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A review of the European species of the genus Tetragoneura Winnertz (Diptera: Mycetophilidae)
No full textPolevoi, Alexei, Jakovlev, Jevgeni (2011): A review of the European species of the genus Tetragoneura Winnertz (Diptera: Mycetophilidae). Zootaxa 3062: 1-12, DOI: 10.5281/zenodo.20704
New data on poorly known species of the genus Leia Meigen (Diptera, Mycetophilidae) from the Palaearctic region
No full textPolevoi, Alexei, Salmela, Jukka (2016): New data on poorly known species of the genus Leia Meigen (Diptera, Mycetophilidae) from the Palaearctic region. Zootaxa 4103 (5): 487-500, DOI: 10.11646/zootaxa.4103.5.
FIGURE 3 in New data on poorly known species of the genus Leia Meigen (Diptera, Mycetophilidae) from the Palaearctic region
No full textFIGURE 3. Leia nigricornis van Duzee, Holotype. (Photo by Mike Narahara).Published as part of Polevoi, Alexei & Salmela, Jukka, 2016, New data on poorly known species of the genus Leia Meigen (Diptera, Mycetophilidae) from the Palaearctic region, pp. 487-500 in Zootaxa 4103 (5) on page 493, DOI: 10.11646/zootaxa.4103.5.7, http://zenodo.org/record/27121
FIGURE 5. Leia rufiptera Ostroverkhova, male terminalia. A. ventral view. B. dorsal view. C. lateral view. D. aedeagal complex, lateral view. E. aedeagal complex, ventral view. F in New data on poorly known species of the genus Leia Meigen (Diptera, Mycetophilidae) from the Palaearctic region
No full textFIGURE 5. Leia rufiptera Ostroverkhova, male terminalia. A. ventral view. B. dorsal view. C. lateral view. D. aedeagal complex, lateral view. E. aedeagal complex, ventral view. F. hypoproct. Scale bars = 0.2 mm. Abbreviations: aed = aedeagus, cerc = cercus, gc = gonocoxite, gst = gonostylus, par = parameres, tg = tergite, vmpg = ventral medial process of gonocoxites.Published as part of Polevoi, Alexei & Salmela, Jukka, 2016, New data on poorly known species of the genus Leia Meigen (Diptera, Mycetophilidae) from the Palaearctic region, pp. 487-500 in Zootaxa 4103 (5) on page 496, DOI: 10.11646/zootaxa.4103.5.7, http://zenodo.org/record/27121
FIGURE 1. Male terminalia. A. T. ambigua, dorsal view. B. T. ambigua, ventral view. C. T. ambigua, lateral view. D. T in A review of the European species of the genus Tetragoneura Winnertz (Diptera: Mycetophilidae)
No full textFIGURE 1. Male terminalia. A. T. ambigua, dorsal view. B. T. ambigua, ventral view. C. T. ambigua, lateral view. D. T. pudogensis sp. n., dorsal view. E. T. pudogensis sp. n., ventral view. F. T. pudogensis sp. n., lateral view. Scale bars = 0.2 mm. Abbreviations: ae = aedeagus, gc = gonocoxite, gst = gonostylus, tg = tergite, sml = submedian lobe.Published as part of Polevoi, Alexei & Jakovlev, Jevgeni, 2011, A review of the European species of the genus Tetragoneura Winnertz (Diptera: Mycetophilidae), pp. 1-12 in Zootaxa 3062 on page 5, DOI: 10.5281/zenodo.20704
Tetragoneura
No full textKey to European species of Tetragoneura 1. R 4 and small cell absent. Sensory organ on dorsal side of male mid tibia present. Mid tibia in male with one spur. Tergite 9 at least half as high as gonocoxites, cerci reduced.............................................................. 2 - R 4 normally developed forming elongated small cell (small cell may be reduced or even absent in one or both wings in T.sylvatica). Sensory organ on dorsal side of male mid tibia not developed. Mid tibia in male with two spurs. Tergite 9 not higher than 1 / 3 of gonocoxites, mostly as a narrow transverse stripe, cerci well developed.................................. 3 2. Gonostylus relatively short with apical hook. Tergite 9 with submedian appendages (Fig. 1: A–C). Female terminalia as in Fig. 3 A, B................................................................... Tetragoneura ambigua Grzegorzek - Gonostylus elongated, angled the middle. Tergite 9 without submedian appendages (Fig. 1: D–F)................................................................................................... Tetragoneura pudogensis sp. n. 3. Tergite 9 divided into two separate sclerites. Gonostylus attenuated to apex, with curved seta at base (Fig. 2: E, F). Female terminalia as in Fig. 3: C, D....................................................... Tetragoneura ruuhijarvii sp. n. - Tergite 9 integrate. Gonostylys different.................................................................... 