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Eutriptus jirinae Lackner & Kapler 2007
Eutriptus jirinae Lackner & Kapler, 2007 Material examined. YEMEN: SOCOTRA: 14 spec., Dixam Plateau, Firmihin, 400–500 m, 12°28′27″N, 54°00′64″E, 22.–25.vi.2009, L. Purchart & J. Vybíral lgt. (1 ex. in TLCM; 13 exs. in NMPC); 9 spec., Dixam Plateau, Firmihin (Dracaena forest), 12°28.6′N, 54°01.1′E, 490 m, 15.–16.xi.2010, J. Hájek lgt. (6 spec. in PVCV; 3 spec. in NMPC); 2 spec., Wadi Zirik, 650–670 m, 12°29′35″N, 53°59′28″E, 16.vi.2009, L. Purchart lgt. (1 spec. in TLCM; 1 spec. in NMPC); 1 spec., Wadi Ayhaft, 24.–26.xi.2003, 12°36′38″N, 53°58′49″E, 190 m, P. Kabátek lgt. (TLCM); 2 spec., Dixam Plateau, 14.–15.vi.2012, Firmihin, Dracaena woodland, 12°28.6′N, 54°01.1′E, 490 m, J. Bezděk, J. Hájek, V. Hula, P. Kment, I. Malenovský, J. Niedobová & L. Purchart lgt. (TLCM). Comments. Its occurrence on the island is mainly on Dixam Plateau, in Wadi Zirik and Wadi Ayhaft (LACKNER & KAPLER 2007, present paper). Endemic to Socotra.Published as part of Lackner, Tomáš & Vienna, Pierpaolo, 2017, Histeridae of Socotra (Coleoptera: Histeroidea), pp. 55-76 in Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) (Acta. Ent. Mus. Natl. Pragae) 57 on page 65, DOI: 10.1515/aemnp-2017-0107, http://zenodo.org/record/532469
Hypocaccus Lackner, 2015, sp. nov.
Hypocaccus (sensu lato) hirsutus sp. nov. Type material examined. Holotype, ♀, side-mounted on a triangular point, left antennal segments 3-11 missing, right protarsus and left mesotarsus broken off, with the following labels: “♀” (printed); followed by: “S India, Kerala, 1994 / SHORANUR 10 º 46 ’N 76 º 16 ’E / bank of Ponnáni riv. / 31.I., Z. Kejval lgt.” (printed); followed by: “ Hypocaccus (s.str.) ? / spec. ?? / det. P. Kanaar 2002 ” (printed-written); followed by: “Gen. nov. / sp. nov. / P. Vienna det., 2003 ” (printed); followed by: “ Hypocaccus (s.l.) / hirsutus sp. nov. / HOLOTYPE / Det. T. Lackner 2015 ” (red label, written) (TLAN). FIGURE 1. Hypocaccus (sensu lato) hirsutus sp. nov., habitus, dorsal view. FIGURE 2.ditto, ventral view. Description. Body (Figs. 1–2) PEL: 2.55 mm; APW: 1.25 mm; PPW: 2.10 mm; EW: 2.20 mm; EL: 1.50 mm; cuticle light to rusty brown, pronotum somewhat darker than elytra, legs, antenna and mouthparts similarly colored. Head (Fig. 3): mandibles rather thin, pointed apically, left mandible with a large triangularly shaped subapical tooth; anterior margin of transverse labrum elevated, keel-like, other mouthparts not examined; clypeus transverse, with faint shallow punctures with keel-like median elevation, surface posterior to this elevation depressed, anterior to it clypeal disc even; frontal stria carinate, outwardly arcuate, supraorbital stria erased, absent; occipital stria complete, feebly carinate; frontal disc with scattered microscopic punctation, anteriorly with an almost straight transverse carina, posteriorly with a carinate chevron; between the two a short transverse carina present; frontal disc posterior to chevron with another transverse carina basically delimiting frontal disc from occiput; eyes flattened, well-visible from above. Antennal scape with numerous long amber setae; 8 th antennomere shaped like a cupule surrounding antennal club up to one-third (Fig. 3), resembling that of Philothis (Atavinus) atavus (for fig. see Lackner 2010, fig. 6). Antennal club globular, conspicuously small; sensory structures of the antennal club examined only externally, due to the unique available specimen, which lacks one antenna. Although the antennal club has not been chemically cleared, a single stipe-shaped vesicle situated on internal distal side of the club was observable under great magnification. Basal half of antennal club glabrous, apical half with intermingled dense short and sparser longer erect sensilla. Pronotum sub-trapezoidal, antennal angles acute, pronotal sides narrowing anteriorly; marginal pronotal stria carinate, complete, becoming rather thin and sub-carinate in posterior angles. Entire pronotal disc with punctation, punctures on most of disc forming elongate confluent rugae; only on ante-scutellar area punctures free, separated by their own to several times their diameter. Pronotal base with a double row of dense punctures; ante-scutellar area with a faint depression; pronotal hypomeron with dense long amber setae. Scutellum rather small, triangular. Elytra: marginal epipleural stria well-impressed; elytral epipleura punctate; marginal elytral stria complete, carinate, continuous with complete apical elytral stria that is continuous with complete and carinate sutural elytral stria. Oblique humeral stria short, double; inner subhumeral stria present as short median fragment; elytral disc with four discal elytral stria 1–4, first stria the longest, surpassing elytral half apically; striae 2–4 somewhat shortened, reaching approximately half of elytral length apically; fourth stria weakly developed, interrupted on its half-length. All striae rather weak and obscured by dense elytral punctation; entire elytral disc densely punctuate, punctures separated by less than their diameter; punctation becomes sparser near elytral flanks and elytral base. Propygidium transverse, its punctation even denser than that of elytra, punctures