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    Pseudolebinthus africanus Robillard 2006

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    Pseudolebinthus africanus Robillard, 2006 (Fig 5C) Pseudolebinthus africanus Robillard, 2006: 677 —Cigliano et al. 2018 (Orthoptera Species File Online) Material examined. Holotype. ZIMBABWE: ♂, Mt. Melsetter 5000 ft [1500 m], Melsetter Dist. Dept. Agric. S. Rhodesia. xi.1957, accession no. 11776 (Whellan) identified Lebinthus africanus by L. Chopard (NHMUK, Brit Mus 1982–221). Paratype. ZIMBABWE: 1♂, Chirinda Forest, Melsetter Dist. Dept. Agric. S. Rhodesia. x.1948, accession no. 11774 (Whellan) 1# identified Lebinthus africanus by L. Chopard (MNHN-EO-ENSIF10684). Type locality. Zimbabwe, Mount Melsetter 5000 ft. Distribution. Zimbabwe Diagnosis. (Emended from Robillard 2006). Coloration contrasted; FIII uniformly brown. Dorsal margin of eyes with small ommatidia thin. Male. Scutellum basal edge sharply raised. FW dorsal field with strong transverse veins. Mirror small, hardly distinct from surrounding cells and crossed by one strong accessory vein; c1 twice as long as c2. Sc with three projections along its length (four in P. whellani, two in P. gorochovi sp. nov.). Male genitalia. Pseudepiphallic lophi (Fig. 5C) longer and thinner than in P. whellani, their apical lobes almost completely fused, as in P. gorochovi sp. nov. Median part of endophallic sclerite well developed.Published as part of Jaiswara, Ranjana, Dong, Jiajia & Robillard, Tony, 2018, Revision of the genus Pseudolebinthus (Orthoptera, Gryllidae, Eneopterinae) with the description of a new species from Southeast Africa, pp. 265-274 in Zootaxa 4521 (2) on pages 267-269, DOI: 10.11646/zootaxa.4521.2.7, http://zenodo.org/record/260988

    Cardiodactylus singapura Robillard, sp. nov.

