1,355,752 research outputs found
Loomisiella pylei Shear & Richart & Wong 2020, new species
No full textLoomisiella pylei, new species Figs. 244–246 Types: Male holotype, male and female paratypes from WASHINGTON: Wahakiakum Co., Lower Hendrickson Canyon, 46.3693°N, - 123.6657°W, elev. 30 m, old-growth forest, collected 23 January 2004, by W. Leonard, M. Leonard, C. Richart, R. M. Pyle, and K. Novoselić. Diagnosis: See under preceding species. Etymology: The species name honors Robert Michael Pyle, a notable lepidopterist, author, conservation advocate, and as is this species, denizen of the Willapa Hills. Description: Male: Length, 5.0 mm. Fourteen ocelli in oval-triangular group. Metazonital shoulders poorly developed; segmental setae curved, acute. Legpairs one, two reduced, pairs three, four encrassate, pairs five to seven near same size as postgonopodal legs; femora three, four with basal femoral knobs acute, posteriorly directed.Anterior gonopods (ag, Figs. 244, 245) with large anterior sternum, widely separated, curving lateral to posterior gonopod coxites, apically decurved but without any apical expansion, small lateral branch tightly appressed to gonopod (free in Fig. 245 due to compression on microscope slide). Posterior gonopod telopodites reduced but relatively larger than in L. evergreen; posterior gonopod coxites (pgc, Figs. 244, 245) short, broad, anteriorly cupped, with broad, lamellate process mesally, basal pseudoflagellum originating posteriorly, two processes distolaterally, more lateral process composed of filaments (seen as separate in Fig. 246 due to compression on microscope slide), mesal process curved, apically fimbriate. Coxae 10, prefemora 11 as usual. Female 5.2 mm long, nonsexual characters as in male. Distribution: WASHINGTON: Lewis Co., FS-25 8.5 mi S of Randle, 46.4409°N, - 121.9966°W, elev. 330 m, 21 December 2003, W. Leonard, C. Richart m f; same as previous but 6 December 2003, m. Pacific Co., Cement Creek, SR-401 2.5 mi S of Parpala Road in Naselle, 46.3341°N, - 123.8002°W, elev. 30 m., 15 January 2006, W. Leonard, C. Richart, from the litter of a riparian forest including Alnus rubra, Picea sitchensis, Sambucus cf. caerulea, and Polystichum munitum, m ff; Trap Creek Road 1.7 mi S of SR-6, Trap Creek Basin, 46.5403°N, - 123.6297°W, elev. 60 m., 19 November 2005, W. Leonard, C. Richart, mm ff. Wahkiakum Co., only known from the type locality.Published as part of Shear, William A., Richart, Casey H. & Wong, Victoria L., 2020, The millipede family Conotylidae in northwestern North America, with a complete bibliography of the family (Diplopoda, Chordeumatida, Heterochordeumatidea, Conotyloidea), pp. 1-78 in Zootaxa 4753 (1) on page 69, DOI: 10.11646/zootaxa.4753.1.1, http://zenodo.org/record/398378
Bifurcatella germania Shear & Richart & Wong 2020, new species
No full textBifurcatella germania, new species Figs. 220, 221, 224, 225 Types: Male holotype from WASHINGTON: Cowlitz Co., Germany Creek Road 5.5 mi N of Stella at SR-4, elev. 140 m., 46.2593°N, - 123.1344°W, collected 22 November 2003, by W. Leonard, C. Richart; male paratype from Germany Creek Road 5.3 mi N of Stella at SR-4, elev. 140 m., 46.2565°N, - 123.1356°W, collected 11 November 2004, by W. Leonard and C. Richart. Diagnosis: The posterior gonopod coxites are similar to those of the preceding species but the anterior gonopods are quite different, lacking the midlength shelf and sharp taper, and with three large subterminal teeth. Etymology: This species takes its adjectival name from Germany Creek, where the holotype and paratype were collected. Description: Male holotype: Length, 9.0 mm. Nineteen ocelli in triangular group. Metazonites with well-developed shoulders, segmental setae long, curved, acute. Color as described for B. pacifica. Legpairs one and