131,315 research outputs found
Eocenomyrma orthospina Dlussky et Radchenko
No full textE. orthospina M a t e r i a l e x a m i n e d. Two workers: 1) Baltic amber, CGC F-6800. Measurements (in mm): HL 0.69, HW 0.60, FW 0.27, FLW 0.31, SL 0.46, ML 0.75, PnW 0.75, PL 0.31, PW 0.23, PPL 0.21, PPW 0.30, ESD 0.27, total length ca. 3.0. Indices: HL/HW 1.15, FW/HW 0.45, FLW/FW 1.15, SL/HL 0.67, SL/HW 0.77, PL/HL 0.45, PPL/HL 0.30. 2) Rovno amber, SIZK K-7026 (specimen damaged, without petiole, postpetiole and gaster). Measurements (in mm): HL 0.66, HW 0.57, SL 0.52, FW 0.23, FLW 0.26, AL 0.85, PnW 0.39, ESL 0.31, ESD 0.30, HTL 0.52, total length ca. 3.5 (by analogy of ML+HL compare to other species). Indices: HL/HW 1.16, SL/HL 0.79, SL/HW 0.91, FW/HW 0.46, FLW/FW 1.13, ESL/ HL 0.47, ESL/HW 0.53. A key for the identification of Eocenomyrma species 1. Head, mesosoma and waist only densely punctated, without rugosity or reticulation (fig. 7). Scape relatively very short, SL/HL 0.52, SL/HW 0.57. Petiole with very short peduncle, PL/ PH 1.13, its anterior surface steep, very slightly concave, meets with dorsal surface of node at an acute angle, dorsal plate short ant strongly declined posteriorly so that petiole seems cuneiform (seen in profile) (figs 8, 10)...................................................................................................................................................... E. breviscapa sp. n. – Head, mesosoma and waist longitudinally rugose or reticulated; petiole of another shape, but in any case with much longer peduncle, PL/ PH> 1.25 (figs 5, 6, 12–17) Scape distinctly longer, SL/HL> 0.58, SL/HW> 0.70...................................................................................................................................................... 2 2(1). Gyne: propodeal spines relatively short (ESL/HL 0.17), straight, slightly widened at the base; head dorsum and mesosoma mostly longitudinally rugose, coarse reticulation presents on occipital area of the head (figs 5, 6)....................................................................................................................... E. ukrainica sp. n. – Workers: propodeal spines longer (ESL/HL> 0.23), more widened at the base, often curved down along their length, if straight, then much longer, ESL/HL> 0.35 (figs 12–17)..................................................... 3 3(2). Head dorsum and mesosoma with longitudinal, slightly sinuous rugosity, without reticulation (fig. 12)......................................................................................................................................... E. rugosostriata (Mayr) – Head dorsum and mesosoma at least partly with reticulation (figs 13–17)................................................ 4 4(3). Whole head dorsum and mesosoma with fine reticulation; petiolar node with well developed, flattened dorsal plate (fig. 13)................................................................................ E. elegantul a Dlussky et Radchenko – Frons with longitudinal, slightly sinuous rugae, remainder part of head dorsum with coarse reticulation; petiolar node with rounded dorsum, without dorsal plate (figs 14–17)............................................ 5 5(4). Propodeal spines thin, not widened at the base, straight, directed backward and upward; petiole with very long peduncle (PL/ PH> 1.90) (figs 14, 15)................................ E. orthospina Dlussky et Radchenko – Propodeal spines massive, widened at the base, slightly curved down along their length, directed mainly backward; petiole with much shorter peduncle (PL/ PH 1.27) (figs 16, 17)........................................................................................................................................................................ E. electrina Dlussky et RadchenkoPublished as part of Radchenko, A. G. & Dlussky, G. M., 2016, Further Data On The Extinct Ant Genus Eocenomyrma (Hymenoptera, Formicidae), pp. 395-406 in Vestnik Zoologii 50 (5) on pages 405-406, DOI: 10.1515/vzoo-2016-0046, http://zenodo.org/record/644928
The Upper Paleolithic rock art of Ukraine between here and nowhere
No full textThe complex of Kamyana Mohyla is the westernmost rock art location of the Eurasian Steppe and the largest
accumulation of cave art sites in the Eastern Europe. So far it has been believed that the complex contains the
Upper Paleolithic cave art images as well as portable art collection that resemble the instances of Upper
Paleolithic worldview. Though this belief lacked the support of archaeological context and chronological attribution it remained neither proved nor disputed. However, the application of digital photogrammetric tools
allowed to perform the sub-millimeter surface modeling of the rock art objects and to re-examine and reconsider
the engravings that were previously attributed to Pleistocene. The modeling results presented in this article
revealed the complete absence of figurative images for the collection of portable art specimens and the dubious
character of those for the cave art one. Therefore, the whole collection should be reconsidered, studied and
attributed according to the state of the art and contemporary archaeological record in the region. This contribution attempts to think over the possible Upper Paleolithic origin of the motifs from Kamyana Mohyla in the
light of new data and proposes three hypotheses towards the understanding of the rock art assemblage from one
of the caves in the complex
Dasypoda michezi Radchenko 2017, sp. n.
No full textDasypoda michezi Radchenko, sp. n. Type material: Holotype ♂ with the printed label: “ Dasypoda ♂, Almograve (Port[ugal].) [37.6524, -8.7960], 21- V-2012, A.Livory & R.Coulomb leg.”. Paratype: 1 ♂ with the handwritten label: “ Dasypoda ♂, Vila Nova de Milfontes (P[ortugal]) [37.723, -8.788], 20-V-2012, A.Livory u R.Coulomb leg.”. Both specimens are stored in the collection of Naturalis Biodiversity Center, Leiden, the Netherlands (RMNH). Etymology. Named after Denis Michez, an authority on Dasypoda and other bees. Diagnosis. Dasypoda michezi sp. nov. may be regarded a representative of the subgenus Heterodasypoda due to the possession of all its diagnostic features, namely the shape and structure of maxillary palpi, galea (Fig. 7), malar space, nervulus (Fig. 1), S6–8 (Figs 23, 27, 31, 35, 39), genitalia (Figs 43, 47, 51, 55), and gonostylus (Figs 59, 63, 67, 68). The new species differs from other species of this subgenus by having: apical projection of S6 apically deeply emarginated in the form of wide triangle (Fig. 23), apicolateral processes of S7 with triangle shaped distal expansion (Fig. 28), inner ventral lobe of gonostylus with only few setae (Figs 63, 71), distal half of external lobe of gonostylus significantly widened and convex in middle of outer surface, diamond-shaped (Figs 43, 47, 51, 55) and its inner surface with several hairs, each of which inserted into crater-like raised alveoli (Figs 67–69, 72). Description. Male: Holotype. Body length ( vertex to T7): 11.8 mm (paratype: 12.0 mm). Mesosoma: W (between tegulae) = 3.0 mm (same in paratype). Head: L = 2.9 mm (paratype: 3.1 mm), W = 3.3 mm (paratype: 3.5 mm); Body black, head slightly wider than long (Figs 3, 4), clypeus, densely punctured by oblique, downward punctures. Lateral edges of clypeus with impunctate bands on sides and apex of clypeus with narrow depressed band (Fig. 4). Face below antennal toruli with relatively wide (equal to the width of scapus), smooth, almost impunctate