4 4. Gonostilus bilobed (Fig. 2: C, D)................................................ Tetragoneura obirata Plassmann - Gonostylus ovate with strongly sclerotised, blade-like sub-apical process (Fig. 2: A, B). Female terminalia as in Fig. 3: E, E.............................................................................. Tetragoneura sylvatica CurtisPublished as part of Polevoi, Alexei & Jakovlev, Jevgeni, 2011, A review of the European species of the genus Tetragoneura Winnertz (Diptera: Mycetophilidae), pp. 1-12 in Zootaxa 3062 on page 10, DOI: 10.5281/zenodo.20704
Leia iturupensis Zaitzev 2001
No full textLeia iturupensis Zaitzev, 2001 (Figs. 2; 6 B) Leia iturupensis Zaitzev, 2001: 457 Material examined: RUSSIA: 1 ♂, Amur province, Klimoutsy, 40 km W of Svobodnyi, 13.8. 1958 (ZIN, Zinovjev leg., pinned). Diagnosis. A yellow species with clear wings and tripartite gonostylus, dorsal branch forked apically. Description. Male (n= 1) Head yellow with darkened spots around ocelli, covered with pale setae. Three ocelli, with laterals close to, but not touching eye margin,. Clypeus, mouthparts and palpus yellow. Palpus slightly longer than head height. Scape and pedicel yellow, flagellum yellow at the base, gradually darkened after segment 4. Pedicel with one elongate seta apically, reaching the apex of flagellomere 2. Middle flagellomeres 0.8 as long as wide. Thorax mainly yellow with pale setae. Mesoscutum yellow with brown patches above wing bases. Scutellum yellow with four strong marginal bristles. Lateral parts of the thorax and mediotergite yellow. Wing (Fig. 6 B). Wing length 3.5 mm. Wing hyaline with small dark spot at base of anal veins. Veins yellow. All, except Sc and A 2 densely setose on dorsal side. Sc 2 well beyond the middle of Sc. Base of M-fork beyond the middle of ta. CuA 1 interrupted at base. R 1 0.8 times as long as ta, which is 0.16 times as long as R 5 and 1.19 times as long as M-stem. Haltere pale yellow. Legs. Coxae yellow, fore coxa with punctiform black spot at the apex, mid and hind coxa with narrowly darkened apical margins. Trochanters yellow with dark ventral spots. Femora yellow, apex of hind femur narrowly darkened. Tibiae and tarsi yellow. Ratio of femur to tibia for fore, mid and hind legs: 1.17; 1.0; 0.79. Ratio of tibia to basitarsus for fore, mid and hind legs: 1.11; 1.46; 2.24. Tibial setae black, spurs yellow. Fore tibia with one spur that is 2.57 times as long as apical tibial diameter. Mid and hind tibia with two spurs. Posteroventral spur on mid tibia 1.24 times longer than anteroventral spur and posteroventral spur on hind tibia 1.23 times longer than anteroventral spur. Abdomen uniformly yellow with pale setae. Terminalia (Fig. 2) yellow. Gonocoxite with only a few setae on the apical part. Medial ventral process of gonocoxite long and slender, with central protrusion bearing a group of long curved setae. Apicoventral corner of gonocoxite attenuated with two apical and several subapical setae. Gonostylus tripartite, dorsal branch forked apically. Tergite 9 small, rounded caudally and with several setae. Cerci elongate, about 2 times as long as tergite 9. Aedeagus with sclerotized, arrow-headed, apical part, ejaculatory apodeme bilobed. Paramere about 1.3 times as long as aedeagus, bowed in lateral view, with sharp apex and ventral flange in the middle. Hypoproct with slightly protruding apicolateral corners and medial group of stout setae. Female. Unknown Biology. Collected by sweep net in mixed oak-pine forest. Distribution. East Russia (Kuril Islands, Amur Province). Remarks. L. iturupensis was only known from the holotype from the Kuril Islands. The specimen studied by us is mostly yellow, while the holotype has brown spots on the thorax and abdomen (Zaitzev 2001). Nevertheless the peculiar structure of the gonostylus leaves