two reduced, pairs three to five enlarged, sixth pair somewhat smaller, seventh pair approximately same size as postgonopodal legs; prefemora three to five with conspicuous knobs. Anterior gonopods (Figs. 220, 222) with mesal branches less than one-third size of lateral branches, broadly flattened in lateral view; lateral branches curved, with large, wing-like expansion anteriorly, bearing three triangular subterminal teeth. Posterior gonopod coxites (Figs. 221, 223) with pseudoflagellar branch tightly reflexed, not strongly tapering, two large fimbriate branches, lateralmost embraces pseudoflagellum, mesal branch subtends small pore. Tenth coxae and eleventh prefemur as usual. Females not collected. Distribution: Known only from the two localities along Germany Creek, which are about 340 meters apart.Published as part of Shear, William A., Richart, Casey H. & Wong, Victoria L., 2020, The millipede family Conotylidae in northwestern North America, with a complete bibliography of the family (Diplopoda, Chordeumatida, Heterochordeumatidea, Conotyloidea), pp. 1-78 in Zootaxa 4753 (1) on page 60, DOI: 10.11646/zootaxa.4753.1.1, http://zenodo.org/record/398378
Bifurcatella germania Shear & Richart & Wong 2020, new species
No full textBifurcatella germania, new species Figs. 220, 221, 224, 225 Types: Male holotype from WASHINGTON: Cowlitz Co., Germany Creek Road 5.5 mi N of Stella at SR-4, elev. 140 m., 46.2593°N, - 123.1344°W, collected 22 November 2003, by W. Leonard, C. Richart; male paratype from Germany Creek Road 5.3 mi N of Stella at SR-4, elev. 140 m., 46.2565°N, - 123.1356°W, collected 11 November 2004, by W. Leonard and C. Richart. Diagnosis: The posterior gonopod coxites are similar to those of the preceding species but the anterior gonopods are quite different, lacking the midlength shelf and sharp taper, and with three large subterminal teeth. Etymology: This species takes its adjectival name from Germany Creek, where the holotype and paratype were collected. Description: Male holotype: Length, 9.0 mm. Nineteen ocelli in triangular group. Metazonites with well-developed shoulders, segmental setae long, curved, acute. Color as described for B. pacifica. Legpairs one and two reduced, pairs three to five enlarged, sixth pair somewhat smaller, seventh pair approximately same size as postgonopodal legs; prefemora three to five with conspicuous knobs. Anterior gonopods (Figs. 220, 222) with mesal branches less than one-third size of lateral branches, broadly flattened in lateral view; lateral branches curved, with large, wing-like expansion anteriorly, bearing three triangular subterminal teeth. Posterior gonopod coxites (Figs. 221, 223) with pseudoflagellar branch tightly reflexed, not strongly tapering, two large fimbriate branches, lateralmost embraces pseudoflagellum, mesal branch subtends small pore. Tenth coxae and eleventh prefemur as usual. Females not collected. Distribution: Known only from the two localities along Germany Creek, which are about 340 meters apart.Published as part of Shear, William A., Richart, Casey H. & Wong, Victoria L., 2020, The millipede family Conotylidae in northwestern North America, with a complete bibliography of the family (Diplopoda, Chordeumatida, Heterochordeumatidea, Conotyloidea), pp. 1-78 in Zootaxa 4753 (1) on page 60, DOI: 10.11646/zootaxa.4753.1.1, http://zenodo.org/record/398378
Bollmanella washingtonensis Shear & Richart & Wong 2020, new species