band reaching base of clypeus. Malar space narrow, 0.15× as long as broad, in 2× shorter than pedicel. Body mostly covered with light yellow-grey hairs (Figs 1–6); head also with such hairs except upper part of face between compound eyes and vertex covered with sparse brown hairs (Figs 3, 4). Back of head and thorax with long, semi-erect hairs. Wings slightly infuscate, notably in distal half. Legs with long greyish-yellow pubescence, apart from dense short dark-yellow pubescence of inner side of metabasitarsus, and very short grey hairs of 2–5th tarsomeres with a sparse sheaf of long hairs on apices of 2–4th tarsi. T1–2 covered by light, long erect hairs, T3–5 with semi-erect hairs and T6–7 with appressed hairs (T5–6 with pale pubescence only on the margin and sides), each hair rising from an oblique backward-directed puncture. Distances between puncture about 2–4× as long as diameter of puncture. Pubescence sparse and revealing surface of tergum. T2–6 with very short, thin branched dark-brown hairs situated in areas between long hairs (length of lateral branches equal to 1/2–2/3 of total hair length). T5–6 with basal parts bearing only dark-brown hairs with small admixture of light hairs on sides. Basal parts of sterna densely punctate, mainly bearing dark brown hairs. Central part of S5–6 with larger sparse punctures. S3–5 with apical bands of appressed dark-brown hair, with longer light yellow-grey hairs on sides of sterna (Fig. 6). Apical part of S6 with wide and deeply emarginated outgrowth (Fig. 23); distal part of latero-apical processes of S7 triangularly expanded (Fig. 28); apex of S8 posterodorsal part with transverse carina slightly notched in center (Fig. 35; carina shown by white arrow). Inner ventral lobe of gonostylus almost bare on apex (Figs 63, 71); distal half of external lobe of gonostylus significantly widened and convex on its outer surface, diamond-shaped (Figs 43, 47, 51, 55), its inner surface with several hairs, each inserted into crater-like raised alveoli (Figs 67–69, 72). Female: unknown. Distribution. South-western Portugal (Fig. 73). Floral visitation. Pollen grains of a Cistus plant (Cistaceae) were found on the body of the paratype (Fig. 70). Other representatives of this subgenus, in particular D. morotei and D. pyrotrichia, are also associated with Cistaceae (Michez et al. 2004b, 2008; Ruiz 2013; Özbek 2014), whereas D. albimana is mostly found on Asteraceae, but also recorded on Cistaceae, Lamiaceae and Rosaceae (Michez et al. 2003, 2004b; Ruiz 2013). Flight period. May. All other Heterodasypoda species also fly in late spring to summer: D. albimana —from May to July (Michez et al. 2003); D. morotei —from May to August, with some sporadic records in April and September and one—later in November (Ornosa & Ortiz-Sánchez 1998, 2004); D. pyrotrichia —from mid May to late August (Grace 2010). Comparative remarks. Dasypoda michezi sp. nov. is most likely to be confused with D. albimana. Both species have similar body length and size of head, approximately same surface punctation of different body parts and colour of pubescence. However, D. michezi sp. nov. is characterized by the shorter first flagellar segment (Figs 15, 19) and differs from all other species of the subgenus in the ratio of different flagellar segments (Figs 17–18, 21–22; Tabl. 1). Flagellomeres 3–8 of D. michezi sp. nov. are cylindrical (Fig. 15), the same as in D. albimana (Fig. 16) and D. pyrotrichia (Fig. 18), in contrast to D. morotei, in which they are slightly curved (Fig. 17). The clypeus of D. michezi sp. nov. (Figs 3, 4) lacks an impunctate median band (this feature is almost invisible when pubescence is undamaged), likewise in D. albimana. Also, the apex of the clypeus is straight in D. michezi sp. nov. (the clypeus of D. morotei and D. pyrotrichia has a more or less distinct, impunctate median band; also it the apex is emarginated in D. morotei). The glossa of D. michezi sp. nov. (Fig. 11) is very narrow and long: 1.5× longer than in D. albimana (Fig. 12) and D. morotei (Fig. 13), but has same length as in D. pyrotrichia (Fig. 14). The most noticeable differences concerns the ratio of the glossa width at base to its length: this proportion is about 1: 6.7 in D. michezi sp. nov., 1: 4.6 in D. pyrotrichia, 1: 3 in D. albimana and 1: 2.3 in D. morotei (Figs 11–14). The surface of the galea of D. michezi sp. nov. (Fig. 7) is clearly structured with many small tubercles merged into sinuous lines in many places, and is less shiny than in D. albimana and D. pyrotrichia (Figs 8, 10). This surface is completely covered with separate small tubercles in D. morotei (Fig. 9). Species Flagomere L: W (µm) Flagomere L: W ratio Flagomeres length ratio 1st* 2nd 1st* 2nd 3rd... 9th 1st: 2nd Dasypoda michezi sp. nov. (n = 2) 412/290 320/295 1.41 1.08 1.1÷1.2 1.28 D. albimana (n = 4) 475/257 268/263 1.85 1.02 1.1÷1.2 1.77 D. morotei (n = 2) 358/293 398/284 1.22 1.40 1.4÷1.5 0.90 D. pyrotrichia (n = 3) 450/290 370/296 1.55 1.25 1.4÷1.6 1.22 *—width on apex. The main significant differences between D. michezi sp. nov. and the other three species of Heterodasypoda concern the structure of S6–8 of males. The outgrowth on apical part of S6 is significantly expanded at apex (ratio between width of outgrowth at base to its width at apex is 1: 1.25), apically deeply emarginated in the form of wide triangle (Fig. 23) in D. michezi sp. nov., unlike other species in which such outgrowth on apex is not expanded (in D. albimana and D. morotei; Figs 24, 25) or almost undeveloped (in D. pyrotrichia; Fig. 26). The new species also differs in the structure of latero-apical process at S7 (Figs 27–30). S8 of D. michezi sp. nov. is chunkier (Figs 31, 39), and superficially appears similar to D. albimana (Figs 32, 40), but significantly differs from that of D. morotei, which has undulating depressions on the sides of basal part of this segment (Figs 33, 41; shown by black arrows), and D. pyrotrichia is possessing more extended sides on the basal part of this segment (Figs 34, 42). The transverse carina situated anteriorly to the apex of the inner part of S8 has a small notch in the center in D. michezi sp. nov. (Fig. 35), (cf. D. albimana and D. morotei (Figs 36, 37) in which it is more deeply cut in or even completely bilobed in D. pyrotrichia (Fig. 38)). The inner ventral lobe of the gonostylus is almost bare in D. michezi sp. nov., with a few thin and short setae (Figs 63, 71), unlike other species, in which this lobe is covered with numerous, more or less long and pointed setae (Figs 64–66). The distal half of external lobes of the gonostylus of D. michezi sp. nov. is considerably widened and convex in the middle of the outer surface, diamond-shaped (Figs 43, 47, 51, 55). Unlike the new species, these lobes are narrow or more strongly elongated toward apex and widely concave in the middle of outer surface in other discussed species (Figs 44–46, 48–50, 52–54, 56–58). The inner medial surface of the outer lateral lobe of gonostylus is bearing several hairs, each of which is inserted into the crater-like raised alveoli (Figs 67–69, 72) in D. michezi sp. nov.; no such hairs are found in other species of Heterodasypoda.Published as part of Radchenko, Vladimir G., 2017, A new bee species of the genus Dasypoda Latreille (Hymenoptera, Apoidea) from Portugal with comparative remarks on the subgenus Heterodasypoda Michez, pp. 164-176 in Zootaxa 4350 (1) on pages 167-174, DOI: 10.11646/zootaxa.4350.1.10, http://zenodo.org/record/105202
Dasypoda morawitzi Radchenko, sp. n.
