)—Toms 1984: 309. Type material. Holotype, ♂, Mozambique: Cabo Delgado, Pantanos de Nhica, zone herbacée à l’Est du camp [herbaceous area E of camp], 10°45’19,1"S 40°13’00"E, 122 m, 29.xi.2009, T. Robillard, nuit, enregistrement appel [call recording MNHN-SO-2018-140] (TR475) (MNHN-EO-ENSIF1517). Allotype, ♀, Mozambique: Cabo Delgado, mare SE de Nhica, bras mort de la Rovuma [pond SE Nhica, dead arm of Rovuma river], 10°45’41,9"S 40°13’31,7"E, 124 m, 27.xi.2009, T. Robillard, mort en élevage [dead in captivity] (MNHN-EO- ENSIF1539). Paratypes (8♂, 4♀): Mozambique: Cabo Delgado, same information as holotype: 1♂, 1♀, morts en élevage [dead in captivity] (MNHN-EO-ENSIF1576-ENSIF1556). Same information as allotype: 2♂, mort en élevage [dead in captivity] (MNHN-EO- ENSIF1557, ENSIF1508). Mare SE de Nhica, bras mort de la Rovuma [pond SE Nhica, dead arm of Rovuma river], 10°45’41,9"S 40°13’31,7"E, 124 m, nuit, zone herbacée en bord de piste [herbaceous area near track] T. Robillard: 22.xi.2009, 1♂ (MNHN-EO-ENSIF1554), 3♀ (MNHN-EO- ENSIF1502, ENSIF1503, ENSIF1505); 23.xi.2009, 1♂ (TR208), enregistrement appel Take 224 [call recording MNHN-SO-2018-133] (MNHN-EO-ENSIF3079); 1♂ (TR206), enregistrement appel Take 222 [call recording MNHN-SO-2018-134], molecular sample X7 - XenMoz (MNHN-EO-ENSIF1515). Mare SW de Nhica, bras mort de la Rovuma [pond SW Nhica, dead arm of Rovuma river], 10°52’09,5"S 40°06’47,1"E, 122 m, 24.xi.2009, nuit, zone herbacée en bord de piste [herbaceous area near track] T. Robillard: 1♂ (TR224), hautes herbes (h=1.3 m) [on high grass], enregistrement appel Take 228 [call recording MNHN-SO-2018-135], molecular sample X23 (MNHN-EO-ENSIF1559); 1♂, mort en élevage (MNHN-EO-ENSIF1581). Additional material examined. Mozambique: Delagoa [Maputo Bay], xii.1898, 1 ♂, identified Phormincter species nova by A. Finot (MNHN). South Africa: Zululand, Mtunzini, 8.i.1952, R. Toms, 1♂, R146, identified Xenogryllus eneopteroides by B. C. Townsend, 1984, B.M.1983-166 (NHMUK 010926568). Malawi: Fish Eagle Bay Lodge, herbaceous area near lake Malawi (Mal 7), S13°02'21.1" E34°19'34.8", 503 m, 6.x.2018, night, 2♂, call recording, 1♀, T. Robillard, K. Salazar & R. Murphy (MNHN). Type locality. Mozambique, Cabo Delgado, Pantanos of Nhica, 10°45’19,1"S 40°13’00"E. Distribution. Mozambique, South Africa, Malawi. Etymology. Species named after the type locality. Diagnosis. Species of average to large size, close to X. eneopteroides and X. maniema n. sp. From X. eneopteroides, the new species differs by rather larger size, face more rounded, almost globular in lateral view, absence of T-shaped median band on pronotum (also absent in X. maniema), larger mirror, and slight differences in male genitalia. From X. maniema n. sp., X. mozambicus mostly differs by male genitalia with larger lophi (Fig. 8F) and presence of transverse carina on ventral face of lophi (also present in X. eneopteroides). As X. maniema and X. eneopteroides, X. mozambicus differs from X. lamottei n. sp. and Asian species by strongly carinated lateral angles of pronotum. Description. Species of average size (Fig. 2 A–D), coloration light brown little contrasted. Eyes little prominent laterally (Fig. 3K), higher than long, occupying almost half of head height in lateral view. Face globular in lateral view (Fig. 3L), with typical whitish mask underlined by a black line below eyes and on mandibles. Pronotum dorsal disc strongly carinated laterally, coloration light brown, with a median dark brown band, not extended laterally near anterior margin; lateral lobes almost homogeneously dark brown. First article of antennae dark brown. Male. FWs very wide, longer than abdomen; dark coloration anterior to 1A including angle of 1A (Fig. 5F). Stridulatory file with 505 teeth (n=1) on transverse part of 1A. FW venation