No full textBollmanella washingtonensis, new species Figs. 115–117 Types: Male holotype, male paratype and two female paratypes from WASHINGTON: Mason Co., W Beerbower Road, at W Schafer State Park Road, elev. 45 m., 47.1001°N, - 123.4706°W, collected 31 December 2004, by W. Leonard. Diagnosis: Unlike any other species of Bollmanella, B. washingtonensis has pregonopodal legpairs three to seven enlarged, and pairs five through seven with femoral knobs. Etymology: The species name notes that this species is endemic to Washington. Description: Male paratype: Length, 10.5 mm. 18–20 ocelli in rounded, triangular eyepatch. Color pale tan, mottled light purplish brown. Segmental shoulders prominent, metazonital setae long, acute. Pregonopodal legpairs three to seven strongly encrassate, legpairs five, six and seven with femoral knobs. Anterior gonopods (Fig. 115) slender, widely divergent, prominently curved near midlength, with acute subapical process as in the foregoing species. Posterior gonopods (Fig. 117) with relatively short telopodites, coxites (Figs. 116, 117) two-branched, posterior branch shorter, tapering, curved laterad, anteromesal branch distally bifurcate, one terminal four times broader than other, slightly curved. Legpairs 10 and 11 as usual for genus. Female 11 mm long, nonsexual characters as in male. Distribution: WASHINGTON: Grays Harbor Co., Middle Satsop Road 2 mi N of Brady near Satsop River, elev. 20 m., 47.0245°N, - 123.5196°W, under maple leaves, 3 January 2003, W. Leonard et al., mm; 10 mi N of US- 12, East band of Wynoochee River, ca. 47.0920°N, - 123.6798°W (estimated coordinates measured 10.0 mi N of US- 12 along Wynoochee Valley Rd), under maple leaves, 3 January 2003, W. Leonard et al., m f; Canyon River, 6 mi W, 1.25 mi N of Matlock, ca. 47.2585°N, - 123.5271°W, in maple and fern litter, 17 January 2003, W. Leonard et al., mm ff; Garrard Road at Weyerhaeuser D-line, elev. 45 m., ca. 46.8041°N, - 123.3033°W, 7 February 2005, W. Leonard, mm ff. Thurston Co., Capitol State Forest, 2 mi S of Rock Candy Mountain, ca. 46.9809°N, - 123.1090°W (estimated coordinates taken from 2 air mi S of Rock Candy Mountain summit), 10 December 2002, W. Leonard et al., mm; Summit Lake, elev. 138 m., 47.0526°N, - 123.1304°W, 18 December 2004, W. Leonard, m f; McAllister Springs S of Steilacoom Rd SE, elev. 50 m., 47.0472°N, - 122.7286°W, 7 February 2004, W. Leonard, C. Richart, from the litter and woody debris of a mixed forest including Acer macrophyllum, Alnus rubra, Pseudotsuga menziesii, Thuja plicata, Polystichum munitum, and Mahonia nervosa, mm f; St. Martin’s College Campus, Woodland Creek, elev. 92 m., 47.0416°N, - 122.8134°W, 12 February 2003, W. Leonard, mm ff. King Co., I-90 at Snoqualmie River, Twin Falls/Iron Horse trailhead, elev. 370 m., 47.4410°N, - 121.6687°W, 25 February 2004, W. Leonard, C. Richart, m f; Along Holder Creek, SR-18 1.5 mi N of the Issaquah exit near Hobart, elev. 310 m., 47.4522°N, - 121.9535°W, 25 February 2004, W. Leonard, C. Richart, m f; Ollalie State Park, elev. 380 m., 47.4364°N, - 121.6594°W, 25 February 2004, W. Leonard, C. Richart, m f; FR-9034 near Granite Mountain Trailhead, Mt. Baker-Snoqualmie National Forest, elev. 580 m., 47.3977°N, -121.4855, 27 October 2007, W. Leonard, mm f; same as previous but 26 March 2004, W. Leonard, C. Richart, m; same as previous but 25 October 2003, m. Cowlitz Co., SR-504 2.1 mi E of Toutle, elev. 152 m., 46.3483°N, - 122.7066°W, 1 March 2004, W. Leonard, C. Richart, from the litter and woody debris of a mixed forest including Alnus rubra, Acer macrophyllum, and Polystichum munitum, m f. Jefferson Co., Falls View Campground, Olympic National Forest, US-101 7.8 mi NW of Brinnon, elev. 150 m., 47.7893°N, - 122.9255°W, 22 February 2003, W. Leonard, m f. Pacific Co., Ellsworth Creek Nature Society Preserve, old growth forest, 14 March 2003, elev. 25 m., 46.3981°N, - 123.8898°W, W. Leonard et al., m ff; same as previous but 23 November 2003, W. Leonard et al., mm; Trap Creek Road/B-line 1.1 mi S of SR-6, elev. 58 m., 46.5432°N, - 123.6151°W, 19 November 2005, W. Leonard, C. Richart, mm ff; same as previous but 1.9 mi S of SR-6, elev. 65 