No full textDasypoda morawitzi Radchenko, sp. n. Type material: Holotype ♂, “Херсонська обл., ЧБЗ, Івано-Рибальчан. дільн. [46.45347, 32.10651], 0 7.08.2015, Котенко А.” [Ukraine, Kherson Region, Black Sea Biosphere Reserve, Ivano-Rybalchanskiy plot, leg. A. Kotenko] (IEEK). Paratypes—293 ♂ 153 ♀: Ukraine: “settlement Donetskoe [48.934722, 37.690556], Slavyansk district, Donetsk Region, 26.07.1973, leg. V. G. Radchenko ”, 1♂ (IEEK); “ Donetsk Botanical Garden [48.01, 37.880556], 8.08.1978 on flowers of Cichorium intybus, leg. V. G. Radchenko ”, 4♂ 1♀ (IEEK); “ Donetsk Region, Enakievo, settlement Olkhovatka [48.23771, 38.42073], 10.08.1978, on Picris sp., leg. V. G. Radchenko ”, 1♀ (IEEK); “ Stalinskaya obl. [Donetsk Region], step Khomutovsky [Ukrainian Steppe Natural Reserve] [47.287, 38.185], 4.vii. [19]48”, 1♂ (SIZK); “ Kherson [Region], Aleshkinskie Peski [Oleshky Sands] [46.583333, 33.05], 26.07.1926, L. Zimin ” 1♂ (ZISP); “ Kherson region, environs of Tsiurupynsk [46.633333, 32.583333]: 0 2.09.2013, leg. A. Yu. Pantova ”, 1♀ (IEEK); 10.09.2014, leg. M. I. Zayika, 1♂ (IEEK); “ Kherson Region, Tsiurupynsk district, [village] Solontsy [46.57732, 32.67538], 28.vi. [19]50 [leg. A. Z. Osychnjuk]”, 1♂ (SIZK); “ Kherson Region, Tsiurupynsk district, environs of village Solontsy [46.57550, 32.64484]”: 0 3.09.2013, leg. M. I. Zayika 1♂ (IEEK); 12.09.2014, leg. M. I. Zayika 1♂ (IEEK); “ Kherson region, Black Sea Biosphere Reserve [46.44827, 32.14668]”: 0 7.08.2015, leg. M. I. Zayika, 2♀ 1♂ (IEEK); 0 5.09.2013, leg. M. I. Zayika, 1♀ (IEEK); “ Kherson region, Black Sea [Biosphere] Reserve, Ivano- Rybalchanskiy plot [46.45347, 32.10651], 10.07.1980, leg. [L.] Bodnarchuk et [I.] Shalimov ”, 1♀ (IEEK); “ Kherson region, ChBZ [Black Sea Biosphere Reserve], Ivano-Rybalchanskiy plot [46.45347, 32.10651], 0 7.08.2015, [leg.] A. Kotenko ”, 1♀ 1♂ (IEEK); “ Kherson region, Black Sea Biosphere Reserve, Solenoozerniy plot, cordon behalf of Parkhomenko [46.45607, 31.97571], 09– 13.08.2014, leg. O. Kumpanenko ”, 1♀ 2♂ (IEEK); “ Kharkiv Region, [settlement] Rohan [49.939722, 36.49], 14.07.1981 on flower of Campanula sp., leg. M. Filatov ”, 1♂ (IEEK); “environs of Kiev [50.35992, 30.45324], 18.iv. [19]54, on flowers of Tussilago farfara ”, 1♀ (SIZK); “ Kiev distr., [village] Hodosovka [50.275, 30.515], 26.07.[20]04, [leg.] M. Nesterov ”, 1♀ (IEEK); “ Kiev Region, Makarov [Kiev-Sviatoshyn] district, village Muzychi [50.35116, 30.11284], 0 1.07. [20]14, leg. M. O. Nesterov ”, 1♀ (IEEK); “ Kiev Region, village Muzychi, 21.07.2012, leg. M. O. Nesterov ”, 2♀ (IEEK); “ Kiev, island Murometz [50.50577, 30.54296], 8.07.2014, leg. A. Yu. Pantova ”, 2♂ (IEEK); “ Kiev, reserve “Bobrovnya” [50.51377, 30.53706], 0 1. 0 8. [20]12, on Aster [aceae]”, 6♀ (IEEK); “[Kiev], Park “Druzhby Narodov” [50.49588, 30.54166], 01.08.[20]12, on Aster [aceae]”, 1♀ (IEEK); “ Zaporoz [hye] Region, Tokmak area, [village] Kohanoe [47.294843, 35.509602], 0 1.08.2001, 0 4.08.2001, leg. Zhakov ”, 2♀ (IEEK); “S[outhern] coast of Crimea from Karatau to Yayla [44.841, 34.522], leg. Vydhalm ” 1♂ (ZISP); “ Crimea, Simferopol district, [village] Urozhaynoye [45.058333, 34.123611]: 0 2.08.2012, leg. V. Zhidkov, 1♀ 1♂ (IEEK); 0 2.08.2012, on Scabiosa sp. leg. S. Ivanov, 4♂ (TNUS); 9.08.1999, leg. S. P. Ivanov 1♂ (TNUS); “ Crimea, Simferopol district, [village] Krasnolesye [44.834722, 34.229167], 16.05.1973 ”, 1♀ (TNUS); “ Crimea, Simferopol, Khoshkeldy [45.00250, 34.17795], 27.07.2004, leg. V. Zhidkov ”, 1♂ (TNUS); “ Crimea, Belogorsk district, [village] Krımskaya Roza [45.054167, 34.359722], 6.viii. [19]69 on Crepis tectorum, [leg.] Turkhanova ”, 1♀ (TNUS); “ Crimea, Belogorsk district, [village] Zuya [45.054167, 34.315278], [leg.] Turkhanova ”: 7.viii. [19]69, on Xeranthemum annuum, 2♀ 6♂ (TNUS); 7.viii. [19]69 on Chondrilla juncea, 2♀4♂ (TNUS); 8.viii. [19]69, on Cichorium sp., 1♀ (TNUS); 15.viii. [19]69, on Cichorium sp., 2♂ (TNUS); 15.viii. [19]69 on Knautia sp., 2♂ (TNUS); “ Crimea, [village] Zolotoe [45.422222, 36.076389], 0 3.06.1999, leg. S. P. Ivanov ”, 1♀ (TNUS); “ Crimea, [village] Kurortnoe [45.472222, 36.3375], 0 1.08.2004, leg. S. Ivanov ”, 2♂ (TNUS); “ Crimea, Chernomorsk district, [village] Gromovo [45.388889, 32.869444], 24.05.1988, leg. S. Ivanov ”, 1♂ (TNUS); “[Crimea], Yevpatoria [45.193889, 33.368056], 27.viii. [19]29”, 1♀ (TNUS); “ Crimea, Krasnogvardeysky district [near] river Salgir [45.35987, 34.31219], 24.07.1985, leg. S. Ivanov ”, 3♀ (TNUS); “ Crimea, Sevastopol, Kamyshovaya bay [44.57332, 33.43434], 27.08.1974, leg. S. P. Ivanov ”, 1♀ (TNUS); “ Crimea, cape Kazantip [45.466667, 35.866667], 7.8.1977, [leg.] M. Filatov ”, 1♀ (IEEK); “ Crimea, cape Kazantip, steppe, 0 2.08.2003, leg. S. Ivanov ”, 1♀ (TNUS); “ Crimea, Tarkhankut [peninsula], the bay Kipchak, [45.48029, 32.59258], 14.06.2012, leg. V. Zhidkov ”, 1♂ (IEEK); “ Crimea, Tarkhankut peninsula, Kipchak valley [45.47217, 32.59929], dry saline, 22.06.2007, leg. Fateryga ”, 1♀ (TNUS); “ Chernigov Region, Menskyi district, village Maksaky [51.424722, 32.186111], 29.vii.1978, meadow, on flowers of Inula brit [annica], leg. [L.] Romasenko ”, 1♂ (SIZK); Ibid. “ 31.vii.1978, leg. [L.] Romasenko”, 1♀ (SIZK); “ Chatki, d. Zenkov [village Zinkiv] [50.2, 34.35], g. Poltava, 14.viii. [19]25, [leg.] Kistyakovskyi ”, 2♀ (SIZK); “[village] Yaresky [49.84617, 33.91617], Myrh [orod] d[istrict], Poltava p[rovince], 24.vii.1925, [leg.] Fabri ”, 1♀ (ZISP); “ Gadyach [50.366667, 34.0] [Poltava Province], 14.08.1909 ”, 1♀ (ZISP); “[Sumy Region, Lebedynskyi district], Mikhaylovskaya celina Nature Reserve [50.833333, 34.083333], 24.vii. [19]49, [leg.] G. Kolmaz ”, 1♀ (SIZK); “[Cherkassy Region], Kaniv biozapov. [Nature Reserve] [49.744444, 31.455833], 24.vi. [19]50 on Stachus [Stachys], [leg.] G. Kolmaz ”, 1♀ (SIZK); “ Kaniv biozapov., a glade, 23.viii. [19]49, [leg.] G. Kolmaz ”, 1♀ (SIZK); “ Kaniv biozapov., Mar'ina g., 17.viii. [19]49 [leg.] G. Kolmaz ”, 1♂ (SIZK); “[Cherkassy Region], Khutir Khmil’na [49.68251, 31.53703], Kaniv district, 9.viii. [19]48, [leg. A. F.] Krishtal ”, 1♂ (SIZK); “ Elisavetgrad [Kirovohrad] [48.509, 32.255], 5.vii. 1902, [leg.] E. Yatsentkovsky, 1♂ (ZISP); “ Kirovograd Region, [Oleksandriia district], village Bandurovka [48.734444, 32.954444], [leg. A. Z. Osychnjuk]”: 3.vii. [19]53, on flowers of Cichorium, 2♀ (SIZK); 13.vii. [19]53 on flowers of Cichorium int [ybus], 1♀ (SIZK); 16.vii. [19]53, on flowers of Echium, 2♀ (SIZK); 18.vii. [19]53, on flowers of Scabiosa, 1♀ (SIZK); 19.vii. [19]53 on flowers of Picris hierac [ioides], 4♀ (SIZK); 24.vii. [19]53 on flowers of Cichorium, Picris hierac [ioides], 4♀ (SIZK); 4.viii. [19]53, on flowers of Scabiosa ochr [oleuca], 1♂ (SIZK); 25.vii. [19]53, on flowers of Carduus, 1♂ (SIZK); 31.vii. [19]53, on flowers of Balota ru [?], 1♂ (SIZK); 10.viii. [19]53, on flowers of Eryngium, 1♂ (SIZK); 11.viii. [19]53, on flowers of Inula br [itannica], 1♀ (SIZK); 12.viii. [19]53, on flowers of Carduus, 3♂ (SIZK); Vinnytsia Region, village Mikhaylivka [48.8075, 28.290556] [leg. A. Z. Osychnjuk]: 15.vi. [19]53, 1♂ (SIZK); 18.vii. [19]53, on flowers of Carduus, 4♂ (SIZK); 2.viii. [19]54, on flowers of Carduus, 1♂ (SIZK); 15.viii. [19]54, on flowers of Carduus, 1♂ (SIZK); “ Ismail region [Odessa Region], Kislitskiy island [45.3916996, 29.0960884], 25.vii. [19]51, on Cirsium, [leg. A. Z. Osychnjuk]”, 1♂ (SIZK); “ Ismail region, Borodinskyi district [Odessa Region, Tarutyne district], village Lesnoe [46.4675, 29.350278], 12.viii. [19]51, [leg. A. Z. Osychnjuk]”, 1♂ (SIZK); Russian Federation: “ Derbent ” [42.0585, 48.276] [Dagestan], “coll. F. Morawitz ” [with the label “ Dasypoda plumipes Pz. ” that handwriting by F. Morawitz], 1♀ (ZISP); “Caucasus [leg.] Bekker”, “coll. F. Morawitz ”, 1♀ 1♂ (ZISP); “[village] Starogladkovskaya [43.633, 46.418], Kizl[yar] Okr. Tersk. [Shelkovskoy District, Chechen Republic], 9.vii. 1927, [leg.] Kiritshenko ”, 1♂ (ZISP); “ Slavyanskaya [45.25, 38.1 167] [Slavyansk-na-Kubani, Krasnodar Krai], N. Cauc [asus], [leg.] Z. Koshur, 19.vii.1937 ”, 1♀ (ZISP); “environs of Orenburg [51.75, 55.13], [river] Ural — meadows: 18.vii.1923, [leg.] Vorontsovskiy”, 1♀ (ZISP); 4.viii.1922, [leg.] Vorontsovskiy ”, 1♀ (ZISP); “ Sarepta ” [Volgograd] [48.512, 44.5499], “coll. F. Morawitz ”, 1♀ 1♂ (ZISP); “ Sarepta 1892 [leg.] Bekker ”, “coll. A. Yakovlev ”, 2♂ (ZISP); “ Sarepta, Saratov prov.”, “coll. L. Wollmann ”, 1♂ (ZISP); “ Saratow ” [51.541, 45.958], “coll. F. Morawitz ”, 1♂ (ZISP); “Urbach [51.237, 46.977], Sarat[ov Region], 24 – 27.07.1926, [leg.] A. Shestakov ” 2♂ (ZISP); “ Orenburg Region [51.76, 55.14], 2.viii.1929, [leg.] P. A. Vorontsovskiy ” 2♂ (ZISP); “ Spask, Jul. ”, [village Spasskoe, Orenburg region] [52.01, 56.53], “coll. Eversmann ”, 3♂ (ZISP); “Luga [58.733333, 29.816667], St-Petersburg, [leg.] Solskiy”, 1♂ (ZISP); “Rostov-on-Don [47. 240556, 39.710556]: 22.07.1963, on Cichorium, [leg. Yu. A.] Pesenko ”, 1♀ (ZISP); 7.08.1963, on Cichorium, [leg. Yu. A.] Pesenko ”, 2♀ (ZISP); “[Bashkortostan, Kuyurgazinsky District] [village] Murap [tal] [52.445944, 55.811472], 22.vii. [19]52, [on] Cich [orium] int [ybus]”, “coll. [K. S.] Nikiforuk ”, 2♀ (SIZK); “[village] Murap [tal], 24.vii. [19]52 [on] Cich [orium] int [ybus]”, “coll. [K. S.] Nikiforuk ”, 1♀ (SIZK); “ Yermekeyevsky [District of Bashkortostan] [54.083333, 53.666667], 10.vii. [19]57, [on] Orig [anum] vulg [are]”, “coll. [K. S.] Nikiforuk ”, 1♀ 1♂ (SIZK); Kazakhstan (all specimens are deposited in ZISP): “ Uralsk [51.233333, 51.366667], [northwestern Kazakhstan], 29 – 30.07.1926, [leg.] A. Shestakov”, 51♂; “ Ber Tschogur Mugodjargebirge [48.459722, 58.553333] [village Birshogyr, Shalkar District, Aktobe Region of Kazakhstan], [leg.] L. Bobyr ”, “coll. Wollmann ”, 1♂; “Kharkin [48.74125, 51.81873], the lower reaches of the Ural River, Kazakhstan ”: 4.vii. [1]951, [leg.] Rudolph, “ Cirsium arvense ”, 1♂; 4.vii. [1]952, [leg.] Rudolph, “ Senecio jacobaea ” 2♂; 4.vii. [1]952 [leg.] Rudolph, “ Convolvulus arvensis ”, 1♂; 10.vii. [1]951, [leg.] Rudolph, “ Senecio jacobaea ”, 1♂; 10.07.1951, [leg.] Rudolph, “ Cichorium intybus ” 1♂; 22.vii. [1]951, [leg.] Popov, “ Cirsium arvense ”, 2♂; 3.viii. [1]951, leg. Rudolph, “ Mulgedium sibiricum ”, 1♂; “South [of the village] Rozhkovo, [51.663333, 52.304722], West Kazakhstan, 8.vii. [1]949, on fly, [leg.] Steinberg ”, 1♂; “Yanvartsevo [51.444253, 52.246097], [Zelenov District, West Kazakhstan Region], right b[ank of the] Ural [River], Kazakhst[an]”: 30.vi. [1]950, [leg.] Rudolph, “ Cichorium intybus ”, 1♂; 4.vii. [1]949, [leg.] Rudolph, “ Cichorium intybus ”, 1♀; 4.vii. [1]949, [leg.] Rudolph, “ Lycopus europaeus ”, 2♂; 4.vii. [1]949, [leg.] Rudolph, “ Cichorium intybus ”, 1♂; 8.vii. [1]949, [leg.] Rudolph], “ Cichorium intybus ”, 11♂; 20.vii. [1]949, [leg.] Popov, “ Cichorium intybus ”, 3♀ 7♂; 20.vii. [1]949, [leg.] Rudolph, “ Cichorium intybus ”, 6♀; 21.vii. [1]949, [leg.] Popov, “ Inula britannica ”, 1♂; 21.vii. [1]949, [leg.] Rudolph, “ Acroptilon picris ”, 4♂; 21.vii. [1]949, [leg.] Rudolph, “ Inula britannica ”, 1♀5♂; 22.vii. [1]949, [leg.] Rudolph, “ Echinops ruthenicus ”, 1♂; 22.vii. [1]950, [leg.] Rudolph, 1♂; 23.vii. [1]949, [leg.] Popov, “ Inula britannica ”, 1♀ 3♂; 24.vii. [1]949, [leg.] Popov, “ Eryngium planum ”, 1♀; 24.vii. [1]949, [leg.] Rudolph, “ Eryngium planum ”, 1♀; 24.vii. [1]949, [leg.] Rudolph, “ Limonium gmelinii ”, 10♀; 24.vii. [1]949, [leg.] Rudolph, “ Cichorium intybus ”, 1♂; 24.vii. [1]949, [leg.] Rudolph, “ Inula britannica ”, 17♀ 8♂; 25.vii. [1]949, [leg.] Rudolph, “ Cichorium intybus ”, 1♂; 27.vii. [1]949, [leg.] Rudolph, “ Inula britannica ”, 1♂; 31.vii. [1]949, [leg.] Popov, “ Limonium gmelinii ”, 1♂; 31.vii. [1]949, [leg.] Rudolph, “ Limonium gmelinii ”, 1♀ 6♂; 31.vii. [1]949, [leg.] Rudolph, “ Cichorium intybus ”, 6♀ 35♂; 2.viii. [1]949, [leg.] Rudolph, “ Inula britannica ”, 1 ♂; 2.viii. [1]949, [leg.] Popov, 1♀; 3.viii. [1]949, [leg.] Popov, “ Limonium gmelinii ”, 6♀ 47♂; 3.viii. [1]949, [leg.] Rudolph, 2♂; 3.viii. [1]949, [leg.] Rudolph, “ Eryngium planum ”, 2♀; 3.viii. [1]949, [leg.] Rudolph, “ Sonchus arvensis ”, 1♀; 3.viii. [1]949, [leg.] Vyrzhikovskaya, 1♀; 4.viii. [1]949, [leg.] Popov, “ Inula britannica ”, 1♀; 4.viii. [1]949, [leg.] Rudolph, “ Cichorium intybus ”, 6 ♂; 4.viii. [1]949, [leg.] Rudolph, “ Inula britannica ”, 6♀; 5.viii. [1]949, [leg.] Rudolph, “ Limonium gmelinii ”, 1 ♂; 5.iii. [1]949, [leg.] Rudolph, “ Cichorium intybus ”, 1♀ 1♂; 10.viii. [1]949, [leg.] Rudolph, “ Cichorium intybus ”, 2♀; 12.viii. [1]949, [leg.] Rudolph, “ Cichorium intybus ”, 9 ♂; 14.viii. [1]949, [leg.] Arnoldi, “ Cichorium intybus ”, 1♀; 14.viii. [1]949, [leg.] Rudolph, “ Inula britannica ”, 2♀; 17.viii. [1]949, [leg.] Rudolph, “ Cichorium intybus ”, 1♀ 2♂; 18.viii. [1]949, [leg.] Rudolph, “ Aster punctatus ”, 2♂; 4.ix. [1]949, [leg.] Rudolph, “ Chondrilla juncea ”, 2♀1♂; 4.ix. [1]949, [leg.] Rudolph, “ Scabiosa isetensis ”, 1♀. Turkey. “TR: Ihlara Vadisi, S Selime (1200 m) [38.3014, 34.2574], 28.07.2016, leg. [M.] Kasparek”, 2♀ (IEEK). Etymology. Named after Ferdinand Morawitz. His collection contains several specimens