almost confluent; pygidium with similar, but sparser punctation, punctures here separated by their own to twice their diameter, becoming sparser apically. Prosternum (Fig. 4): entire prosternal surface with alutaceous microsculpture, prosternal process with punctures; surface between carinate and complete carinal prosternal striae densely punctate; lateral prosternal striae complete and carinate, convergent anteriorly, united in front of united carinal prosternal striae, passing mesad to small but deep prosternal foveae. Mesoventrite (Fig. 4): transverse, punctate, punctures separated by about their own to twice their diameter; marginal stria of mesoventrite carinate laterally, anteriorly obsolete, antero-medially absent; meso-metaventral stria undulate, concealing meso-metaventral suture. Metaventrite (Fig. 4) covered with scattered fine punctures separated by several times their diameter, surface along median longitudinal line slightly depressed; lateral metaventral stria straight, carinate, not reaching metacoxa. Lateral disc of metaventrite with rather deep punctures of various sizes, punctures without setae; metepisternum with similar punctation becoming weaker and sparser on fused metepimeron; lateral metepisternal stria present as a short vague fragment on fused metepimeron. First visible abdominal ventrite with punctation similar to that of metaventrite, completely striate laterally; punctation becoming microscopic and very sparse apically. Legs: protibia: outer margin apically with two rather approximate low teeth topped by large round denticle; this double-tooth rather widely separated from the following 6 proximal low teeth each topped by round denticle diminishing size in proximal direction. Protarsal groove shallow, protibial spur large, bent, growing out from apical protibial margin; posterior face of protibia rugulose-lacunose; apical margin ventrally with two tiny denticles, posterior protibial stria carinate and complete, separating rugulose-lacunose outer part of protibia from smooth median part of prosternal surface; inner row of setae sparse; inner posterior denticles absent. Mesotibia and metatibia similar, outer margin of both with dense row of long thick denticles supplemented by another, sparser row of shorter denticles on anterior face; both meso-and metatibial spurs well developed and rather long, straight; each meso-and metatarsomere with two long strongly sclerotized setae; tarsal claws in both cases approximately as long as half-length of their respective apical-most tarsomeres, slightly bent apically. Male unknown. Differential diagnosis. Hypocaccus hirsutus sp. nov. differs markedly from all species of the genus Hypocaccus from India by densely ciliate pronotal hypomeron as well as by strongly elevated anterior margin of clypeus. Biology. Unknown, the type specimen was collected on the bank of Ponnáni River. Distribution. Known only from the type locality, village of Shoranur in the Indian province of Kerala (Fig. 5). Discussion. The genus Hypocaccus C. Thomson, 1867 contains currently three subgenera: Hypocaccus, Baeckmanniolus Reichardt, 1926 and Nessus Reichardt, 1932. The primarily Holarctic nominotypical subgenus FIGURE 3. Hypocaccus (sensu lato) hirsutus sp. nov., head, dorsal view. FIGURE 4. Hypocaccus (sensu lato) hirsutus sp. nov., prosternum, meso+metaventrite. FIGURE 5. Type locality of Hypocaccus (sensu lato) hirsutus sp. nov. in India. Hypocaccus is normally found on sandy soils, banks of rivers, and seashores, and includes 44 described species (Lackner 2014 d). The pronotal hypomeron of Hypocaccus s. str. is normally asetose; however, several Nearctic psammophilous representatives (e.g., H. texaco Mazur, 1991) do possess ciliate pronotal hypomeron. From India, three species have been recorded so far: H. brahminius (Marseul, 1864), H. brasiliensis (Paykull, 1811) and H. sinae (Marseul, 1862). Another subgenus, the strictly beach-dwelling Baeckmanniolus, with 11 described species (Lackner 2014 d), occurs mainly on seashores of all continents (except Antarctica) but has not yet been recorded from India, although one species, H. (B.) varians (Schmidt, 1890) is widely spread in Indo-Malayan as well as Australian regions. Mazur (2011) reports it from neighboring Sri Lanka. Species of Baeckmanniolus do not have a ciliate pronotal hypomeron. The two subgenera, Hypocaccus and Baeckmaniolus are separated based on their pronotal punctation (species in the subgenus Baeckmanniolus possess impunctate pronotum vs. punctate in the subgenus Hypocaccus) and number of rows of denticles on the metatibia (two in Hypocaccus vs. three in Baeckmanniolus) (Lackner 2010, Bousquet & Laplante 2006). The third subgenus is the species-rich Nessus containing 53 species distributed almost exclusively in the Old World and occurring chiefly on carcasses, mammal excrement, and in rodent nests, with some specialized Middle- Asian psammophilous forms (Lackner 2014 d). Four species: H. (N.) fugax (Marseul, 1857), H. (N.) fugitivus (Desbordes, 1925), H. (N.) malabaricus (Reichardt, 1932) and H. (N.) rubripes (Erichson, 1834) are known from India (Mazur 2011). Nessus was historically treated as a subgenus of the genus Hypocacculus Bickhardt, 1914 but was moved into Hypocaccus by Mazur (2011) without