us in no doubt about the species determination. Based on male terminalia, L. iturupensis resembles L. graeca Bechev, L. umbrosa Caspers and L. subfasciata (Meigen). Principal differences are found in the gonostylus. In L. iturupensis the dorsal branch of the gonostylus is forked apically, while in other species it is simple (L. umbrosa, L. subfasciata) or with a small preapical tooth (L. graeca) (see Caspers 1991, p. 329, Fig. 13, Bechev 1997, p. 180, Figs. 1–3).Published as part of Polevoi, Alexei & Salmela, Jukka, 2016, New data on poorly known species of the genus Leia Meigen (Diptera, Mycetophilidae) from the Palaearctic region, pp. 487-500 in Zootaxa 4103 (5) on pages 490-492, DOI: 10.11646/zootaxa.4103.5.7, http://zenodo.org/record/27121
Leia bimaculata Meigen 1804
No full textVariation in the male terminalia of Leia bimaculata (Meigen, 1804) Leia bimaculata is a common European species, which is variable in body coloration (from almost completely yellow to mostly black) and wing markings (from distinct dark bands to completely clear wing). Male terminalia of this species have been illustrated by several authors. None of the available figures give details of the aedeagal complex structure, but there is always a well visible dorsal projection at the base of the gonostylus (Plassmann 1973; Hutson et al. 1980; Zaitzev 1994). Considering stable interpretation of the species, we call such specimens "typical". During our study we noticed four specimens from Finland and Russia with small differences in the male terminalia. The most distinctive characters are as follows. Typical L. bimaculata: lobes of ejaculatory apodeme straight (Fig. 7: E), gonostylus with conspicuous dorsal projection at base (Fig. 7: A, H–J). Deviating specimens: lobes of ejaculatory apodeme protruding caudally (Fig. 7: G), gonostylus with no trace of dorsal projection at base (Fig. 7: K) Other differences are visible e.g. in the shape of parameres (Fig. 7: D–G), but this may depend on the position of terminalia on the slide and the angle of view. Small variation in hypoproct structure (Fig. 7: B, C) is observed in both typical and deviating speciemen groups. All deviating specimens have clear wings, while studied typical specimens have wings with distinct (though sometimes quite faint) preapical band. Variable body coloration (from rather narrow dark bands on tergites and mostly yellow thorax, to considerably darkened abdomen and thorax) is observed in both typical and deviating specimen groups. Though differences in the gonostylus and ejaculatory apodeme seem to be stable, we did not feel confident enough to describe this as a new species until additional material is available. Material examined. Typical specimens: AUSTRIA: 2 ♂, Igls, Tirol, 900 m, 16.8. 1953, J. R. Vockeroth leg. (CNC, pinned). ITALY: 1 ♂, Caserta, Campania Prov., 30.4. 1844, (CNC, coll. G. E. Shewell, pinned); 1 ♂, Bozen, 13.6.1873? (ZIN, # 217, pinned). LITHUANIA: 1 ♂, Vilnius distr. Karmazinai, 54 ° 49 '08,4": 24 ° 55 '51,9", 19.9 – 16.10. 2003, rearing from Paxillus involutus (LZM, J.Rimsaite leg., pinned). SLOVAKIA: 1 ♂, Muranska planina Nat. Park Muran, Hrdzava dolina 26.7. 2010, Malaise trap, (JES, J. Ševčík leg. #DIPT-JS- 2014 -0297); 1 ♂, Bratislava env., Devinska Kobyla hill, 17.6. 1994, Malaise trap (JES, J. Ševčík leg., #DIPT-JS- 2014 -0298). CZECH REPUBLIC: 1 ♂, 1 ♀, Bohemia, Zivohost, near Slapy dam, 22.8. 2010, rearing from fungi, ex Lactarius vellereus, (JES, J. Ševčík leg., DIPT-JS- 2014 -0299). SPAIN: 1 ♂, Granada, 700 m, 19.7. 1960, J. R. Vockeroth leg. (CNC, pinned). SWEDEN: 2 ♂, Sm: Ulvsdal, 14.5.– 14.8.2002, Malaise trap, (IZBE, N. Franc & Co. leg., pinned). UNKNOWN LOCALITY: 1 ♂, 24.7.1842? (ZIN, coll. Osten-Sacken, pinned). Deviating specimens: FINLAND: 2 ♂, Al: Saltvik, (ZMH, Frey leg. # 3760 and # 4016, pinned). RUSSIA: 2 ♂, Leningrad province, Luga 13 – 14.8.1953 (ZIN, Stackelberg leg., pinned).Published as part of Polevoi, Alexei & Salmela, Jukka, 2016, New data on poorly known species of the genus Leia Meigen (Diptera, Mycetophilidae) from the Palaearctic region, pp. 487-500 in Zootaxa 4103 (5) on pages 497-499, DOI: 10.11646/zootaxa.4103.5.7, http://zenodo.org/record/27121