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    Cardiodactylus singapura Robillard, sp. nov. (Figs 1–5) Type material. Holotype male: Singapore. Central Catchment Nature Reserve, 01° 22 ' 58.3 ''N 103 ° 48 ' 50.3 ''E, 37 m, 4.VII. 2009, nuit, adulte en élevage, enregistrement appel (F0-male 4), T. Robillard (ZRC). Allotype female: Singapore. Bukit Timah, 01° 21 '01.0''N 103 ° 46 ' 44.3 ''E, 95 m, VII. 2009, adulte en élevage, T. Robillard (ZRC). Paratypes: Singapore. MacRitchie Reservoir, 01° 21 ' 10.2 ''N 103 ° 49 ' 32.2 ''E, 36 m, 2.VII. 2009, nuit, adultes en élevage, T. Robillard: 23, enregistrement appel (F0-males 1-2) (MNHN-ENSIF 2754-2755); 1 Ƥ (MNHN- ENSIF 2756); 1 Ƥ (ZRC). Central Catchment Nature Reserve, 01° 22 ' 58.3 ''N 103 ° 48 ' 50.3 ''E, 37 m, 4.VII. 2009, nuit, adultes en élevage, T. Robillard: 13, enregistrement appel (F0-male 3) (MNHN-ENSIF 2757); 1 Ƥ (MNHN- ENSIF 2758), 13, 1Ƥ (ZRC). Bukit Timah, 01° 21 '08.4''N 103 ° 46 ' 43.8 ''E, 116 m, VII. 2009, nuit (8: 30 PM), T. Robillard: 13, adulte en élevage, enregistrement appel (F0-male 6) (MNHN-ENSIF 2759); 13 (TR 39), sur plante de sous-bois (feuille, h= 1.5 m), 2 Ƥ, adultes en élevage, (ZRC). Singapore [no precision], 13, Coll. Baker, 14618 (MNHN-ENSIF 2753). Singapore [no precision], VII- 2009, T. Robillard, nuit, adulte en élevage, 13, enregistrement appel (F0-male 5) (MNHN-ENSIF 2760) Type locality. Singapore, Central Catchment Nature Reserve, 01° 22 ' 58.3 ''N 103 ° 48 ' 50.3 ''E, 37 m. Etymology. Named after the type locality. Other material examined. Singapore. Bukit Timah, 01° 21 '08.4''N 103 ° 46 ' 43.8 ''E, 116 m, 1.VII. 2009, nuit, T. Robillard: 1 juvenile (MNHN). Central Catchment Nature Reserve, 01° 22 ' 58.3 ''N 103 ° 48 ' 50.3 ''E, 37 m, 4.VII. 2009, nuit, T. Robillard: 1 juvenile (MNHN). Singapore [no precision], reared specimens (F 1 generation, TR- 2010): 53, 2Ƥ (MNHN). Distribution. Singapore island. Diagnosis. Species small with contrasted coloration, close to C. pelagus Otte, 2007 from Borneo, from which it differs by the shape of the pseudepiphallus in male genitalia and lighter colour pattern. Description. Size small for the species group. General coloration contrasted, light brown with dark brown patterns. Head dorsum yellow brown with 4 wide dark brown bands, more or less punctuated, bordered by dark brown triangles posterior to eyes and a yellow line more externally (Figs 1 A, 2 A). Fastigium mottled with dark brown. Scapes light brown with a transverse dark brown line anteriorly (Figs 1 A, 2 B). Antennae heterogeneously brown. Face yellowish to light brown, diversely mottled with brown, with 2 parallel lines on the fastigium, 4 dark spots on face, and 2 dark brown spots ventral to eyes. Mouthparts yellow brown. Maxillary palpi yellowish, dark brown apically. Head lateral side with a dark brown area posterior to eyes, progressively lighter ventrally. Pronotum: Dorsal disk trapezoidal, posterior margin straight or slightly bisinuated; light brown mottled with greyish brown and dark brown spots; anterior corners yellowish. Lateral lobes dark brown, ventral edges yellow. Legs light brown to yellowish brown, femora mottled with dark brown; tibiae more or less banded; tarsomeres with dark brown ends. Tarsomeres III- 1 with 2–3 spines on dorso-external edges (m= 2.2, n= 10). Hind wings longer than FWs, the dark brown tail exceeding the forewings twice longer than the pronotum. Cerci light brown with black rings. Tergites light brown mottled with black. Male: FW coloration (Figs 1 B, 2 C): Dorsal field cells dark brown, veins mostly orange brown, except for whitish areas in the chords, part of anal veins and in the region posterior to mirror. Base of chords with a brown sclerotization. Harp posterior angle with a whitish sclerotized area. Lateral field dark brown; veins MA, MP, R orange brown, transverse veins and projections of R whitish. FW venation: 1 A slightly bisinuated; stridulatory vein (Figs 2 C, 2 D) with 156–171 teeth (Fig. 2 E) (m= 164, n= 4), distributed on both transverse (123–143 teeth, m= 133) and longitudinal (15–20 teeth, m= 19) parts of 1 A, plus a group of 9–16 teeth near base of 1 A (m= 13, n= 4). CuP missing. Harp longer than wide, with 2 w-shaped harp veins; posterior