explanation. Although Mazur (2011) did not give any reasons for inclusion of Nessus in Hypocaccus, he apparently did so based on few differences between the two taxa (the densely punctate-variolate structure of frons, occasionally with transverse rugae is often shared between the two taxa), with the exception of perhaps the body size, that is normally much smaller in Nessus than in Hypocaccus (Mazur pers. comm.). There are, however, also rather ‘small’ Hypocaccus species, e.g., an undescribed species from Mongolia, which only reaches 2.00 mm, a length ‘normal’ for species of Nessus (Lackner unpublished). There are also ‘large’ species of Nessus, e.g., the psammophilous H. (N.) vlasovi Kryzhanovskij, 1966 from Turkmenistan, that reaches up to 3.00 mm, a body length ‘normal’ for most species of Hypocaccus s. str. (Kryzhnovskij & Reichardt 1976). The newly described species H. hirsutus does not fit neatly into any current subgenus of Old World Hypocaccus based on the setose pronotal hypomeron, as well as strongly elevated anterior clypeal margin. It is morphologically most similar to members of Hypocaccus s. str. On the other hand, the elevated anterior clypeal margin resembles the monotypic African genus Parahypocaccus Vienna, 1995, but differs from it by the presence of prosternal foveae. Apparently, the external morphological characters (body size, pronotal pilosity, etc.) vary between species of both Hypocaccus s. str. and Nessus, most likely convergent adaptations to the environment. Recent phylogenetic analysis of the Saprininae genera and subgenera (Lackner 2014 d) based on morphological characters failed to clarify the relationships between the taxa Nessus, Hypocaccus or Baeckmanniolus, recovering them in a large unresolved clade of taxa sharing a single synapomorphy of single pear or stipe-shaped vesicle inside their antennal club. The newly-described species lacks numerous characters (e.g. male genitalia, mouthparts) that would be needed for inclusion in a cladistic analysis for proper (sub)generic placement. A subsequent phylogenetic study based on molecular characters is in progress.Published as part of Lackner, Tomáš, 2015, Hypocaccus (s. l.) hirsutus sp. nov., an atypical new species of the genus Hypocaccus C. Thomson, 1867 from India (Coleoptera: Histeridae: Saprininae), pp. 589-595 in Zootaxa 4040 (5) on pages 590-594, DOI: 10.11646/zootaxa.4040.5.8, http://zenodo.org/record/24246
Euspilotus (Platysaprinus) ferreri Lackner & Arriagada 2020, sp. nov.
Euspilotus (Platysaprinus) ferreri sp. nov. (Figs 34–44, 56) Type locality. Brazil: Minas Gerais, Parque National da Serra do Cipó. Type material examined. HoLoTYP E:, glued onto the tip of the mounting point, genitalia extracted and disarticulated, glued onto a separate mounting card under the specimen, ‘ Brazil Minas Gerais | P. N. Serra do Cipó | Campo Rupestre | 15 Tctubre 2004 | Leg. Gustavo Schiffer [printed] || Euspilotus | (Platysaprinus) | ferreri | nov. sp. | Det. G. Arriagada, 2018 [printed] || Euspilotus (Platysaprinus) | ferreri sp. nov. | HTLTToPE 2019 | des. T. Lackner & G. | Arriagada [red label, written]’ (CEMT). PARATYP Es: ♀, ‘ BRASIL: Minas Gerais. | Mata do Paraíso. 20º47’ | S; 42º51’W. F.Z.WII- | 1998 | Vaz de Mello [printed] || Euspilotus | (Platysaprinus) | ferreri | nov. sp. | Det. G. Arriagada, 2018 [printed] || Euspilotus (Platysaprinus) | ferreri sp. nov. | PARAToPE | des.T. Lackner & G. | Arriagada 2019 [red label, written]’ (CVMD);, ‘ BRASIL Minas Gerais | Viçosa | Noviembre 1999 | Leg. A. Bello [printed] || Euspilotus | ( Platysaprinus ) | ferreri | nov. sp. | Det. G. Arriagada, 2018 [printed] || Euspilotus (Platysaprinus) | ferreri sp. nov. | PARAToPE | des.T. Lackner & G. | Arriagada 2019 [red label, written]’ (CTLA); ♀, ‘ BRASIL Río de Janeiro | Nova Friburgo | Marzo 2012 | Leg. E. Grossi [printed] || Euspilotus | (Platysaprinus) | ferreri | nov. sp. | Det. G. Arriagada, 2018 [printed] || Euspilotus (Platysaprinus) | ferreri sp. nov. | PARAToPE | des. T. Lackner & G. | Arriagada 2019 [red label, written]’ (CND);, ‘ Brazil Mato Grosso | Lavras | Enero 2003 | leg. C. Cambraia [printed] || Euspilotus | (Platysaprinus) | ferreri | nov. sp. | Det. G. Arriagada, 2018 [printed] || Euspilotus (Platysaprinus) | ferreri sp. nov. | PARAToPE | des. T. Lackner & G. | Arriagada 2019 [red label, written]’ (MNHNC). Description. Body (Fig. 34). PEL: 3.50–3.60 mm; APW: 1.40–1.50 mm; PPW: 2.50–2.75 mm; EW: 2.90–3.10 mm; EL: 2.10–2.20 mm; rectangular oval, dark brown; body appendages castaneous, antennal club lighter. Head. Tccipital stria well impressed, supra-orbital stria indistinct, frontal stria present in antero-lateral angles, for short distance prolonged onto epistoma. Frons with slight lateral and posterior depressions, wholly punctate, punctures separated by their own to twice their diameter; epistoma depressed, punctate. Eyes large, protruding. Labrum and mandibles similar to preceding species; other mouthparts not examined. Antennal scape dilated and thickened, with single long seta on anterior surface and single shorter seta dorsally. Antennal club circular, depressed dorso-ventrally, tomentose, without visible external sensory structures. Pronotum (Fig. 34). Anterior pronotal angles obtuse, marginal pronotal stria complete, pronotum