of the new species. Diagnosis. The species is very similar to D. hirtipes, hardly distinguishable from it in general appearance. Specimens of D. morawitzi are slightly smaller (body length: female 9–14 mm [N = 50], fig. 1–4; male 9.5–13.5 mm [N = 50], fig. 9, 10, 13, 14, 17) than D. hirtipes (female 11.5–15.5 mm; male 10–15 mm), but still broadly overlap in size. The main character, which sharply distinguishes these species is the structure of galeal surface in both males and females, rippled by wavelike lines in D. morawitzi (fig. 20, 21) and completely covered with small tubercles in D. hirtipes (fig. 22, 23) as well as in D. oraniensis (fig. 26, 27) and D. albipila (fig. 28). The galeal surface is intermediate in D. sinuata (fig. 24, 25) and partly D. tubera Warncke, 1973 (fig. 29). However, the latter species differs sharply by the structure of the gonostylus, which has three well-developed processes. In comparison with D. hirtipes, the central apical part of male S 6 in D. morawitzi is more produced anteriorly (fig. 35, 38), the apical projections on S7 are more broadly rounded (fig. 36, 39), S8 is wider, with more projected outgrowths at each side (fig. 37, 40–42), the apical part of S8 is wider and rounded, with two separated teeth on its the ventral side (fig. 43), which are shorter and closer than those of D. hirtipes (fig. 44) and D. sinuata (teeth connected in D. albipila). The inner process of the gonostylus has a shorter fringe of hairs on the margin (fig. 45, 47, 49, 50, 61 – 66), similar to that of D. albipila (fig. 49, 50) and D. sinuata, but distinct from the very long branched hairs in D. hirtipes (fig. 46, 48, 55, 56). The base of the gonostylus in D. morawitzi is untoothed, only with an elongated area that is rounded, crest-shaped, and protruded apically (fig. 51, 61); in D. albipila the base of the gonostylus has protrusion in the form of a small flat visor, which is widely rounded (fig. 54), whereas in D. hirtipes is a triangular tooth forming a curved inward axil, fig. 57, 58). In D. oraniensis this tooth is more rounded apically (fig. 59, 60). Description. Male: holotype—one of the most recently collected specimens which by the size of the body and the color of hairs (fig. 17–19) occupies an intermediate position between the forms that were at the extremes by these parameters (fig. 9–16). Body length ( vertex–T7): 11 mm (paratypes: 9.5–13.5 mm), Mesosoma: W (between tegulae) = 2.5 mm (paratypes: 2.3–2.8 mm). Head: L = 2.5 (paratypes: 2.4–2.6 mm), W = 2.8 mm (paratypes: 2.7– 3.1 mm). Face mainly covered with long white, grayish-white and small admixture (in some paratypes with large admixture or without it) of dirty yellow hairs adjacent to clypeus obscuring integument (fig. 12, 16, 18). Pubescence sparse on vertex near ocelli, with admixture of dark brown and black hairs (fig. 19). Malar distance very short, as in female (see below). Thorax dorsally mostly with dirty yellowish hairs, ventrally and laterally with light white ones (scutum in some paratypes covered with grayish-white hairs), center of scutum with little admixture of dark hairs. Hind tibia (fig. 32–34) intermediate structure between D. hirtipes and D. sinuata. Apical bands on terga consisting of different hairs, from white to dirty yellow (or almost reddish): on T1 widely interrupted, T2 whole (or interrupted narrowly), T3–T6 with continuous apical bands. T1–T3 at base with long erect white and dirty yellow (or grayish-white) hairs, on T4–T6 with such erecte light hairs located at sides of metasoma, otherwise covered with short dark brown or black hairs (on T4–T5 often with admixture of erect light hairs anterior to apical band), T7 covered with black and dark brown, relatively long, appressed hairs with great admixture of light hairs on the sides of terga. All sterna apically lightened, translucent, with narrow band of short, light white (or gray) hairs, on S5 with admixture of reddish-yellow hairs on sides (fig. 11, 15) (certain paratypes, particularly small-sized, with long and densely pubescent sterna). Inner process of gonostylus with short, thick, almost unbranched hairs (fig. 63), except some hairs with small number of short side branches (Fig. 66). All different color variants of body pubescence (same as at females) are observed throughout the range of the species. Lighter forms usually characteristic for smaller specimens. Female (fig. 1 – 8). Body length: 10.5–14 mm. Mesosoma: W (between tegula) = 2.6–3.3 mm. Head: L = 2.8– 3.2; W = 3.3–3.7 (in SIZK was found 2 very small specimens of females, with body length only 8.5 and 9 mm; mesosoma W = 2.0– 2.1 mm; head: L = 2.3–2.4; W = 2.8–3.0 mm; one collected in environs of Kiev, and second in village Zinkiv, Poltava Region of Ukraine). Face mainly with light-colored (white or gray) hairs, with little (fig. 8) or significant (fig. 7) admixture of dark hairs on frons and vertex. Malar distance very short (0.87× shorter than mandible width at the base; fig. 30), as in other representatives of this group. Mesonotum with light gray, yellowish-gray or yellowish-brown (almost reddish) hairs, at middle with little (fig. 2, 4) or greater admixture of dark brown or black hairs (fig. 1, 3). Metasoma with broad, white, apical hair bands on T2–T4 (on T2 often narrowly interrupted), on basal part covered by erecte and decumbent black hairs, T5 with very dense, appressed, dark gray (or grayish-brown) hairs apically, remaining surface covered with long, dark hairs; prepygidial and pygidial fimbria dark gray or grayish-brown. Pygidial plate narrowly elongated at apex, relatively deeply excised apically (fig. 31). Apical parts of sterna slightly enlightened, S3–S5 with continuous and wide apical bands of dark gray-brown (fig. 5) or lighter yellowbrown (fig. 6) plumose erect hairs. Scopa on hind legs golden-yellow, sometimes lighter: grayish-yellow to grayish-white (fig. 1–4). Distribution. The new species occurs from Vinnitsa Region of Ukraine to western Kazakhstan, and from St. Petersburg to the south of the Crimean Peninsula, the Caucasus and Turkey (fig. 67). Most likely, D. morawitzi is rather widespread, but study of European collections is needed to clarify occurrence across the Palaearctic realm. In some regions, for example in western Kazakhstan and in the Crimea, specimens of D. morawitzi are more common than those of D. hirtipes. Floral visitation. Based on the label data, females prefer to collect pollen from plants of the families Asteraceae (mostly Inula britannica, Cichorium intybus and Picris hieracioides) and Plumbaginaceae (Limonium gmelini), although some females were recorded on Apiaceae (Eryngium planum), Boraginaceae (Echium sp.), Caprifoliaceae (Scabiosa isetensis, S. ochroleuca) and Lamiaceae (Lycopus europaeus, Origanum vulgare, Stachys sp.). In addition, single males were collected on plants of Campanulaceae (Campanula sp.) and Convolvulaceae (Convolvulus arvensis). Dasyposa morawitzi has supposedly a wider range of trophic links than D. hirtipes, but these data also need to be clarified because these two species have long been confused. Flight period. End of May (in the Crimea from mid-May) — mid- September. There is one unusually smallsized female specimen (9 mm long) in the collection of SIZK that was collected in Kiev (Ukraine) on 18.04.1954. Such early flight of this species is probably an artifact. Comments. The discovery of this new species r