margin raised along diagonal vein. Mirror area variable, c 1 long and narrow, c 2 quite large; mirror (d 1) longer than wide, not rounded, generally separated in two parts by a transverse vein, the posterior part triangular; e 1 cell very elongated, along the mirror. Apical field long and triangular, with 3 wide cell alignments posterior to mirror and a narrow apical alignment. Lateral field: R with 6–7 projections (m= 6, n= 5). Subgenital plate yellow brown, the median area dark brown. Male genitalia (Fig. 3): Pseudepiphallus very sclerotized, little setose, slightly narrowed at mid-length. Dorsal ridges parallel, forming a wide gutter, flattened anteriorly, their external edges carenated. Pseudepiphallic sclerite with latero-anterior expansions, but without a membranous sac as in C. novaeguineae; membrane between the expansions with short strong setae. Apex of pseudepiphallus spoon-like, not rounded, and slightly indented. Rami narrow, with wide preapical plates. Pseudepiphallic parameres trilobate, the posterior lobe wide. Ectophallic arc complete, with a short posterior expansion. Ectophallic apodemes thin, divergent. Apex of ectophallic fold almost hidden by pseudepiphallic parameres, trilobate, membranous. Base of ectophallic apodemes with short sclerotized expansions oriented posteriorly. Endophallic sclerite small. Endophallic apodeme with both lateral lamellas and a short medio-dorsal crest. Membrane of endophallic cavity smooth. Female: FW coloration: cells dark brown, veins orange brown, with whitish sclerotization on distal half of CuA and MP, including cells between CuA and MA. MA and R orange brown, R bifurcations whitish. Anterior part of FW with variable number of whitish transverse veins. FW venation (Fig. 4 A): 9–11 (m= 10, n= 5) strong longitudinal veins on dorsal field; lateral field with 9–11 (m= 9.5, n= 5) longitudinal veins including 5–6 R bifurcations and 3–4 ventral veins. Ovipositor: Shorter than hind femora; apex with both dorsal and ventral edges denticulate (Fig. 4 E). Female genitalia (Fig. 4 B–D): Copulatory papilla triangular with baso-lateral sclerotizations; apex rounded and folded ventrally. Juvenile: First instar after hatching observed in laboratory, similar in coloration to that of C. novaeguineae, yellow with brown longitudinal bands. Later instars with more contrasted coloration, body black, legs and head yellow brown (Fig. 1 C– 2 C). Black coloration reverses in sub-adults, characterized by a more homogeneous brown coloration (Fig. 1 C–D). Measurements. See Table 1. PronL PronW FWL FWW HWT FIIIL FIIIW TIIIL Male holotype 2.4 4 14.1 4.3 4.2 14 3.7 11.5 Males (n= 5) 2.2–2.4 3.7–4.2 13.3–14.1 4.1–4.3 4.2–5.4 13.4–15 3.6–4.1 11.5–13.1 (Mean) (2.4) (4) (13.7) (4.2) (4.9) (14.3) (3.9) (12.5) Female allotype 2.5 4.3 14.5 4.2 5.5 15.5 3.9 13.4 Females (n= 5) 2.2–2.7 4.1–4.5 13.5–15.2 3.9–4.4 4.5–5.5 14.1–15.5 3.9–4.4 12.5–13.5 (Mean) (2.4 (4.3) (14.3) (4.1) (5.1) (14.9) (4.1) (13) continued. TIIIs TaIIIs ST OL Ias Ibs Oas Obs Tt Lt Bt Male holotype 8 6 11 7 3 ??? - Males (n= 5) 8–10 6 11 – 13 7 2–3 - (Mean) (8.8) (6) (11.8) (7) 2.2 - Female allotype 6 6 11 7 3 - - - 10.9 Females (n= 5) 6–10 6–7 11 – 13 5–7 2–3 - - - 9.7–10.9 (Mean) (8) (6.4) (11.8) (6.4) (2.2) - - - (10.2) Habitat and life history traits. Cardiodactylus singapura is a nocturnal species living in the forested areas. Males produce their discreet calling songs at night, from low branches and leaves (Fig. 1). Juveniles are found day and night in the leaf litter and on leaves of low plants. Behavior. Calling song (Fig. 5): High speed video observations at 1250 frames per second confirm that each calling song corresponds to only one FW closure, the gaps within the syllable being made by the jerking movement of the FWs. At 26 °C, this mono-syllabic calling song (TR-male 1 -F0male: MNHN-ENSIF 2754) lasts for 87.0 ± 11.8 ms with a period of 8.2 ± 2.2 s and a duty cycle of 1.06 %. The dominant frequency is 19.0 ± 0.7 kHz and corresponds to the third frequency peak of the song.Published as part of Robillard, Tony, 2011, New Cardiodactylus species from unsuspected places in Southeast Asia (Orthoptera, Grylloidea, Eneopterinae), pp. 14-26 in Zootaxa 2909 on pages 16-22, DOI: 10.5281/zenodo.20145