wholly punctate, punctures separated by 2–3 times their diameter. Pronotal hypomeron asetose, scutellum small, triangular. Elytra (Fig. 34). Elytral epipleuron weakly punctuate, marginal epipleural stria very thin; marginal elytral stria weakly cariniform, complete, continued along elytral apex for short distance; apical elytral stria otherwise absent. A dense row of prominent punctures present along marginal elytral stria. Inner subhumeral stria present as long median fragment, deeply impressed, with tiny punctures. Humeral elytral stria weakly impressed on basal elytral third; dorsal elytral striae well impressed, punctate, first longest, slightly surpassing elytral half apically, striae 2–4 slightly shorter, stopping short of half of elytral length apically, basal ends of all striae curved mediad. Fourth dorsal elytral stria curved inwardly and shortly continued as apical fragment of sutural elytral stria; sutural elytra stria otherwise present only as short and weak fragment on apical elytral half. Elytra wholly punctate, punctures on basal elytral half (chiefly among elytral striae) almost microscopic; on apical elytral half much denser and prominent, separated by once to twice their diameter, extreme elytral apex glabrous. Propygidium and pygidium densely punctate, punctation of pygidium becoming finer and sparser apically; pygidium slightly depressed laterally. Prosternum (Fig. 35). Prosternal process strongly compressed, keel-like, with fine scattered punctures; marginal prosternal stria present, terminating in large prosternal foveae; carinal prosternal striae divergent apically, terminating in prosternal foveae; lateral prosternal striae carinate, apically joining carinal striae at two-thirds point. Mesoventrite (Fig. 35) approximately four times as wide as long, with scattered microscopic punctuation; marginal mesoventral stria complete, slightly carinate; meso-metaventral stria complete, undulate. Metaventrite (Fig. 35). Disc of metaventrite with sparse microscopic punctation, along apical margin double row of larger punctures present; lateral metaventral stria almost complete, slightly carinate, stopping short of metacoxa. Lateral disc of metaventrite with three rows of large deep punctures. Abdomen. First visible abdominal ventrite slightly depressed on basal third, almost completely striate laterally, disc along margins with larger punctures, medially only with sparse microscopic punctures. Legs. Protibia dilated, on outer margin with 4–5 low teeth, each topped by short amber denticle, outer row of setae regular, protarsal groove shallow, protibial stria complete and carinate, median row of setae very short. Protibial spur short, growing out from apical protibial margin; outer part of posterior surface of protibia rugulose-lacunose; median part of posterior surface glabrous; posterior protibial stria complete, with microscopic setae; setae of inner protibial margin double, dense and regular. Mesotibia slightly dilated, on outer margin with six prominent denticles, outer row of setae sparse; median row of setae microscopic and scattered. Posterior mesotibial stria complete; protibial spur conspicuous. Anterior mesotibial stria complete; a single inner row of setae. Metatibia slightly dilated, on outer margin with seven denticles progressively increasing in length apically; metatibia otherwise similar to mesotibia. Male genitalia. Sternite VIII (Figs 36–37) strongly sclerotized, separated medially, extreme apices with tuft of microscopic setae. Tergite VIII (Fig. 37) apically almost straight, not arcuate as in preceding species; sternite VIII and tergite VIII not fused laterally (Fig. 38). Tergite W (Fig. 39) apically inwardly arcuate; tergite IW (Fig. 39) somewhat more parallel-sided than in preceding species. Spiculum gastrale (Figs 41–42) dilated on both ends, its apex (I ‘head’) strongly sclerotized. Aedeagus (Figs 43–44) with slightly larger phallobase than with preceding species; ratio phallobase: parameres 1.0: 2.5; otherwise similar to preceding species, strongly curved from lateral view, apex not as pointed as with preceding species. Differential diagnosis. Differs from E. (P.) latimanus in the absence of costate first dorsal elytral stria as well as by the non-costate lateral pygidial margins. Differs from E. (P.) myrmecophilus in the impunctate basal third to half of the elytra, and from E. (P.) vazdemelloi sp. nov. in the broadly-oval body and basally non-abbreviated dorsal elytral striae 3–4 (abbreviated in case of E. (P.) vazdemelloi sp. nov.)). Biology. Unknown. Etymology. Patronymic, this new species is named after the Spanish entomologist Juan de Ferrer Andreu (1924– 2008), a specialist on Histeridae. Distribution. Known only from Brazil (Minas Gerais, Mato Grosso and Rio de Janeiro).Published as part of Lackner, Tomáš & Arriagada, Gerardo, 2020, Revision of Euspilotus, subgenus Platysaprinus, with description of two new species (Coleoptera: Histeridae), pp. 303-317 in Acta Entomologica Musei Nationalis Pragae 60 (1) on pages 312-314, DOI: 10.37520/aemnp.2020.017, http://zenodo.org/record/387984
Otto Mazal (1932-2008)
Lackner Franz. Otto Mazal (1932-2008). In: Gazette du livre médiéval, n°54. 2009, fasc. 1. p. 124
Hypocaccus (Nessus) curtus Lackner & Seres 2018, comb. nov.