m, 46.5401°N, - 123.6296°W. Notes: This widespread species could be considered as a Taiyutyla, since the pregonopodal leg modifications do not conform to the other species of Bollmanella. Nevertheless, the form of the gonopods indicates that it belongs in the latter genus.Published as part of Shear, William A., Richart, Casey H. & Wong, Victoria L., 2020, The millipede family Conotylidae in northwestern North America, with a complete bibliography of the family (Diplopoda, Chordeumatida, Heterochordeumatidea, Conotyloidea), pp. 1-78 in Zootaxa 4753 (1) on pages 32-34, DOI: 10.11646/zootaxa.4753.1.1, http://zenodo.org/record/398378
Complicatella complicata Shear & Richart & Wong 2020, new combination
No full textComplicatella complicata (Shear, 1974), new combination Bollmanella complicata Shear 1974, p. 146. Figs. 192–193 New Records: WASHINGTON: Thurston Co., Black Lake and Belmore Road at 66 th Avenue, Tumwater, elev. 60 m., 46.9889°N, - 122.9659°W, 19 November 2003, W. Leonard, mm; Hospital Creek, above confluence with Skookumchuck River, 5 horiz mi S, 3 horiz mi E of Vail, elev. 245 m., 46.7732°N, - 122.5855°W, 15 December 2003, W. Leonard, K. McAllister, mm ff; Rainier Road SE, 3.0 mi NW of Military Road SE, Fort Lewis Military Reservation, elev. ca. 100 m., ca. 46.9351°N, - 122.7600°W, 29 November 2003, W. Leonard, m; The Evergreen State College campus, Beach Trail N of Driftwood Road, Olympia, elev. 30 m., 47.0791°N, - 122.9776°W, 26 January 2003, W. Leonard, mm ff; same as previous but 16 March 2003, mm; same as previous but 23 March 2003; same as previous but 22 December 2003, mm ff; Priest Point Park, Olympia, elev. ca. 65 m., ca. 47.07°N, - 122.89°W, 5 January 2003, W. Leonard, mm f; same as previous but 2 February 2003, m; same as previous but 14 November 2004, mm ff; along Rock Candy Mtn. Road, Capitol State Forest, elev. ca. 300 m., 47.0256°N, - 123.0735°W, 18 December 2004, W. Leonard, m. Grays Harbor Co., Porter Creek Campground, Capitol State Forest, elev. 90 m., 46.9779°N, - 123.2565°W, 24 January 2005, W. Leonard, m. Cowlitz Co., Germany Creek Road 5.5 mi N of Stella at SR-4, elev. 110 m., 46.2593°N, - 123.1344°W, 22 November 2003, W. Leonard, C. Richart, mm; same as previous but 8 December 2003, m f; same as previous but 11 November 2004, mm ff. Jefferson Co., along Dosewallips River, Dosewallips Road 7.0 mi W of US-101, Olympic National Forest, elev. 115 m., 47.7326°N, - 123.0201°W, 19 November 2003, W. Leonard, m; Falls View Campground, US-101 7.8 mi NW of Brinnon, elev. 150 m., 47.7893°N, - 122.9255°W, 22 February 2003, W. Leonard, m f. Mason Co., SR-119 6 mi W of US-101 at Hoodsport, elev. 310 m., 47.7326°N, - 123.2097°W, 7 February 2003, W. Leonard, mm. King Co., FR-9034 near Granite Mountain Trailhead, Mt. Baker-Snoqualmie National Forest, elev. 580 m., 47.3977°N, - 121.4855°W, W. Leonard, 27 October 2007, mm f; same as previous but 26 March 2004, W. Leonard, C. Richart, m; same as previous but 25 October 2003. Lewis Co., 604 Roswell Road, Centralia, elev. 65 m., 46.7227°N, - 122.9452°W, 25 January 2004, C. Richart, W. Leonard, mm ff; SE end of Riffle Lake, elev. 250 m., 46.4644°N, - 122.1705°W, 9 April 2004, W. Leonard, C. Richart, m; Slide Creek, Weyerhaeuser 4000 Rd 0.8 mi SW of Pe-Ell-McDonald Rd., elev. 190 m., 46.5250°N, - 123.1913°W, 4 December 2004, W. Leonard, C. Richart, mm ff; same as previous but 3 January 2005, W. Leonard; Stillman Basin, 1.8 mi on 4000 Weyerhaeuser from Pe-Ell McDonald Rd., elev. 290 m., 46.5145°N, - 123.2015°W, 4 December 2004, W. Leonard, C. Richart, from the litter and woody debris of a mixed forest including Thuja plicata, Alnus rubra, Rubus spectabilus, Pseudotsuga menziesii, and Polystichum munitum, m f; North Fork of Newaukum River, below confluence with Middle Fork, elev. 88 m., 46.6046°N, - 