    Lebinthus estrellae Robillard 2015

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    Lebinthus estrellae Robillard, 2015 (Fig. 1) Lebinthus estrellae Robillard, 2015, in Robillard & Yap, 2015: 63 (original description); Baroga et al., 2016: 94 (taxonomic key). Type material. Holotype male. Philippines. [Visayas] Leyte: Burauen, Barangay Villa Corazon, forêt secondaire sur pente [secondary forest on slope] (GPS Bar2), 10°57’52.1”N 124°46’39.8”E, 345 m, (TR29), iii.2013, coll. T. Robillard (UPLBMNH) [examined]. Paratypes (2♂, 15♀): same information as holotype (MNHN) [examined]. Type locality. Leyte: Philippines. Burauen, Barangay Villa Corazon, secondary forest on slope Distribution. Philippines: Leyte, Barauen Diagnosis. This species can be recognized from other species of the sanchezi group by its dark brown colouration, short rounded lophi in male genitalia, presence of vertical whitish bands below eyes, clypeus black with yellow edge; male genitalia with short rounded lophi, and very short female FWs (reaching posterior margin of first tergite). Habitat. Found in bushes and leaf litter.Published as part of Baroga-Barbecho, Jessica B., Tan, Ming Kai, Yap, Sheryl A. & Robillard, Tony, 2020, Taxonomic study of Lebinthus Stål, 1877 (Orthoptera: Gryllidae: Eneopterinae) with description of six new species in the Philippines, pp. 401-438 in Zootaxa 4816 (4) on page 415, DOI: 10.11646/zootaxa.4816.4.1, http://zenodo.org/record/395452