Hypocaccus (Nessus) curtus (Rosenhauer, 1847) comb. nov. (Figs 32, 40–44, 46–54) Saprinus curtus Rosenhauer, 1847: 26 (original description). MARSEUL (1855): 751 (redescription). Saprinus (Hypocaccus) curtus: GANGLBAUER (1899): 389 (redescription). Hypocacculus (Nessus) curtus: REICHARDT (1932): 49, 122 (keyed, redescription, incl. pl. IV, fig. 9); REICHARDT (1941): 285, 300 (keyed, redescription, incl. fig. 147C). Saprinus puncticollis Küster, 1849: 30 (original description). MARSEUL (1855): 755 (redescription); BICKHARDT (1916): 96 (synonymy). Saprinus (Hypocaccus) puncticollis: GANGLBAUER (1899): 389 (redescription). Hypocacculus (Nessus) puncticollis: KRYZHANOVSKIJ & REICHARDT (1976): 204, 213 (keyed, redescription); VIENNA (1980): 179, 181 (keyed, redescription, incl. fig. 64b); MAZUR (1984): 89 (catalogue); MAZUR (1997): 254 (catalogue); YÉLAMOS (2002): 320 (keyed, redescription, incl. fig. 157f); MAZUR (2004): 94 (catalogue); MAZUR (2011): 209 (catalogue); LACKNER et al. (2015): 118 (catalogue). Saprinus cribellaticollis Jacquelin du Val, 1858:99 (original description). Fauvel in GOZIS (1886): 202 (as synonym of Saprinus puncticollis). MARSEUL (1862): 509 (redescription). Saprinus (Hypocaccus) cribellaticollis: SCHMIDT (1885): 312 (keyed). Saprinus sicanus Marseul, 1862: 490 (original description, incl. pl. XVII, fig. 47). BAUDI DI SELVE (1864): 233 (as synonym of Saprinus puncticollis). Saprinus kuesteri Marseul, 1862: 715 (catalogue; unecessary replacement name for S. puncticollis Küster, 1849). Saprinus revisus Marseul, 1876: 39 (original description). BICKHARDT (1916): 97 (as synonym of Saprinus curtus). Type material examined. Saprinus curtus Rosenhauer, 1847. LECTOTYPE (present designation): ♁ (Fig. 40), originally pinned with pin-hole in its right elytron, mounted on a rectangular mounting card, right antennal funicle and left mesotarsus missing, genitalia extracted and disarticulated, glued to the same mounting card as the specimen, ‘curtus / Rosenh. [written] // Hungaria [written] // herbeus Mars. [written] // Ex Musaeo / Rosenhauer [black-margined, printed label] // pas synonime / d’Herbeus Mars. / Dr. Auzat 1917 [written-printed] // Hongrie / Ex-Musaeo / ROSENHAUER [printed] // Hypocacculus / (Nannolepidius) curtus / (Rosenhauer, 1847) / Dr. Auzat Dét. 1917 [printed] // Exemplaire provenant de la / collection Vauloger de Beaupré / Marcel (1862-1904) et inclus dans / la collection S. Risser en 2011 [black-margined, printed label] // Saprinus curtus / Rosenhauer, 1847 / LECTOTYPE / Des. T. Lackner 2017 [red label, printed]’ (ZSM). Saprinus puncticollis Küster, 1849. LECTOTYPE (present designation): ♁ (Fig. 42), glued onto a rectangular mounting card, two left and three right mesotarsomeres missing, genitalia extracted, disarticulated and glued to the same mounting card as the specimen, ‘Typ! [written] // Cagliari / Dr. Küster [written] // puncticollis / Küst. [written] // Saprinus / curtus Rosenh. [written] // Saprinus puncticollis / Küster, 1849 / LECTOTYPE / Des. T. Lackner 2017 [red label, printed]’ (ZSM). Saprinus cribellaticollis Jacquelin du Val, 1858. LECTOTYPE (present designation): ♀ (Fig.41), glued on a rectangular mounting card, both antennal funicles broken off; legs: except for right foreleg and left foretibia, all tibiae broken off; with the following labels:tiny, green rectangular label that is glued onto much larger translucent plastic mounting card (original mounting card of J. du Val) and tiny, red, quadrate label, followed by, ‘ Saprinus cribellaticollis / Jacquelin du Val, 1858 / LECTOTYPE / Des. T. Lackner 2017 [red label, printed]’ (MNHN; coll. Jacquelin du Val). Saprinus sicanus Marseul, 1862. LECTOTYPE (present designation):♁ (Fig. 43), glued onto a rectangular mounting card, right antennal funicle, both protarsi, two segments of right mesotarsus, as well as both metatibiae missing, male genitalia extracted, disarticulated and glued onto the same mounting card as the specimen, with the following labels: small, square-shaped blue label, followed by, ‘ Saprinus / sicanus m. / Schaum ‘59 [round label, written] // 129c / Saprinus / sicanus m. / Sicile / Schm 679 [round label, written] // 47 (129c) Saprin / sicanus m60 / Sicil. [written] // MUSEUM PARIS / Coll. De Marseul / 2842-90 [printed] // TYPE [red-printed label; followed by: “ Saprinus sicanus / Marseul, 1862 / LECTOTYPE / Des. T. Lackner 2017 [red label, printed]’ (MNHN). Saprinus revisus Marseul, 1876. LECTOTYPE (present designation): ♀ (Fig. 44), left antennal funicle, left protarsus, and left metatarsus missing, glued onto a rectangular mounting card, female genitalia extracted, glued to the same card as the specimen, ‘ Saprinus / revisus / rest of label illegible [round, blue label, written] // MUSEUM PARIS / Coll. / De Marseul 1890 [light-green label, printed] // TYPE [red-printed label] // Saprinus revisus / Marseul, 1876 / LECTOTYPE / Des. T. Lackner 2017 [red label, printed]’ (MNHN). Additional material examined. ALGERIA: ANNABA: Bône [= Annaba], 1 ♀, coll. Dr. Buysson (MNHN; coll. Thérond); Bône [= Annaba], 1 ♁, Desbr. (MFNB). EGYPT: Egypt, no further data, 1 ♀, coll.Ancey, (MNHN; coll. Thérond). FRANCE: BOUCHES- DU- RHÔNE: Camargue, 2 ♁♁, L. Puel lgt., Auzat coll. (MNHN; coll. Thérond); Camargue, Vaccares, no date, 1 ♁, 29.v.1937, 1 ♁, J. Thérond lgt. (MNHN; coll. Thérond); Camargue, La Sauvage, 1.v.1928, 1 ♁, L. Puel lgt. (MNHN; coll. Thérond); St. Maries de la Mer, 18.vii.1922, 1 ♀, Dr. A. Chobaut lgt., coll.Dr.Auzat (MNHN; coll.Thérond). ITALY: SARDINIA: Cagliari, Saline di Stato, 10.v.1989, 1 ♁, 3 ♀♀, C. Meloni lgt. (1 ♁ in CTLA, 3 ♀♀ in MSNG); Stagno di Molentargius, 27.iii.1979, 1 ♁, C. Meloni lgt. (CPVV), 29.v.1988, 1 ♁, 1 ♀, C. Meloni lgt. (MSNG); Serdiana, 8.vi.2003, 6 ♁♁, 6 ♀♀, Fancello lgt. (MSNG); Molentargius, 31.i.1979, 1 ♁, C. Meloni lgt. (MSNG); Cagliari, Campo Santa Gilla, 28.iii.1983, 2 ♀♀, C. Meloni lgt.