Fig. 3 in New combination and redescription of Brachyponera mesoponeroides Radchenko, 1993 (Hymenoptera: Formicidae: Ponerinae)
No full textFig. 3. Hypoponera mesoponeroides (Radchenko, 1993) comb. nov., paratype workers. (A) Body in lateral view. (B) Petiole in lateral view. (C) Mandible and gena in lateral view. (D) Metatibial spur. (E), (F) Labels of a paratype. (A), (F) specimen code: IMG20160605-01; (B), (C), (D), (E) IMG20160605-02.Published as part of Dang ,An Van, Yamane, Seiki,, Nguyen, Anh D., & Eguchi, Katsuyuki, 2018, New combination and redescription of Brachyponera mesoponeroides Radchenko, 1993 (Hymenoptera: Formicidae: Ponerinae), pp. 221-229 in Revue suisse de Zoologie 125 (2) on page 226, DOI: 10.5281/zenodo.141420
Tetramorium exile S., Radchenko & Schulz, 2007, New species
No full text5. Tetramorium exile Csösz & Radchenko New species (figs 39-42) Description of Worker (figs 39-42). Morphometric data of the holotype worker: CL: 725; CW: 675; FR: 240; FL: 240; SL: 585; ML: 790; MW: 440; PEW: 215; PEH: 240; NOH: 130; NOL: 130; PEL: 155; PPW: 240; PPL: 160; PPH: 210; SPL: 90; SPSP: 130; EL: 135; EH: 100; ED: 170; Small size, CS 695 [620, 743]. Whole body and appendages brown to dark brown. Head elongated, CL/CW 1.06 [1.04, 1.08], with very feebly convex sides, straight occipital margin and narrowly rounded occipital corners. Eyes small, EYE 0.167 [0.158, 0.175]. Frons very narrow, FR/CS 0.34 [0.33, 0.35], frontal lobes as wide as frons, FL/FR 1.0 [1.0, 1.02]. Scape very long, SL/CS 0.84 [0.82, 0.85], without longitudinal dorsal carina basally, smooth and shiny. Promesonotal dorsum slightly convex, metanotal groove very shallow or completely absent. Propodeal teeth moderately long and acute, directed mainly upwards. Petiolar node trapezoidal in profile, NOH/NOL 1.04 [0.90, 1.16], petiole relatively high, PEH/NOL 1.83 [1.70, 1.96]. General appearance moderately rugose, ground surface microreticulate, dull. Head dorsum longitudinally rugose, ground surface microreticulate, occiput rugoso-reticulate. Alitrunk dorsum and petiolar node rugoso-reticulate ground surface microreticulate, mesopleuron usually rugulose and microreticulate, dorsum of postpetiole longitudinally rugulose and microreticulate. Polygonal striation continuous on 1st gastral tergite (see fig. 8.). Ventral surface of head with several short and few moderately long, straight setae arising posterior to buccal cavity (see fig. 5.). Gynes and Males are unknown. Material examined: HOLOTYPE [[worker]]: AFGHANISTAN - O-Afghanistan, Walang, 2520m, Salangtal, Hindukusch, 29.09.1952. leg. J. Klapperich (HNHM); PARATYPES: 35 [[workers]] from the same nest (3[[workers]] / HNHM, 4[[workers]] / MIZ, 25[[workers]] / NHMW 1[[worker]] / PCAS). Morphometrics: (33 workers were measured). Diagnosis. According to the literature (Collingwood 1961a, 1961b, Pisarski 1967a, 1967b, 1969) and collection materials there are no similar Tetramorium species, described from Afghanistan. Workers of T. exile n. sp. differ from related species by absence of psammophore, relatively small eyes, (EYE, Table 1.), extremely narrow firons, the very long and smooth scape (FR/CS and SL/CS, Table 1.), and by the trapezoidal petiolar node (NOH/NOL and PEH/NOL, Table 1.). Tetramorium exile n. sp. is mostly similar to T. anatolicum sp.n. and T. chefketi, but differs from the latter by the shape of petiolar node: in T. exile n. sp. it is relatively high and short, trapezoidal in profile, while in T. chefketi it is relatively low and longer, cubic in profile, (PEH/NOL and NOH/NOL, Table 1.). The following Discriminant D(3a) function proves the separation between T. exile n. sp. and T. chefketi: D(3a) = 0.052 PPW -0.057 FL -0.081 NOL +14.667, T. exile n. sp. D(3a) = +3.418 ±0.901 [+2.169, +4.756] (n=33), T. exile n. sp., holotype D(3a) = +3.023, p<0.001, T. anatolicum n. sp. D(3a) = +3.164 ±0.796 [+1.746, +4.606] (n=38), T. anatolicum n. sp., holotype D(3a) = +3.246, p<0.001, T. chefketi D(3a) = -3.417 ±1.004 [-5.680, -0.524] (n=107), T. chefketi lectotype D(3a) = -4.179, p<0.001; T. sarkissiani lectotype D(3a) = -2.809, p<0.001; T. turcomanicum syntype worker D(3a) = -4.169, p<0.001. T. exile n. sp. differs from T. anatolicum n. sp. by its longer scape and narrower firons (SL/CS and FR/CS, Table 1.). The following Discriminant D(2a) function provides the separation between T. exile n. sp. and T. anatolicum n. sp.: D(2a) = 0.081 SL - 0.151 FL - 7.652 T. exile n. sp. D(2a) = +2.683 ±0.762 [+1.193, +3.493] (n=33), T. exile n. sp., holotype D(2a) = +3.493, p<0.001, T. anatolicum n. sp. D(2a) = -2.683 ±1.064 [-4.682, -0.042] (n=38), T. anatolicum n. sp., holotype D(2a) = -3.117, p<0.001. For further combination of morphometric characters see Table 1-2. Distribution. Known from the type locality only. Etymology. This adjective [exilis / -e; exile (neutrum) = slender] refers to the elongate body of this species.Published as part of Csösz S., Radchenko, A. & Schulz, A., 2007, Taxonomic revision of the Palaearctic Tetramorium chefketi species complex (Hymenoptera: Formicidae)., pp. 1-38 in Zootaxa 1405 on pages 22-2
Fig. 2 in New combination and redescription of Brachyponera mesoponeroides Radchenko, 1993 (Hymenoptera: Formicidae: Ponerinae)
No full textFig. 2. Hypoponera mesoponeroides (Radchenko, 1993) comb. nov., paratype workers. (A) Head in full-face view. (B) Head in lateral view. (C) Mesosoma in lateral view. (D) Body in dorsal view. (B), (C), (D) specimen code: IMG20160605-01; (A) IMG20160605-02.Published as part of Dang ,An Van, Yamane, Seiki,, Nguyen, Anh D., & Eguchi, Katsuyuki, 2018, New combination and redescription of Brachyponera mesoponeroides Radchenko, 1993 (Hymenoptera: Formicidae: Ponerinae), pp. 221-229 in Revue suisse de Zoologie 125 (2) on page 225, DOI: 10.5281/zenodo.141420