122.8481°W, 29 December 2004, W. Leonard, m. Notes: This species was described from 1 mi W of Bayshore and 4 mi N of Shelton, Mason Co., Washington, and heretofore was known only from the type locality. New collecting revealed a much wider distribution, detailed above and in the supplemental KML map, and show this species to be the most widely distributed and most commonly collected conotylid in western Washington. All these populations are linked by identical anterior gonopods (Fig. 192), but the posterior gonopod coxites (i.e., Fig. 193) are so complicated that it is difficult to view them from exactly the same angle and make comparisons. Thus it is not possible really to be sure that all collections belong to the same species. This can be worked out later with genetic data. Descriptions of new species of ComplicatellaPublished as part of Shear, William A., Richart, Casey H. & Wong, Victoria L., 2020, The millipede family Conotylidae in northwestern North America, with a complete bibliography of the family (Diplopoda, Chordeumatida, Heterochordeumatidea, Conotyloidea), pp. 1-78 in Zootaxa 4753 (1) on pages 48-49, DOI: 10.11646/zootaxa.4753.1.1, http://zenodo.org/record/398378
Vancouvereuma shawi Shear & Richart & Wong 2020
No full textVancouvereuma shawi (Shear, 2004) Figs. 102–104 Taiyutyla shawi Shear, 2004:16. Type locality: BRITISH COLUMBIA: Vancouver Island, Lower Hanging Sump Cave, 16 km SW of Port McNeil, male holotype, juvenile male paratype (VMNH). New record: WASHINGTON: Grays Harbor Co., unnamed tributary of Wynoochee River, Olympic National Forest, elev. 457 m., 47.4852°N, - 123.5231°W, 28 September 2003, W. Leonard, m f. Notes: Vancouvereuma shawi has also been collected in Thurston Co., Washington (Shear 2004), and females resembling this species have been collected as far north as southern Alaska.Published as part of Shear, William A., Richart, Casey H. & Wong, Victoria L., 2020, The millipede family Conotylidae in northwestern North America, with a complete bibliography of the family (Diplopoda, Chordeumatida, Heterochordeumatidea, Conotyloidea), pp. 1-78 in Zootaxa 4753 (1) on pages 26-28, DOI: 10.11646/zootaxa.4753.1.1, http://zenodo.org/record/398378
Taiyutyla amicitia Shear & Richart & Wong 2020, new species
No full textTaiyutyla amicitia, new species Figs. 27–36 Types: Male holotype, 16 male and 8 female paratypes from IDAHO: Kootenai Co., Beauty Creek, Idaho Panhandle National Forests, 47.6031°N, - 116.6453°W, collected 8 November 2004, by C. Richart, W. Leonard, J. Baugh. Diagnosis: Taiyutyla amicitia is unique in having femoral knobs only on legpairs four and five. Etymology: The species is named to commemorate the friendship that formed over millipede species discovery (L., amicitia, a noun in apposition) between WAS, CHR and William P. Leonard. Description: Paratype male from Beauty Creek: Length, 10.0 mm. Twenty-two ocelli in triangular eyepatch. Metazonites with low shoulders on all trunk rings from three to 25. Color light brown, marked darker purplish brown, markings more distinct anteriorly. Legpairs one and two reduced, three to seven enlarged, mesal knobs present only on femora four and five, mesal and large on both (Figs. 32–36). Anterior gonopods (Figs. 27, 28, 36) simple, with short, triangular process subdistally. Posterior gonopod coxites (Fig. 29, 31) complicated, with T-shaped branch (T, Fig. 29) originating subdistally, a strong, triangular tooth at its base; this branch with two long, thin processes, the more distal one set with small teeth. Terminus of coxite strongly hooked. Legpair 10 coxae of normal size, with small glands, legpair 11 femora with long, thin, dorsally directed knobs. Female 14.5 mm long, similar to male in nonsexual characters. Distribution: IDAHO: Shoshone Co., Hobo Cedar Grove Botanical