(MSNG). SICILY: Sicily, no further data, 1♁., 1 spec., Krtz. (MNHN); Sicilia, no further data, 1 ♀ (MFNB). LIBYA: TRIPOLI: Tripolis, no further data, 1 ♀ (MFNB). SPAIN: ANDALUSIA: Andalusia, no further data, 1♀ (MFNB). TUNISIA: TUNIS: Tunis, 1 spec., collector unknown, Reitter coll. (ZSM); Tint, i.–ii.1882, 1 ♁, G. & L. Doria lgt. (ZIN); Carthage, vii. 1914, 1♁, Novak lgt. (ZIN); Tunis, no further data, iv.[18]83, 1 ♁ (MFNB); Tunis, no further data, 6 ♁♁, 3 ♀♀ (MFNB); Tunis, ii.–iii.1882, 1♁, G. & L. Doria lgt. (MFNB); Radès, iv.1933, 1♁, M. Grossclaude (MNHN; coll. Thérond). SOUSSE: Sebkha Kelbia lake near Sousse, 8.iv.1962, 1 ♁, Cl. Besuchet lgt. (MSNG). Redescription. PEL: 1.60–2.00 mm; APW: 0.75–1.00 mm; PPW: 1.40–1.60 mm; EW: 1.50–1.75 mm; EL: 1.00–1.40 mm. Body (Fig. 40) oblong, oval, rather convex, cuticle dark-brown to black with faint to pronounced greenish hue; legs and antennal funicle light reddish-brown; antennal scape somewhat darker. Head: mandibles densely punctate dorsally; clypeus densely and coarsely punctate, almost rugose-lacunose, anterior margin slightly elevated; frontal disc with similar, if somewhat weaker punctation; occasionally this punctation is confluent and forms tiny rugae; frontal stria slightly outwardly arcuate, complete to reduced to interrupted medially, supraorbital stria well developed; eyes flattened, but visible from above. Basal third of frontal disc with irregular rounded glabrous area; occipital stria weak, but visible. Antennal scape somewhat darker than reddish antennal funicle, antennae similar to other species of the subgenus, sensory structures of the antennal club studied by DE MARZO & VIENNA (1982). Pronotum convex, lateral sides slightly narrowing anteriorly; anterior pronotal angles obtuse, marginal pronotal stria complete, its lateral portion observable in some cases from lateral view only. Entire pronotal disc covered with punctures separated by one to several times their diameter, punctation weakens medially. Scutellum very small, triangular. Elytra: elytral epipleuron impunctate, marginal epipleural stria complete, marginal elytral stria well developed, complete, continued as apical elytral stria for short distance. Humeral elytral stria well developed, present on basal elytral third; internal subhumeral stria present as a median fragment. Dorsal elytral striae 1–4 well developed, first the longest, slightly bisinuate, reaching approximately two-thirds of elytral length apically, occasionally even slightly longer, striae 2–4 shorter, reaching approximately elytral mid-length apically, while second stria may be longer than striae 3–4; fourth stria usually the shortest, formed in most cases of beads of punctures, stopping short of elytral mid-length apically. Fourth dorsal elytral stria usually not connected (connected in specimens that belong to form ‘ cribellaticollis ’) with the basal end of (in)complete sutural elytral stria, which is in punctures and can be basally shortened. Elytral punctation covers approximately apical half of elytral disc, slightly surpassing elytral mid-length basally, slightly and scatteredly entering elytral intervals in some specimens; punctation rather dense, punctures separated by approximately their own diameter. Basal elytral fifth, fourth elytral interval, elytral flanks and extreme elytral apex impunctate, or with scattered microscopic punctation only. Propygidium and pygidium: propygidium covered with punctation similar to that of elytra; pygidium with much finer and sparser punctation. Prosternum: prosternal process slightly to moderately concave (observed from lateral view); carinal prosternal striae carinate, divergent on prosternal apophysis, running convergent to sub-parallel to almost approximate apically; from mid-length of prosternal process slightly divergent anteriorly, apically united under tiny loop; interspaces between carinal prosternal striae with scattered punctures. Lateral prosternal stria strongly carinate, convergent apically, united in front of united carinal prosternal striae; lateral sides of prosternal process densely punctate; prosternal foveae moderately large, deep. Mesoventrite: disc of mesoventrite approximately three times as wide as long, with scattered punctures (occasionally almost glabrous); marginal mesoventral stria complete, slightly inwardly arcuate medially; meso-metaventral stria undulate, bisinuate, in punctures, slightly distanced from meso-metaventral suture medially. Metaventrite: disc of metaventrite apart from several rows of tiny punctures situated along basal margin entirely glabrous; lateral metaventral stria almost straight, slightly bisinuate, deeply impressed, in punctures, stopping short of