Fig. 4 in New combination and redescription of Brachyponera mesoponeroides Radchenko, 1993 (Hymenoptera: Formicidae: Ponerinae)
No full textFig. 4. Hypoponera mesoponeroides (Radchenko, 1993) comb. nov., nontype worker. (A) Head in full-face view. (B) Body in lateral view. (C) Antenna. (D) Petiole. (E) Body in dorsal view. (A), (B), (C), (D) colony code: AD16CP60, specimen code: CAP20171220-01; (E) AD16CP58, CAP20170714-01.Published as part of Dang ,An Van, Yamane, Seiki,, Nguyen, Anh D., & Eguchi, Katsuyuki, 2018, New combination and redescription of Brachyponera mesoponeroides Radchenko, 1993 (Hymenoptera: Formicidae: Ponerinae), pp. 221-229 in Revue suisse de Zoologie 125 (2) on page 227, DOI: 10.5281/zenodo.141420
Dasypoda schwarzi Radchenko et Michez 2022, sp. nov.
No full textDasypoda schwarzi Radchenko et Michez sp. nov. Type material. Holotype ♂ with the printed label: " Tunesien, Thala 20 km S [35°23'N 8°41'E], 12.4.1981, leg. Max. Schwarz " (MSAA). Paratypes: 2 ♂ with the printed label: " Morocco, 40 km S Guercif [33°53'N 3°22'W], 15.- 17.5.1995, Ma. Halada lgt." (UMONS); Tunesien, Thala 20 km S, 12.4.1981, leg. J. Gusenleitner (OÖLM). Etymology. The species is named after Maximilian Schwarz, an authority on bee systematics. Subgeneric affinity. Dasypoda schwarzi sp. nov. can be regarded as a representative of the subgenus Microdasypoda Michez (in Michez et al. 2004b) due to the combination of its diagnostic features. First, representatives of the subgenus Microdasypoda are characterized by a relatively small body size of approximately 7-12 mm (contrast Megadasypoda and many species of Dasypoda s.str. and Heterodasypoda which are significantly larger). Then, the subgenus is most significantly distinguished from all other Dasypoda subgenera by the structure of the male genitalia and S7: the gonostylus of Microdasypoda is either unilobed with one basal tooth, or bilobed without a membranous structure that connects these lobes (contrast Dasypoda s.str. in which the gonostylus is bilobed with such membranous structure, and Heterodasypoda and Megadasypoda in which it is trilobed), and the S7 does not bear latero-apical processes, which are characteristic of all other subgenera of Dasypoda. The previously cited apomorphies of Microdasypoda should however always be regarded with a combination of other characters that are individually shared by other subgenera, but not all. First, the malar space of Microdasypoda is always much shorter than the pedicel (contrast most Megadasypoda) and their maxillary palpi and galea are of a sub-equal length (contrast Dasypoda s.str. and Megadasypoda). The nervulus (cu-v) of Microdasypoda is strongly antefurcal (contrast most Megadasypoda). The apex of S6 of Microdasypoda has a long, dense pubescence (contrast Dasypoda s.str. and Megadasypoda which have short, sparse ones). Microdasypoda lack lateral hooks at the basal half of S8 (contrast Dasypoda s.str. and some Megadasypoda). The external lobe of the gonostyle of Microdasypoda is not lanceolate (contrast Megadasypoda and some Heterodasypoda), and the inner lobe of the gonostyle of Microdasypoda does not present a scaly surface (contrast Heterodasypoda in which this scaly surface is clearly visible). The new species described in this work, Dasypoda schwarzi sp. nov., clearly corresponds to all the features that characterize the subgenus Microdasypoda. Description. Male (Figs 1, 2): Holotype. Body black, its length (vertex to T7) 11.9 mm (paratypes: 11.5 mm and 10.5 mm). Head slightly wider than long (Fig. 6): L = 2.9 mm (paratypes: 2.8 mm and 2.7 mm), W = 3.4 mm (paratypes: 3.2 and 3.1 mm, respectively); clypeus densely punctured by oblique downward hair-bearing punctures separated by 0.5–1 puncture diameter with abundant long, dense, white adjacent hair obscuring underlying surface, apex of clypeus with narrow depressed impunctate band. Paraocular area irregularly punctate, punctures separated by 1–3 puncture diameter. Paraocular and supraclypeal areas and basal part of frons with long erect pale-white plumose hairs, intermixed on upper part near vertex with dark brown hair. Genal area, apical parts of vertex, and occiput with long, erect pale-white pubescence intermixed with dark brown hair. Central part of frons with narrowly depressed, median stripe, upper part of frons medially polished, unpunctured; laterally irregularly, sparsely punctate and very weakly shagreened. Malar space narrow, 4 times as broad as long, (W = 0.43 mm, L = 0.11 mm) (Fig. 4). Antennal scape anteriorly with long erect white hair. Flagellum moderately long, first flagellomere 1.7 times as long as its apex width and 1.25 times longer than second one (Fig. 3, 44). Labrum polished and shining, apical edge with long dense fringe of golden-yellow or pale-yellow thick hair. Glossa short: L = 0.65 mm, widened at base: W (at base) = 0.23 mm (Fig. 5). 2 nd and 3 rd segments of labial palpi apically swollen. Galea weakly shagreened with very sparse and superficial punctation (Fig. 9); maxillary palpi subequal in length with galea. Mesosoma: W (between tegulae) = 2.9 mm (paratypes: 2.9 mm and 2.7 mm). Scutum with hair-bearing punctation, punctures separated by 1–1.5 puncture diameters with exception of sparsely punctate (3–4 puncture diameters) central part (Fig. 7); mesosoma ventrally covered with very long, erect white hair, laterally and dorsally with pale-yellow hair, on scutum intermixed with shorter dark brown hairs; scutellum and metanotum with rufous hair. Propodeal triangle finely shagreened, basally with narrow transverse rugosity; external margins well-defined by lateral lines (Fig. 8). Propodeum laterally very weakly shagreened and sparsely punctured with long erect yellow hair. Wings hyaline with light brown tint; tegulae, venation and stigmata dark brown. Nervulus (cu-v) antefurcal. Legs with long pale-yellow pubescence, apart from greyish pubescence of inner side of tibiae and dense short ginger-yellow pubescence of inner side of metabasitarsus. Metasoma (Figs 10, 11): L = 5.8 mm (paratypes: 5.7 and 5.3 mm); W = 4.3 mm (paratypes: 4.3 mm and 3.8 mm, respectively). T1 basal part with oblique, moderately sparse punctation with long, semierect hair; separated by 1–3 puncture diameters; marginal part of T1 depressed, almost unpunctured and very thinly and densely transversely wrinkled with sparse superficial weakly noticeable punctation bearing short brown hair (Fig. 12); narrow apical