    Cardiodactylus thailandia Robillard 2011

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    Cardiodactylus thailandia Robillard, 2011 Cardiodactylus thailandia Robillard, 2011: 22; Eades et al. 2013 (Orthoptera Species File Online). Type material. Male holotype: Thailand. Khaoyai [Khao Yai] National Park, about 14.VII.1986 (M. Takeda) (OMNH) [examined]. Type locality. Thailand, Khao Yai National Park. New signalizations: Thailand. Trat Province, eastern Thailand, Chang I. in Siam Bay, low mountains near sea, primary forest, 5–20.XI.2000, night, A. Gorochov and L. Anisyutkin: 1♂, 1♂ juvenile (ZIN). Cambodia. 10–15 km NEE of Sihanoukville (City near Siam Bay), environs of waterfall, 200 m, secondary forest, 11–12.IX.2003, night, A. Gorochov: 1♂ (adult in captivity), 1♀ juvenile (ZIN). Distribution. Thailand, Cambodia. Diagnosis. Species of average size for the genus, with very contrasted black and vivid yellow coloration, close to C. borneoe n. sp. from Borneo, C. admirabilis from Singapore, C. guttulus from Ryukyu Islands (Japan), and C. kondoi from the Philippines, with differences on color pattern and the shape of the pseudepiphallic sclerite in male genitalia.Published as part of Robillard, Tony, Gorochov, Andrej V., Poulain, Simon & Suhardjono, Yayuk R., 2014, Revision of the cricket genus Cardiodactylus (Orthoptera, Eneopterinae, Lebinthini): the species from both sides of the Wallace line, with description of 25 new species, pp. 1-104 in Zootaxa 3854 (1) on page 100, DOI: 10.11646/zootaxa.3854.1.1, http://zenodo.org/record/492933

    Lebinthus puyos Robillard 2013

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    Lebinthus puyos Robillard, 2013 (Figs. 1, 6, 7) Lebinthus puyos Robillard, 2013: 49 (original description); Baroga et al. 2016: 94 (taxonomic key). Type material. Holotype male. Philippines. Luzon: Laguna, Los Baños, Mount Makiling, base, secondary forest on campus, 14°09’12.9”N 121°14’05.0”E, 168 m (GPS Maki1), 27.vi.2011, night, leaf litter (TR652), coll. T. Robillard (UPLBMNH) [examined]; Allotype female: same information as holotype, TR653 (UPLBMNH) [examined]; Paratypes (5♂): same information as holotype (MNHN) [examined]. Other material examined. Philippines. [Luzon]: Laguna: College, 50 m, 1♂ (JBB318), 20.ii.1958, coll. E.S. Novero (UPLBMNH ORT-01161); UPLB, Mt. Makiling, 1♀ (JBB106), 06.viii.2014, coll. J.B. Baroga (UPLBMNH); Los Baños, 300 m, 1♀ (JBB349), 21.iv.1960, coll. A. Djamin (UPLBMNH ORT-01192); [Quezon Province], Mt. Banahaw, 1♂ (JBB297), 2.vii.1994, coll. O. Reyes (UPLBMNH ORT-01140); same information as holotype, 6♂, lab colony, call recording (F0-male1, F0-male1-3, F1-male1-2, F5-male1) (MNHN). Type locality. Philippines. Luzon: Laguna, Los Baños, Mount Makiling, base, secondary forest on campus. Distribution. Philippines: Laguna: Mt. Makiling and Los Baños; Quezon: Mt. Banahaw (new record). Diagnosis. This species differs from L. sanchezi by its whitish face, a fastigium that is not orange apically, without clear longitudinal bands in the vertex (Robillard et al., 2013), and male genitalia with long and rounded pseudepiphallic lophi and C-shaped pseudepiphallic parameres. Calling song. The song of L. puyos consists of long trills (duration = 3.1± 0.8 s [1.7– 4.0 s]; period = 25.6± 2.8 s [22.4– 33.1 s]) made up of more than 200 syllables of increasing intensity (Figs. 7 A–C). Syllable duration = 12.6±1.2 ms (10–15 ms). The spectrogram analysis reveals that the frequency spectrum is completely ultrasonic. The syllables show a clear dominant peak at 23.12±1.06 kHz typically corresponding to the first peak of a harmonic series with an intermediary peak at ca. 25 to 30 kHz (Figs. 7D, 7E). Habitat. The species is found in secondary habitats in the type locality, i.e., foot of Mt. Makiling; specimens were found in leaf litter and on top of leaves of small plants (Fig. 6A).Published as part of Baroga-Barbecho, Jessica B., Tan, Ming Kai, Yap, Sheryl A. & Robillard, Tony, 2020, Taxonomic study of Lebinthus Stål, 1877 (Orthoptera: Gryllidae: Eneopterinae) with description of six new species in the Philippines, pp. 401-438 in Zootaxa 4816 (4) on page 412, DOI: 10.11646/zootaxa.4816.4.1, http://zenodo.org/record/395452