metacoxa; lateral disc of metaventrite depressed, with large oval deep punctures separated by less than their diameter; metepisternum with similar punctation, punctures of smaller sizes than those of lateral disc of metaventrite. First visible abdominal ventrite striate laterally, with scattered fine punctation, occasionally almost impunctate. Legs: protibia (Fig. 32) on outer margin with 8–11 short to moderately long denticles diminishing in size proximally, protibial groove deep; rest of leg characters similar to preceding species. Male genitalia: sternite VIII (Figs 46–47) narrowing apically; sternite VIII and tergite VIII fused laterally (Fig. 48). Tergite IX medio-laterally with tiny acute projection (Figs 49–50). Spiculum gastrale (Figs 51–52) similar to other congeners. Aedeagus (Figs 53–54) almost subparallel, bluntly pointed apically. Distribution. Hungary (?), France, Italy: Sardinia, Sicily, Spain, Portugal, Greece, Malta, Cyprus, Turkey, Tunisia, Algeria, Libya, Egypt. Biology. According to VIENNA (1980), who repeats THÉROND (1975), H. (N.) curtus is found under detritus in sand near the seacoast, where it was collected from near Suaeda sp. and Statice virgata W. plant roots. Remarks. The type specimen was part of Rosenhauer’s collection, which later became partly a part of R. Oberthür’s collection (A. Taghavian, pers. comm. 2017), currently housed in MNHN. The senior author has visited MNHN multiple times and failed to locate the type specimen(s) of this species in the collections of MNHN (including R. Oberthür’s collection). Mr. Serge Risser (Pleucadeuc, France) recently purchased the Histeridae collection of the late Marcel René Paul de Vauloger de Beaupré and published its contents in two separate papers (RISSER 2013a,b). When reading RISSER’ S paper (2013a) we were intrigued by a specimen identified as Hypocacculus (Nannolepidius !) curtus originating from Hungary and from ‘Musaeo Rosenhauer’. Mr. Risser was kind enough to send this specimen to one of us (T. L.). Having examined it as well as compared it to Rosenhauer’s original description we concluded that this is the long-lost type specimen of Rosenhauer’s species Saprinus curtus. This species was described based on an unspecified number of specimens and therefore we designate a lectotype to fix the species identity. Saprinus curtus has become a mystery practically since its description, which was, however, rather detailed and served the purpose well. The reason for this was probably the fact that the type specimen(s) were unavailable for comparison and perhaps also because no more specimens matching this species were ever reported from ‘Hungary’. Based on the description alone, BICKHARDT (1916) correctly synonymized the H. (N.) puncticollis (Küster, 1849) with H. (N.) curtus, which was also followed by REICHARDT (1932). MÜLLER (1937), however, doubted the two species are synonymous since the apical elytral stria in H. (N.) curtus reaches only mid-length of elytral apex, while, according to MÜLLER (1937) it is complete in H. (N.) puncticollis. Furthermore, MÜLLER (1937) advocated using Küster’s H. (N.) puncticollis as the valid (albeit not the earliest) name for this species and suggested, perhaps because of the incomplete description or the absence of the type material, that H. (N.) curtus was a dubious taxon. In the latest treatise on the Histeridae of the USSR (KRYZHANOVSKIJ & REICHARDT 1976), which in fact included almost the entire Palaearctic fauna, Kryzhanovskij upheld MÜLLER’ S (1937) opinion, and the name Hypocaccus (Nessus) puncticollis gained priority. This was followed by MAZUR (1984, 1997, 2011) in all three editions of his world catalogue of the Histeridae as well as by the latest edition of the Palaearctic Catalogue by LACKNER et al. (2015). Having examined both type specimens as well as numerous non-type specimens we can conclude that the two species are synonymous, and the earlier described taxon (H. (N.) curtus) has the priority. Regarding external morphological variation of this species, see Remarks section of H. (N.) curtus . Saprinus puncticollis was described from a specimen found in Cagliari by Küster himself, as well as from specimen(s) brought by Mr. Handschuh from Cartagena (Spain) (KÜSTER 1849). The depository of the Spanish specimens is unknown and hence we designate the male specimen from Cagliari (Sardinia) as the lectotype to fix the identity of this taxon for purpose of synonymy. Saprinus cribellaticollis was described based on unknown number of specimens.A single specimen was located in the original collection of Jacquelin du Val, deposited in MNHN, under the label ‘ Saprinus cribellaticollis ’. Jacquelin du Val did not provide his specimens with any labels, but, according to the curator of Coleoptera in MNHN, A. Taghavian, he kept his types in his private collection. Therefore we presume that this specimen, which completely matches J. du Val’s description, is a syntype. The species was described based on an unknown number of specimens and therefore we designate the lectotype to fix the taxon identity for purpose of synonymy. Saprinus sicanus was described from Sicily (Italy) based on an unspecified number of specimens, therefore we designate the lectotype to fix the taxon identity for purpose of synonymy. Saprinus revisus was