Area, Idaho Panhandle National Forests, elev. 1295 m., 47.0880°N - 116.1152°W, 14 October 2006, W. Leonard, C. Richart, A. Fusek, Habitat: oldgrowth Tsuga heterophylla, Thuja plicata, Abies grandis, under litter and woody debris, m f (CAS). Notes: This species, the first Taiyutyla from Idaho, is syntopic with Brunsonia pulchra, n. sp. (see below) at Beauty Creek, the type locality for both of them. The original collection was mixed, and after the two species were sorted out by size, the gonopod differences became apparent. Taiyutyla amicitia is perhaps the Taiyutyla species closest in gonopod form (particularly the posterior gonopod coxites) to some Bifurcatella, n. gen. species, but the large femoral knobs on legpair five and the somewhat larger body size argue for including it in Taiyutyla.Published as part of Shear, William A., Richart, Casey H. & Wong, Victoria L., 2020, The millipede family Conotylidae in northwestern North America, with a complete bibliography of the family (Diplopoda, Chordeumatida, Heterochordeumatidea, Conotyloidea), pp. 1-78 in Zootaxa 4753 (1) on page 13, DOI: 10.11646/zootaxa.4753.1.1, http://zenodo.org/record/398378
Bifurcatella hoh Shear & Richart & Wong 2020, new species
No full textBifurcatella hoh, new species Figs. 210, 211, 214, 215 Types: Male holotype, female paratype from WASHINGTON: Jefferson Co., Alder Creek at Hoh River Road, elev. 125 m., 47.8301°N, - 124.2272°W, collected 14 December 2003, by W. Leonard, C. Richart. Diagnosis: The unique feature of this species is the thin mesobasal process (mbp, Fig. 215) of the posterior gonopod coxite, found in no others; further, the apical process of the coxite is sharply curved anteromesally. The mesal branch of the anterior gonopod is more than 2/3 the length of the lateral branch and is straight and blunt. Etymology: Named for the Hoh River of Olympic National Park; a noun in apposition. Description: Male holotype: Length, 11.0 mm. Ocelli 22 in triangular patch. Metazonites with moderately well-developed shoulders, segmental setae acute, curved. Color pale to medium tan, marked darker purplish brown, mostly on dorsum. Legpairs one and two reduced, three to seven enlarged, three to five with swollen femora, femora of pair seven slender; large femoral knobs on pairs five and six. Anterior gonopods (Figs. 210, 214) with lateral branches robust, apically expanded and decurved, terminus cap-like; mesal branches straight, cylindrical, blunt, two-thirds to three-quarters length of lateral branches. Posterior gonopod coxites (Figs. 211, 215) with evenly curved and tapered pseudoflagellar process, fimbriate branch small, subtending shallow, rimmed pit with pore, mesobasally arising thin process. Terminal processes both strongly recurved. Tenth coxae and eleventh prefemora as usual. Female: Length, 11.5 mm, nonsexual characters as in male. Distribution: WASHINGTON: Jefferson Co., Olympic National Park, near Hoh Rain Forest Visitor Center, elev. 180 m., 47.8597°N, - 123.9413°W, 14 December 2003, W. Leonard, C. Richart, mm; Clallam Co., Olympic National Park, Heart of the Hills Campground, elev. 545 m., 48.0355°N, - 123.4263°W, 18 November 2003, W. Leonard, m f.Published as part of Shear, William A., Richart, Casey H. & Wong, Victoria L., 2020, The millipede family Conotylidae in northwestern North America, with a complete bibliography of the family (Diplopoda, Chordeumatida, Heterochordeumatidea, Conotyloidea), pp. 1-78 in Zootaxa 4753 (1) on pages 56-57, DOI: 10.11646/zootaxa.4753.1.1, http://zenodo.org/record/398378
Bifurcatella angulata Shear & Richart & Wong 2020, new species