Leia rufiptera Ostroverkhova 1977
No full textLeia rufiptera Ostroverkhova, 1977. (Figs. 5; 6 D) Leia rufiptera Ostroverkhova, 1977: 30. Material examined: RUSSIA: 1 ♂, Primorje territory, east slope of Sikhote-Alin ridge, basin of the river Sankobe, 43 km along river Sitsa, 1.8. 1941 (ZIN, Grunin leg., pinned). Diagnosis. A yellow species, wings with preapical dark spot and darkened areas along veins, terminalia densely covered with strong setae. Description. Male (n= 1) Head yellow with darkened spots around ocelli, covered with pale and brownish setae. Three ocelli, with laterals very close to (almost touching) eye margins. Clypeus, mouthparts and palpus yellow. Palpus slightly longer than head height. Scape and pedicel yellow, flagellum yellow with darker dorsal spots (more extensively darkened in apical segments). Middle flagellomeres 1.4 times as long as wide. Thorax mainly yellow with brownish setae. Mesoscutum yellow with three darker longitudinal stripes and small brown patches above wing bases. Scutellum yellow with four strong marginal bristles. Lateral parts of thorax yellow. Mediotergite slightly darkened caudally. Wing (Fig. 6 D). Wing length 5.37 mm. Wing greyish with yellowish anterior margin, narrow apical cloud, dark preapical spot, darkened areas at base of median fork, along ta, CuA 2 and apical portion of M 1 and with small dark spot at base of anal veins. Veins yellowish-brown, all, except Sc and A 2, densely setose. Sc 2 near the middle of Sc. Base of M-fork well beyond the middle of ta, slightly before Rs. CuA 1 not interrupted at base. R 1 1.05 times as long as ta, which is 0.27 times as long as R 5 and 1.07 times as long as M-stem. Haltere yellow. Legs. Coxae yellow, fore coxa with punctiform black spot at apex, mid and hind coxa with narrowly darkened apical margins. Trochanters yellow with dark ventral spots. Femora yellow, mid and hind femora darkened at base. Tibiae and tarsi yellow. Ratio of fore femur to fore tibia: 1.13. Ratio of fore tibia to fore basitarsus: 1.01. Tibial setae black, spurs yellow. Fore tibia with one spur, that is 1.75 times as long as apical tibial diameter. Abdomen yellow with pale setae, tergites and sternites darkened posteriorly. Terminalia (Fig. 5) yellow. Gonocoxite densely covered with strong setae. Ventromedial process of gonocoxite also densely bristled, with widened apical process produced caudally. Gonostylus with strongly sclerotised hook-like ventral portion. Tergite 9 subrectangular, with slightly emarginated caudal margin, setose apically. Cerci pointed at apices, about as long as tergite 9. Aedeagus with T-shaped apical portion. Parameres short, fused, forming apical triangular incision and paired apicolateral openings. Hypoproct with smooth apicolateral corners and medial group of numerous thin setae. Female. Unknown Biology. According to Ostroverkhova (1979), species was found in Larch and mixed forests. Distribution. East Russia. Remarks. Three syntypes of L. rufiptera are located in the collection of Tomsk University, but terminalia preparations are severely damaged (E. Subbotina, pers. comm.). Nevertheless, the specimen examined by us fits the original description and male terminalia agree well. Several distinctive characters mentioned by G. Ostroverkhova (wing markings, dense bristles on gonocoxites, paired apicolateral openings in aedeagal complex), leave little doubt about the identity of the specimens. Leia rufiptera is easily distinguished from other representatives of the genus by the peculiar wing markings and structures of the male terminalia (e.g. densely setose gonocoxite, strongly sclerotized hook-like ventral portion of gonostylus and fused parameres).Published as part of Polevoi, Alexei & Salmela, Jukka, 2016, New data on poorly known species of the genus Leia Meigen (Diptera, Mycetophilidae) from the Palaearctic region, pp. 487-500 in Zootaxa 4103 (5) on pages 495-496, DOI: 10.11646/zootaxa.4103.5.7, http://zenodo.org/record/27121
Tetragoneura pudogensis Polevoi & Jakovlev, 2011, sp.n.