    Nécrologie. M. Eugène de Robillard de Beaurepaire

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    Travers Émile. Nécrologie. M. Eugène de Robillard de Beaurepaire. In: Bulletin Monumental, tome 65, année 1901. pp. 478-491

    Cardiodactylus borneoe Robillard & Gorochov 2014

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    Cardiodactylus borneoe Robillard & Gorochov, 2014 (Figs. 11, 12) Robillard et al., 2014: 12; Dong et al., 2018: 6 (molecular phylogeny); Cigliano et al., 2019 (Orthoptera Species File). Material examined: Malaysia, Sabah, Sandakan, Rainforest Discovery Centre, 1 female (SDK.19.6), N5.87410, E117.94366, 50.2± 5.1 m.a.s.l., 8 January 2019, 2102 hours, on foliage, coll. M. K. Tan & S. T. Toh (ZRC); 1 female (SDK.19.21, Fig. 11A) N5.87542, E117.94231, 31.1± 4.8 m.a.s.l., 9 January 2019, 1930 hours, on foliage, coll. M. K. Tan, R. Japir, Momin Binti & J. Y. Lee (FRC); 1 male (SDK.19.36, Figs. 9 B–9D), N5.87547, E117.94110, 34.1± 5.4 m.a.s.l., 9 January 2019, 2046 hours, under foliage calling, coll. M. K. Tan, R. Japir, Momin Binti & J. Y. Lee (ZRC). Calling song (Fig. 12): At 28.4°C and RH 74%, the calling song of C. borneoe consists of doublets (or rarely triplets) of long syllables including strong amplitude modulation forming pulse-like structures. Echemes (doublet) last 660 ± 75 ms with a period of 3.5 ± 1.2 s. Within doublets, syllables last 196 ± 43 ms, with a period of 372 ± 47 ms. Dominant frequency value is 17.6 ± 0.4 kHz with no clear harmonic structure. Habitat and life history traits. C. borneoe inhabits the dipterocarp forest in Sandakan and is typically found on or under the foliage of tree sapling or shrub at night. Remark. The calling song of C. borneoe differs from that of Cardiodactylus admirabilis Tan & Robillard, 2014 from Singapore by consisting of doublets with longer echeme duration (instead of triplets with shorter echeme duration of 205.6 ms) and higher dominant frequency (instead of 14.3 kHz). This is despite of the similarities in the morphology and genitalia of these two species. Another morphologically similar species is Cardiodactylus thailandia Robillard, 2011 from Thailand but its song is unknown.Published as part of Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C. & Robillard, Tony, 2019, Crickets of the subfamily Eneopterinae (Orthoptera: Grylloidea) from Sandakan, Sabah: one new species and calling songs of a sympatric species, pp. 347-363 in Zootaxa 4619 (2) on pages 355-362, DOI: 10.11646/zootaxa.4619.2.9, http://zenodo.org/record/324967

    Cardiodactylus admirabilis Tan & Robillard, n. sp.