No full textBifurcatella angulata, new species Figs. 212, 213, 216, 217 Types: Male holotype, male and female paratypes from WASHINGTON: Grays Harbor Co., FS-2153 5.3 mi N of Kelley Road, Canyon River basin, elev. 160 m., 47.3035°N, - 123.5081°W, collected 12 December 2004, by W. Leonard. Diagnosis: The mesal branches of the anterior gonopods are about half the length of the lateral branches, which are curved at the tips and with a subterminal tooth. The mesal side of the posterior gonopod coxites is developed into a large, serrate flange (Fig. 217) lying parallel to the tip of the pseudoflagellar branch. The pseudoflagellar branch is evenly curved from its base but midway makes a sharp angle before tapering to a thin tip. No other species has this combination of characters. Etymology: The species name refers to the sharp, angular bend in the pseudoflagellar branch of the posterior gonopod coxite. Description: Male paratype: Length, 9.5 mm. Twenty ocelli in triangular group. Metazonites with moderately well-developed shoulders, segmental setae curved, acute. Color pale tan marked darker purplish brown. Legpairs one and two reduced, three through six enlarged, seventh near size of postgonopodal legs; fifth femora conspicuously swollen, with long, distomesal knob. Anterior gonopods (Figs. 212, 216) long, narrow, mesal branch half length of lateral, cylindrical, blunt, slightly clavate, anterior branch slender, curved at tip, with subterminal tooth. Posterior gonopod coxite (Fig. 213, 217) with pseudoflagellar branch evenly curved from base, reflexed at acute angle, tapering to finely pointed tip, lying alongside serrate lamella on mesal side. Fimbriate branch relatively large. Main branch of coxite with two hooked terminal processes, subterminal tooth large, broad. Tenth coxae, eleventh prefemora as usual. Female 9.0 mm long, nonsexual characters as in male. Distribution: WASHINGTON: Grays Harbor Co., Middle Satsop Road 2 mi N of Brady near Satsop River, elev. 25 m., 47.0245°N, - 123.5196°W, under maple leaves, 3 January 2003, W. Leonard et al., mm ff. Lewis Co., Iron Creek Campground, FS-25 9.6 mi S of Randle, Gifford Pinchot National Forest, elev. 340 m., 46.4289°N, - 121.9868°W, 21 December 2003, W. Leonard, C. Richart, m. Notes: molecular phylogenetics and integrative taxonomy of other poorly dispersing arthropods only known from the Southern Cascades and the Olympic Peninsula have found that these regions support populations that are independently evolving species (e.g., Richart and Hedin 2013). Thus, we recommend that future research evaluate our B. angulata hypothesis with molecular data.Published as part of Shear, William A., Richart, Casey H. & Wong, Victoria L., 2020, The millipede family Conotylidae in northwestern North America, with a complete bibliography of the family (Diplopoda, Chordeumatida, Heterochordeumatidea, Conotyloidea), pp. 1-78 in Zootaxa 4753 (1) on page 58, DOI: 10.11646/zootaxa.4753.1.1, http://zenodo.org/record/398378
Bifurcatella olympiana Shear & Richart & Wong 2020, new species
No full text<i>Bifurcatella olympiana,</i> new species <p>Figs. Figs. 206–208</p> <p> <b>Types</b>: Male holotype and male and female paratypes from WASHINGTON: <i>Thurston Co.</i>, Olympia, Allison Springs, elev., 55 m., 47.0454°N, - 122.9821°W, collected 4 December 2004, by W. Leonard, C. Richart.</p> <p> <b>Diagnosis</b>: Separable from all congeners in that the mesal branch of the anterior gonopod is as long as (and from some angles may appear longer than) the lateral branch and is curved and acute at the tip as opposed to straight and blunt.</p> <p> <b>Etymology</b>: The species name is an adjective referring to the city of Olympia, Washington, from which many records of this species are known.