as in X. eneopteroides, mirror wider and apical field longer, forming a long triangle including 5–6 (n=8) cell alignments. Male genitalia (Fig. 7 J–K): Similar to X. eneopteroides, except longer pseudepiphallic lophi (Fig 8F); transverse carina on ventral face of lophi present but weak. Female (Fig. 2 C–D) FW dorsal field elongate, with 8–9 longitudinal veins (m = 9, n = 6). Subgenital plate with a shallow apical indentation (Fig. 9D). Ovipositor short, about 0.6 times FIII length. Female genitalia: Copulatory papilla (Fig. 11 H–J) as in X. eneopteroides, conical and narrow, well-sclerotized except base and apex. Life history traits. X. mozambicus lives in areas of wet savannah (Fig. 15) or on the edge ponds and lakes. Individuals of both sexes are found only at night in herbaceous vegetation, and males sing at night from the vegetation (ca. 30–80 cm high). Toms (1984) demonstrated that this species has directional calls and turs while calling, giving rise to changes in sound intensity. Calling song (Figs 12C, 16). At 21.5 °C, males of X. mozambicus emit almost continuously long bouts of highly musical calling songs. After a warming phase, call bouts are made of successions of echemes of 23 ± 2 syllables, barely separated from each other (echeme duration = 3.13 ± 0.29 s; echeme period 3.21 ± 0.30 s, echeme duty cycle = 97.6 %), with a regular amplitude profile, except for the last syllable, which is more intense. Within echemes, syllables are initially organized in 4 ± 1 doublets, which are followed by a series of similar syllables. Syllables are very long (syllable duration = 126 ± 5 ms), with a relatively short syllable period of 137 ± 8 ms (syllable duty cycle = 92%). All syllables are characterized by a large amplitude modulation: in initial doublets within echemes, the first syllables are less intense than the second ones and they start with a lower amplitude than their end (initial amplitude ending amplitude); the rest of the syllables have a higher starting amplitude, except for the last syllable of each echeme, which has a higher ending amplitude. The frequency spectrum shows a pure-tone dominant frequency at 3.3 ± 0.01 kHz, followed by a rich harmonic content including three powerful harmonics at ca. 6.6, 9.9, 13.2 kHz, especially visible in ending syllables. Measurements. See Table 7. Taxonomic discussion. The specimens observed by Toms (1984) and identified Xenogryllus eniopteroides (wrong spelling for X. eneopteroides) from Zululand (Mtunzini and Eastern Transvaal (Clanor)), South Africa, belong to X. mozambicus n. sp. according to one male specimen observed from the series of Toms (NHMUK 010926568).Published as part of Jaiswara, Ranjana, Dong, Jiajia, Ma, Libin, Yin, Haisheng & Robillard, Tony, 2019, Taxonomic revision of the genus Xenogryllus Bolívar, 1890 (Orthoptera, Gryllidae, Eneopterinae, Xenogryllini), pp. 301-338 in Zootaxa 4545 (3) on pages 328-331, DOI: 10.11646/zootaxa.4545.3.1, http://zenodo.org/record/261887
FIG. 18. — Lebinthus santoensis n in Eneopterinae crickets (Insecta, Orthoptera, Grylloidea) from Vanuatu
No full textFIG. 18. — Lebinthus santoensis n. sp.: A, C, E, in Matantas, pointe de Quirros, Espiritu Santo, Vanuatu, by day; A, habitat; C, habitat close up; E, male; B, D, in Big Bay, Vatthé, Espiritu Santo, Vanuatu,by day, along beach; B, habitat; D, habitat close up with juveniles and adults in leaf litter; F, female juvenile, in Peavot, Espiritu Santo, Vanuatu, by day. Photos: S. Hugel (A, C, E, SH-19X06a; F, SH-20x06a) and T. Robillard (B, TR35; D, TR65).Published as part of Robillard, Tony, 2009, Eneopterinae crickets (Insecta, Orthoptera, Grylloidea) from Vanuatu, pp. 577-618 in Zoosystema 31 (3) on page 614, DOI: 10.5252/z2009n3a11, http://zenodo.org/record/539817
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