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    Cardiodactylus admirabilis, Tan & Robillard, n. sp. (Figs. 1 A, 2–7) Cardiodactylus singapura Robillard, 2011 —Tan et al. 2012: 66 (juvenile specimens wrongly identified). Material examined. Holotype (male): Singapore, Admiralty Park (Nature Park), secondary forest near mangroves, N01.44639, E 103.77930, 11.2 ± 5.4 m, on Dillenia leaf, 25 June 2013, 2017 hours, coll. H. Yeo & M. K. Tan (ZRC.ORT. 1020). Paratypes. Singapore, same locality as holotype, secondary forest near mangroves, 3 males, coll. H. Yeo & M. K. Tan: (1) N01.44637, E 103.77926, 27.4 ± 5.6 m, on Dillenia leaf, 15 June 2013, 2120 hours, collected as the final instar, moulted into adult on 16 June 2013 (ZRC.ORT. 1021); (2) N01.44676, E 103.77919, 10.7 ± 5.2 m, among herbs, 25 June 2013, 2042 hours (ZRC.ORT. 1022); (3) N01.44682, E 103.77908, 17.2 ± 6.4 m, calling on leaf of tree sapling, calling song recorded in-situ, 6 July 2013, 2145 hours (MNHN-EO-ENSIF 3441). Diagnosis. Species of average size for the species group, characterised by contrasted coloration dark brown and yellow, with a wide rounded yellow area posterior and male genitalia more or less triangular, with a little rounded posterior apex. Very close to Cardiodactylus thailandia Robillard, 2011 from which it differs the following characters: general colouration brown and whitish while C. thailandia is almost black and vivid yellow; male FW: diagonal not bicolour; fore and middle legs: pale yellow brown with dark brown patterns (spots on femur, rings on tibiae) while legs of C. thailandia are almost entirely orange brown; male genitalia differ by pseudephiphallus posterior region (posterior lobe wider than in C. thailandia) and dorsal edge of dorsal ridges with different shape. C. admirabilis differs strikingly from Cardiodactylus singapura Robillard, 2011 by its colouration more contrasted on FW dorsal field, with a wide whitish area posterior to mirror (Fig. 1); and male genitalia: rami wide (narrow in C. singapura), with preapical plate more sclerotised along edge (equally sclerotised in C. singapura); ectophallic arc angulated (more broadly rounded in C. singapura); ectophallic apodemes lamellate (thin in C. singapura); endophallic sclerite sclerotized and rather wide, not tapering anteriorly but truncated with the apex acute (small and tapering anteriorly to an acute apex in C. singapura). Contrary to most species of the species group described from this geographic area (Robillard, 2011; Robillard et al. in prep), C. admirabilis shows a multisyllabic calling song and not a monosyllabic song. Description. Size average for the species group. General colouration contrasted, light brown with dark brown patterns (Fig. 2). Head dorsum yellow brown with 4 wide dark brown bands, more or less punctuated, bordered by dark brown triangles posterior to eyes and a yellow line more externally (Fig. 3 A). Fastigium yellow laterally, median area dark brown, joint with the 2 outermost dark brown bands on vertex. Scapus light brown. Antennae heterogeneously brown. Face yellowish to light brown, diversely mottled with brown, with 2 parallel lines on the fastigium (Fig. 3 B). Maxillary palpi yellowish. Head laterally sometimes with a dark brown area posterior to eyes and genae bordering the eyes, progressively lighter ventrally (Fig. 3 C). Pronotum: Dorsal disk trapezoidal, anterior margin concave, posterior margin straight or slightly bisinuated; dark brown mottled with light brown or yellow spots. Lateral lobes dark brown, with a black longitudinal band on dorsal half, ventral edges and dorsal margins yellow (Fig. 3 C). Legs light brown to yellow brown, knees dark brown, femora mottled with dark brown; tibiae darker, more or less banded; tarsomeres with dark brown ends (Fig. 2). Tarsomeres III- 1 with 3 spines on dorsoexternal edges (m= 3, n= 4). Hind wings longer than FWs, the dark brown tail exceeding the forewings less than twice longer than the pronotum (Fig. 2 A). Cerci light brown with faint black rings. Tergites dark brown or black; sternites light brown to brown. Subgenital plate yellow brown. Male. FW as shown in Fig. 3 D. Colouration: Dorsal field cells dark brown, veins mostly orange brown, some areas white with yellow veins, including base of FWs and anal veins, area near plectrum, part of the chords, and wide semi-circular area posterior to mirror. Base of chords with a dark brown sclerotization. Harp posterior angle and anterior area with a white and yellow sclerotized area. Lateral field with cells mostly dark brown, except areas near projections of Sc; veins M, R, Sc dark brown, transverse veins and projections of Sc pale yellow. FW venation: 1 A slightly bisinuated; stridulatory vein with transverse part 3.3 mm, with 176 teeth on the transverse area of the file, 18 on the curved area, and 22 near base of 1 A (n= 1, holotype) (Fig. 3 E). CuP absent. Harp longer than wide, with 2 w-shaped harp veins; posterior margin raised along diagonal vein. Mirror area variable, c 1 very long and very narrow and fairly straight to slightly sinuous, c 2 quite large; mirror (d 1) longer than wide, not rounded, generally separated in four parts by transverse and oblique veins, separating vein more posterior than anterior, the posterior part triangular; e 1 cell very elongated, along the mirror. Apical field long and triangular; with 3 wide cell alignments posterior to mirror, 2 apical most wide cells with dividing veins. Lateral field: Sc with 7–8 projections and 3 more ventral veins (m= 7.8, 3, n= 4). Male genitalia as shown in Fig. 4: Pseudepiphallus very sclerotized and little setose, more or less triangular, only slightly narrowed new mid-length. Dorsal ridges divergent posteriorly, their dorsal edge bean-like, slightly asymmetrical, carinated interiorly, the left one curved, the right one straight; pseudepiphallus bisinuated laterally. Pseudepiphallic sclerite with latero-anterior triangular expansions; the membrane between them with short strong setae. Apex of pseudepiphallus spoon-like, not rounded. Rami wide, with wide triangular preapical plate; plate with lateral edge rounded and more sclerotised; with apex lamellate and acute. Pseudepiphallic parameres trilobate, the posterior lobe wide. Ectophallic arc complete, with a short posterior expansion. Ectophallic apodemes lamellate, diverging, their apex curved dorso-interiorly. Apex of ectophallic fold trilobate, membranous. Base of ectophallic apodemes with thin sclerotized expansions oriented posteriorly. Endophallic sclerite small, with a medio posterior anpansion and 2 thin lateral arms sclerotised. Endophallic apodeme with both lateral lamellas, and a short medio-dorsal crest. Membrane of endophallic cavity smooth. Female. Unknown. Juvenile (Fig. 5). Early instars generally pale with irregular spots of different size; dorsum of head mostly dark brown with faint yellow bands; with black longitudinal bands from genae towards abdominal apex (Fig. 5 A, B). Later instars with more contrasted colouration, body darker with spots denser; dorsum of head with more contrasted yellow bands (Fig. 5 C). Measurements. See Table 1. Males (n = 4) 3.1–3.9 4.6–4.9 14.9–16.3 4.5–4.7 5.0– 5.5 15.8–15.9 - 13.6–14.6 (3.5) (4.8) (15.4) (4.6) (5.1) (15.9) (14.1) TIIIs TaIIIs Ias Ibs Oas Obs Holotype 9 2 13 2–3 5 Males (n = 4) 8–10 (8.8) 2 (2) 10–13 (12) - 3–5 (3.8) * Hind femur shrunk upon preservation. Etymology. The name reflects the surprise to discover a new species from the non-reserve parkland in a highly urbanised Singapore. The name also coincide with part of the name the type locality, Admiralty Park; from Latin, admirabilis = surprising. Distribution. This species is so far known from Admiralty Park in the north of Singapore and not found in central areas (Bukit Timah Nature Reserve and Central Catchment Nature Reserve) in which Cardiodactylus singapura exists. Conversely, Cardiodactylus singapura was not sighted and recorded from Admiralty Park. FIGURE 3. Cardiodactylus admirabilis n. sp. male: head in dorsal (A) and anterior (B) views; head and pronotum in lateral view (C); FW in dorsal view (D); stridulatory file (E). Scale bar: A–D, 1 mm; E, 0.5 mm. Habit and life history traits. Cardiodactylus admirabilis n. sp. is a nocturnal species, although the nymphs are sometimes sighted in the day. In captivity, the adults were fed with Dillenia fruits, flowers and leaves in which the species may be found. Behaviour. Calling song (Figs. 6, 7): The calling song of Cardiodactylus admirabilis n. sp. is a short echeme made of a triplet of syllables emitted by groups of 3 or 4, unlike the other Cardiodactylus species described from the region, including C. singapura, which have mono-syllabic calling songs (Robillard, 2011; Robillard et al. in prep). The calling song has the following features: echeme duration = 205.6 ± 7 ms, echeme period = 2.11 ± 1.3 s, echeme duty cycle = 11.7 ± 4.4 %; syllable duration = 36.9 ± 3.9 ms, syllable period = 84.8 ± 0.8 ms, syllable duty cycle = 43.5 %; dominant frequency (3 rd peak of the spectrum) = 14.26 0.74 kHz. The syllables within the echeme tend to be slightly different, as shown in Table 2: the frequency increases by more than 1.5 kHz between syllable 1 an d syllable 3; syllable 2 tend to be longer then the others, and syllables 1 and 2 include gaps, while syllable 3 is continuous and more intense. Discussion. The nymphs of this species were previously sighted in Admiralty Park and wrongly recorded in Tan et al. (2012) as Cardiodactylus singapura.Published as part of Tan, Ming Kai & Robillard, Tony, 2014, A new species of Cardiodactylus (Orthoptera: Grylloidea: Eneopterinae) from Singapore, pp. 364-376 in Zootaxa 3764 (3) on pages 365-371, DOI: 10.11646/zootaxa.3764.3.6, http://zenodo.org/record/25085