</p> <p> <b>Description</b>: <i>Holotype male:</i> Length, 11.5 mm. 20–22 ocelli in triangular eye group. Metazonites with moderately well-developed shoulders, segmental setae curved, acute. Color light grayish brown, marked darker purplish brown; from above four dark spots on each ring give the impression of a dark median stripe. Legpairs one and two reduced, three to seven thickened; femoral knobs distal on femora four to seven. Anterior gonopods (Figs. 206, 208) with two subequal branches, lateral branch decurved and expanded at tip, mesal branch tapering, decurved and acute at tip. Posterior gonopod coxites (Fig. 207, 209) with pseudoflagellar branch uniquely “kinked” about midlength, firmbriate branch small, pore mesal of fimbriate branch. Body of coxite with broad serrate lamella, two subterminal hooks, slightly expanded tip. Coxae 10 and prefemora 11 as usual.</p> <p>Female 11.0 mm long, nonsexual characters as in male.</p> <p> <b>Distribution</b>: WASHINGTON: <i>Thurston Co.</i>, Rainier Road SE, 3.0 mi NW of Military Road SE, Fort Lewis Military Reservation, elev. ca. 100 m., ca. 46.9351°N, - 122.7600°W, 29 November 2003, W. Leonard, mm ff; Hospital Creek, above confluence with Skookumchuck River, 5 horiz mi S, 3 horiz mi E of Vail, elev. ca. 240 m., ca. 46.7732°N, - 122.5855°W, 12 October 2003, W. Leonard, m; same as previous but, 15 December 2003, W. Leonard, K. McAllister, mm ff; Capitol State Forest, Potosi Creek ca. 2 mi S of Rock Candy Mountain, elev. ca. 290 m., ca. 46.9809°N, -123.109, estimated coordinates from Potosi Creek 1 horiz mi S of Rock Candy Mountain Summit, 18 December 2002, J. Ziegltrum, R. Shoat, mm; Capitol State Forest, 1.7 horiz mi ENE of Rock Candy Mtn., elev. 450 m., 47.0170°N, - 123.0776°W, 27 October 2003, W. Leonard, m f; Capitol State Forest, unnamed tributary to Perry Creek, elev. 260 m., 47.0262°N, - 123.0692°W, 27 October 2003, W. Leonard, m; Priest Point Park, Olympia, elev. ca. 30 m., ca. 47.07°N, - 122.896°W, 23 February 2003, W. Leonard, m f; same as previous but 5 January 2003, m; same as previous but 2 February 2003, f; same as previous but 11 January 2004, m; Olympia, The Evergreen State College campus, Beach Trail N of Driftwood Road, elev. ca. 30 m., 47.0791°N, - 122.9776°W, 26 January 2003, W. Leonard, mm ff; same as previous but 27 December 2003, mm ff; McAllister Springs S of Steilacoom Road SE, elev. ca. 30 m., 47.0472°N, - 122.7286°W, 22 February 2004, W. Leonard, from the litter and woody debris of a mixed forest including <i>Acer macrophyllum</i>, <i>Alnus rubra</i>, <i>Pseudotsuga menziesii</i>, <i>Thuja plicata</i>, <i>Polystichum munitum</i>, and <i>Mahonia nervosa</i>, mm; same as previous but 7 February 2004, W. Leonard, C. Richart, mm; same as previous but 11 December 2004, W. Leonard, mm ff; Tumwater, Black Lake and Belmore Road at 66 th Avenue, elev. ca. 50 m., 46.9890°N, - 122.9665°W, 19 November 2004, W. Leonard, m; as in the type locality but 25 November 2004, W. Leonard. <i>Lewis Co.</i>, Gifford Pinchot National Forest, FS-25 8 mi S of Randle, elev. ca. 330 m., ca. 46.4475, -121.9984, 6 December 2003, W. Leonard, C. Richart, mm ff; Gifford Pinchot National Forest, FS-25 8.5 mi S of Randle, elev. 320 m., 46.4409°N, - 121.9966°W, 6 December 2003, W. Leonard, C. Richart, mm ff; same as previous but 21 December 2003, m f; SR-508 at Bremer, elev. 220 m., 46.5883°N, - 122.4258°W, 6 December 2003, W. Leonard, C. Richart, mm ff. <i>Grays Harbor Co.</i>, FS-2153 5.3 mi N of Kelley Road, Canyon River basin, elev. 156 m., 47.3035°N, - 123.5081°W, 13 October 2003, W. Leonard, m.</p>Published as part of <i>Shear, William A., Richart, Casey H. & Wong, Victoria L., 2020, The millipede family Conotylidae in northwestern North America, with a complete bibliography of the family (Diplopoda, Chordeumatida, Heterochordeumatidea, Conotyloidea), pp. 1-78 in Zootaxa 4753 (1)</i> on pages 55-56, DOI: 10.11646/zootaxa.4753.1.1, <a href="http://zenodo.org/record/3983782">http://zenodo.org/record/3983782</a>
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