No full text5. Tetragoneura pudogensis sp.n. (Fig. 1: D–F) Holotype: 3, RUSSIA, KARELIA, Kp: Prirechnyi, 3 km SW (alt. 100 m), 24.6 – 13.8.2009, (ZISP, A.Polevoi leg.) Paratypes: 3 3, same data (ZISP, A.Polevoi leg.). Other material examined. FINLAND: 1 3, Ab: Karjalohja, Karkali Strict Nature Reserve, 24.8.– 05.10.2004 (METLA, J. Jakovlev leg.). Etymology. Named after locality Pudozh (also administrative district and biogeographical province, where type material originates from) in the South–East of Republic Karelia, using Latin suffix -ensis designating the place of origin. Diagnostic characters. This species is closely related to T.ambigua Grzegorzek and T. sasakawai Oliveira & Amorim. Reliably distinguished only by the structure of male terminalia. Description. Male (n= 4) Head black, densely gray dusted (posterior part looks thinly dusted in some angles), with numerous long black setae on vertex. Lateral ocellus separated from eye margin by the distance of 1.1–1.5 times its own diameter. Palpus yellow. Clypeus black, clothed with short hairs. Antennae brown, pedicel and two basal flagellomeres yellow. Fifth flagellomere 1.7–2.3 times as long as wide. Scape and pedicel with long black setae apically. Eyes densely covered with minute hairs. Thorax. Scutum and scutellum black with gray polinosity. Scutum with short biserial ac, biserial dc and some long black setae by lateral margin. Pleurae dark-brown, thinly grey dusted. Antepronotum and proepisternum haired and with black bristles, the rest of the pleurae bare. Wing length 2.38–2.65 [mean value 2.56] mm. Wing hyaline, veins yellow, costa and radius brownish. C extending well beyond the middle of the distance between R 5 and M 1. Sc short, ending free. R 4 absent. R 1 1–1.3 times as long as ta. Stem of M-fork 1.2–1.4 times as long as ta. Base of CuA-fork lies under the middle of ta. Macrotichia well developed on all veins with exception of proximal part of M. Haltere yellow. Legs yellow, hind femur darkened apically. Front tibia with small pv. Mid tibia with 1 a, 1 ad, 2 pd, 3–4 p and pv. Hind tibia with 6–8 ad, 12–13 pd. Ratio of tibia to first tarsomere for front, mid and hind legs: 1.37–1.45 [1.41]; 1.7–1.86 [1.77]; 1.84–2.05 [1.99]. Front and mid tibia with one spur, hind tibia with two spurs of subequal length. Mid tibia with well developed ovate sensory organ dorsally on basal third. Abdomen dark-brown, tergites II and III with yellow spots at base. Terminalia (Fig. 1: D–F), brown. Gonocoxites with narrow median cleft almost reaching their base. Gonostylus rather long, strongly sclerotised, bearing sparse short hairs apically. Aedeagus long well protruding beyond apical margin of tergite 9, reminds jumping dolphin in lateral view. Tergite 9 with wide triangular distal excavation, cerci reduced. Female. Unknown. Biology. All specimens were obtained with Malaise trap in herb-rich mixed secondary forest. Distribution. Only known from south–east part of Republic Karelia (north–west Russia) and south Finland.Published as part of Polevoi, Alexei & Jakovlev, Jevgeni, 2011, A review of the European species of the genus Tetragoneura Winnertz (Diptera: Mycetophilidae), pp. 1-12 in Zootaxa 3062 on pages 9-10, DOI: 10.5281/zenodo.20704
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