    Paranisitra leytensis Robillard & Yap 2015

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    Paranisitra leytensis Robillard, 2015 (Fig. 1) Paranisitra leytensis Robillard & Yap, 2015: 85; Baroga et al. 2016: 94 (taxonomic key). Type material. Holotype male. Philippines. [Visayas]: Leyte [Island], Burauen, Barangay Villa Corazon, zone herbacée [herbaceous area] (GPS Bar1), 10°57’50.5”N 124°46’ 35.3”E, 282 m (TR14), iii.2013, coll. T. Robillard (UPLBMNH). Allotype female. Philippines. [Visayas]: Leyte [Island]: Buo, zone secondaire près de la route [secondary area near road], (TR34), iii.2013, coll. T. Robillard (UPLBMNH). Paratypes (1 male, 5 females). Philippines. same information as HT and AT (MNHN-EO-ENSIF-3162–3166). [examined] Other material examined. Philippines. [Mindanao]: [Surigao del Norte]: Siargao Island, Del Carmen, Brgy. Esperanza, 1♂ (JBB139), 01–05.x.2016, coll. S.A. Yap (UPLBMNH), right middle leg for molecular analysis (N40); Del Carmen, Brgy. Katipunan, 2♀, 07.iv.2018, rainy, coll. J.B. Baroga-Barbecho, S.A. Yap, M.K. Tan, & H. Yeo (UPLBMNH, ZRC); 2♀ (Siargao18_44, 84), 14–17.x.2018, coll. M.K. Tan, J.B. Baroga-Barbecho & S.A. Yap (ZRC). [Mindanao Island: Agusan del Sur]: Casawangan Lake, 1♂ (JBB363), 22.vii.2015, coll. N.M. Barbecho (NMP 13579), left middle leg for molecular analysis (N39). Type locality. Philippines. Visayas, Leyte Island, Burauen, Barangay Villa Corazon, 10°57’50.5”N 124°46’ 35.3”E, 282 m. Distribution. Philippines. Visayas: Leyte Island; Mindanao: Siargao Island (new record); Mindanao Island: Agusan del Sur (new record). Diagnosis. Species close to P. longipes in terms of face coloration with a wide yellow band; differing by its smaller size and male genitalia with smaller and triangular pseudepiphallic lophi, shorter endophallic sclerite, and narrower endophallic apodemes. Remarks. The male specimens from Siargao Island and Mindanao Island (Agusan del Sur) were identified as P. leytensis based on the features of the body, face, size and male genitalia. They slightly differ from the specimens from Leyte by the following characters: in Siargao specimens, male genitalia wider than long, short and less setaceous lophi, and short rami; in Agusan specimen, thin and long male genitalia, ectophallic fold longer and narrowing basally. Male specimen from Mindanao Island is the smallest among examined specimens of the species.Published as part of Baroga-Barbecho, Jessica B., Yap, Sheryl A., Tan, Ming Kai & Robillard, Tony, 2019, Taxonomic review of the genus Paranisitra Chopard (Orthoptera: Gryllidae: Eneopterinae: Nisitrini) with description of a new species from Mindanao, pp. 81-96 in Zootaxa 4568 (1) on page 86, DOI: 10.11646/zootaxa.4568.1.5, http://zenodo.org/record/259926

    Matuanus rectinervus Robillard & Desutter-Grandcolas 2008, n. sp.

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    Matuanus rectinervus Robillard n. sp. Figs 3-6C TYPE MATERIAL. — Holotype male (MNHN-ENSIF 1096), 8 km NW Col d’Amieu, N la Foa, abattage, Pied Table Unio, 22.II.1994 (16) (L. Desutter-Grandcolas), identified M. elegans Otte, 1987 by L. Desutter-Grandcolas. Paratype male (MNHN-ENSIF 1097), Massif de l’Aoupinié, 400 m, 11 km SW Ponérihouen, milieu ouvert, tronc mort vertical, 21.III.1994 (5) jour (L. Desutter-Grandcolas), identified M. elegans Otte, 1987 by L. Desutter-Grandcolas. MNHN. TYPE LOCALITY. — New Caledonia, 8 km NW Col d’Amieu, N la Foa, Pied Table Unio. DESCRIPTION. — Very similar to M. elegans and M. priapus in coloration, body size and shape. Fastigium shorter than in M. elegans and M. priapus (Fig. 6C). Coloration very contrasted; face light yellow with dark patterns as in M. elegans and M. priapus, dorsal disk of pronotum red brown, lateral lobes yellow; legs I-II light yellow, femora with dark brown spots and tibiae with two dark brown rings; FIII yellow brown with two brown dorsal spots, a brown line on outer face and a very contrasted dark brown band on inner face, barely visible in M. elegans (Otte et al. 1987) and M. priapus. Male: FW venation (Fig. 3) almost similar to M. elegans and M. priapus, but 1Aa prolonged up to the apical field (compare Figs 1-2). Stridulatory file with 93 (HT) to 102 teeth (PT) (m = 98, n = 2). Male genitalia (Figs 4-5): very similar to M. priapus laterally except for the lateral edge of pseudepiphallic sclerite, straight and folded ventrally, while it is rounded in M. elegans and M. priapus. Endophallic apodeme as in M. priapus, wider and longer than in M. elegans and raised dorsally. Female: unknown. MEASUREMENTS. — Holotype male. PronL: 3.0 mm; PronW: 3.9 mm; FIIIL: 14.5 mm; FIIIW: 3.4 mm; TIIIL: 13.3 mm; FWL: 19.5 mm; FWW: 3.7 mm.Published as part of Robillard, Tony & Desutter-Grandcolas, Laure, 2008, Systematics of Matuanus Gorochov (Grylloidea, Podoscirtidae, Podoscirtinae) from New Caledonia: new data and the analysis of venation diversity, pp. 273-290 in Mémoires du Muséum national d'Histoire naturelle 196 on page 27
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