margin of all terga light, translucent. Basal parts of T2–T5 moderately dense punctured, punctures separated by 1–2 puncture diameter with long semi-adjacent yellow hair, underlying surface with sparse short brown hair. Apical margins of T2–T5 slightly depressed with entire bands of very short dense white hairs covered from above by apical parts of long yellow hair. Basal part of T6 densely punctate by hair-bearing puncture with semi-adjacent long yellow hair (in holotype specimen this pubescence erased), laterally with long erected pale-yellow hair intermixed with sparse brown hair; apical margin of this tergum polished, impunctate. Basal parts of sterna moderately sparse punctured (denser on S2–S4 and sparser on S5) by very small oblique hair-bearing punctures with very short dark brown hair; underlying surface weakly shagreen (Fig. 13). Sternal margins with sparse, yellow hair fringes longer laterally and shorter centrally, narrowly interrupted at the centre of S1–S4, and widely on S5. Marginal parts of S1–S5 very sparse punctured, polished, its narrow apical margin light, translucent, on S2–S5 medially widely roundly notched. S6 marginal part medially with wide projection deeply triangularly emarginated on apex (Figs 14, 15). S7 (Fig. 17) latero-apically with short, weakly sclerotized semicircular projections bent towards inner side of sternum (Fig. 18). S8 relatively short, stumpy, deeply semicircularly concave laterally before widened basal part (Figs 19, 20); apex of S8 posterodorsal part with whole transverse carina not notched or emarginated in centre (Fig. 16). Genitalia with bilobed gonostyli (Figs 21–34). Inner lobe of gonostylus moderately wide and long, widened in a circular shape at apex and covered with sparse long setae on surface facing penis valves and on margins (Figs 33, 34, 85, 86); external lobe wide, its apex obliquely truncated (Fig. 32), ventral part narrowly, triangularly elongated, inner surface with several hairs, each inserted into crater-like raised alveoli (Figs 35–37). Female unknown. Distribution. Northwest Africa: Atlas Mountains area (Morocco and Tunisia) (Fig. 89). Floral visitation. Pollen grains of the Malvaceae plant family were mainly found on the body of the paratype (Fig. 38), and only a few grains of Asteraceae. Species diagnosis. The main significant differences D. schwarzi sp. nov. from the other known species of Microdasypoda concern the structure of genitalia. Other Microdasypoda species have single-lobed gonostyli with a spine-like tooth on the inner basal part (in D. cingulata, D. crassicornis and D. iberica; Figs 82–84) or bilobed gonostyli with a thickened internal lobe that has a very dense, long pubescence on surface facing penis valves (in D. brevicornis; Figs 75, 80, 81). Unlike other representatives of this subgenus, the genitalia of D. schwarzi have bilobed gonostyli with the inner lobes that are laterally flattened and dorso-ventrally widened in a circular shape at the apical part, and with sparse setae on the surface facing penis valves (Figs 85, 86). By the structure of its genitals and sterna, D. schwarzi is most similar to D. brevicornis, sharing the same structure of S6, S7, dorso-apical part of S8, glossa and galea. In both species, unlike other Microdasypoda, the setae along the lower edge of the galea are longer in the apical half than in the basal half where they are very short and poorly visible (Figs 9, 39) (Michez et al. 2004b indicated the absence of setae in the basal half of galea, probably due to the relatively lower resolution of optics used at that time). Dasypoda schwarzi can also be separated from D. brevicornis by the following features: the external lobe of the gonostyle in D. schwarzi with a strongly oblique apex (Figs 21, 23, 29, 32); the digitus of its volsella shorter and wider (Fig. 28); the 1 st flagellomere relatively longer (Figs 43, 44); the mandibular palpus with thicker segments (Figs 5, 6, 9); the apical projection of the S6 with right angles on the outer edges (rather than rounded lateral corners as in D. brevicornis) (Figs 55, 59); the lateral margins of the S8 deeply semicircularly notched before the apodemes (straight-beveled lateral margins of S 8 in D. brevicornis) (Figs 65, 69); and the lower half of the face with light pubescence without dark brown or black hair (the latter being typical in D. brevicornis). Remarks on the comparative morphology of Microdasypoda. The use of modern optics with a higher resolution made it possible to refine the original diagnosis of the subgenus Microdasypoda Michez (Michez et al. 2004b). In particular, the outer lobe of the gonostyle may have a lanceolate shape (in D. crassicornis), the S 7 in almost all species of the subgenus (with the exception of D. iberica) has a very small almost non-chitinous lateroapical outgrowths (Fig. 18); the dorso-apical part of S8 has an entire transverse carina or two partly or completely separated teeth (Figs 70–74); the S6 apicoventrally may have not only brown pubescence (in most species), but can also present a complete silver-white pubescence (in D. iberica; Fig. 53; Radchenko et al. 2019). This updated clarification of the diagnostic characters of all species belonging to the subgenus Microdasypoda, as well as the detailed morphological characters of the male of D. iberica that was recently published (Radchenko et al. 2019) made it possible to compose a corrected and updated key for the males.Published as part of Radchenko, Vladimir G., Ghisbain, Guillaume & Michez, Denis, 2022, A new bee species of the genus Dasypoda Latreille (Hymenoptera, Apoidea) from Northwest Africa with comparative remarks on the subgenus Microdasypoda Michez, pp. 74-86 in Zootaxa 5188 (1) on pages 75-80, DOI: 10.11646/zootaxa.5188.1.4, http://zenodo.org/record/708741
Fig. 5 in New combination and redescription of Brachyponera mesoponeroides Radchenko, 1993 (Hymenoptera: Formicidae: Ponerinae)
No full textFig. 5. Hypoponera mesoponeroides (Radchenko, 1993) comb. nov., nontype queen, colony code: AD16CP57; specimen code: CAP20170716-01. (A) Head in full-face view. (B) Body in dorsal view. (C) Antenna. (D) Body in lateral view. (E) Compoud eye. (F) Mesosoma in lateral view.Published as part of Dang ,An Van, Yamane, Seiki,, Nguyen, Anh D., & Eguchi, Katsuyuki, 2018, New combination and redescription of Brachyponera mesoponeroides Radchenko, 1993 (Hymenoptera: Formicidae: Ponerinae), pp. 221-229 in Revue suisse de Zoologie 125 (2) on page 228, DOI: 10.5281/zenodo.141420
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