described from Algiers (Algeria) based on an unknown number of specimens, therefore we designate the lectotype to fix the taxon identity for purpose of synonymy. The type of S. curtus was found in mid-19 th century ‘Hungary’. This vague locality could refer to anywhere in the former Hungarian monarchy, which stretched south to the Adriatic Sea. It is possible that this species will be discovered in countries of the former Yugoslavia. It is a rather rare and seldom-collected species apparently spread around the Mediterranean Sea. Its rarity and slight morphological differences regarding dorsal punctation or course of carinal prosternal striae probably account for its numerous synonymies. Hypocaccus (Nessus) controversus (G. Müller, 1937) (Figs 45, 55–63) Hypocacculus controversus G. Müller, 1937: 115 (original description). Hypocacculus (Nessus) controversus: KRYZHANOVSKIJ & REICHARDT (1976): 204, 212 (keyed, redescription); MAZUR (1984): 89 (catalogue); MAZUR (1997): 252 (catalogue); MAZUR (2004):93 (catalogue). Hypocaccus (Nessus) controversus: MAZUR (2011): 208 (catalogue); LACKNER et al. (2015): 117 (catalogue). Type material examined. Hypocacculus controversus. LECTOTYPE (present designation): ♀ (Fig. 45), mounted on a triangular mounting card, right metatarsus missing, ‘ ♀ [written] // Banat 1909 / Herkulesbad / leg. M. Hilf / Coll. O. Leonhardt [printed] // sbsp. / controversus [written] // TYPUS [light-ochre label, printed] // scat. / Hist. 6 [yellow label, written] // Hypocacculus / (Nessus) / controversus / G. Müller, 1937 / LECTOTYPE / des. T. Lackner 2017 [red label, written]’ (CST). PARALECTOTYPES: 1 ♀, side-mounted on a triangular mounting point, left meso- and metatarsus missing, ‘Athen / Phaleron [written] // Da Scat. / 6 [yellow label, written] // Hypocacculus (Nessus) / controversus Müll. / Det. T. Lackner 2017 [printed-written] // Hypocacculus / (Nessus) / controversus / G. Müller, 1937 / PARALECTOTYPE / des. T. Lackner 2017 [red label, written]’ (CST). 1 ♀, ‘Saloniki / Schatzmayr [written] // Da Scat. / 6 [yellow label, written] // Hypocacculus (Nessus) / controversus Müll. / Det. T. Lackner 2017 [printed-written] // Hypocacculus / (Nessus) / controversus / G. Müller, 1937 / PARALECTOTYPE / des. T. Lackner 2017 [red label, written]’ (CST). Additional material examined. CYPRUS: Cyprus, no further data, 1 spec. (probably a male, genitalia lost), Baudi, (MFNB). GREECE: Greece, 1 ♁ (genitalia lost, sexed by the protarsi), 1 ♀, Emge lgt., C. & O. Vogt coll. (1 ♀ in CTLA, 1 ♁ in MSNG); Greece, 1 ♁, (MFNB). ATTICA: Attica, no further data, 2 ♀♀ (MFNB). CRETE: Lerapetra E, 0–20 m, 17.–23.iv.2000, 1 ♀, A. Kopetz ltg. (MSNG). IONIAN ISLANDS: Zante [=Zakynthos],Kalamaki,1909, 1♁, M.Hilf lgt., Coll. O. Leonhard (MNFB). JORDAN: IRBID: 5 km NE of El Karama, 31.iii.1994, 31.58°N, 35.36°E, 200 m, 1 ♀, S. Bečvář jun. & sen. lgt. (dubious identification) (MSNG); Toten Meer [= Dead Sea], 10.v.1963, 1 ♀, J. Klapperich lgt. (dubious identification) (MSNG). ROMANIA: BANAT: Banat, Orșova, 1909, 1 ♀, M. Hilf lgt., coll. O. Leonhard (MFNB). TUNISIA: DJERBA: Rass Taguernes, 10.–20.ii.1997, 1 ♀, Egger Manfred lgt. (dubious identification) (MSNG). TURKEY: IZMIR: Smyrna? [=Izmir], no further data, 1 ♁ (MFNB). Diagnostic description. This species is externally rather similar to the preceding species and therefore here we provide only the diagnostic description outlining the differences between the two taxa. Body (Fig. 45) somewhat more round and more flattened, light to dark brown, with light bronze hue (never with greenish hue). PEL: 2.00–2.30 mm; APW: 1.00–1.10 mm; PPW: 1.50–1.70 mm; EW: 1.65–1.90 mm; EL: 1.25–1.50 mm. Frontal disc more finely punctate than the one of H. (N.) curtus; pronotum medially almost impunctate. The first dorsal elytral stria is only slightly longer than the second (apically both striae 1–2 surpass slightly elytral half), never reaching ¾ of the elytral length apically (in H. curtus the first dorsal elytral stria is substantially longer, occasionally surpassing ¾ of elytral length apically). Sutural elytral stria always connected basally with fourth dorsal elytral stria (in H. curtus these two striae are joined only in specimens that belong to the ‘ cribellaticollis ’ form), can occasionally be shortened apically. Carinal prosternal striae strongly convergent apically, their apices very approximate, stopping posterad of united lateral prosternal striae; their united apices not forming a ‘loop’ as in H. curtus. MÜLLER (1937) mentioned another character: the mesoventral punctation is supposed to be denser and coarser in ‘ controversus ’ than in ‘ puncticollis ’ (= H. curtus). According to our observations, this is a valid, but not entirely stable character, since even among the few ‘ controversus ’ specimens we were able to examine we saw a specimen with only weak mesoventral punctation; the majority of specimens had their mesoventrite densely punctate. Male genitalia (Figs 55–63) are generally similar to the preceding species, the aedeagi differ most markedly: the one of H. (N.) curtus is sub-parallel and blunted apically, while the one of H. (N.) controversus is shorter, stouter, slightly dilated in apical third with acutely pointed apex (compare Figs 53 and 62). Note. The two female specimens from Jordan as well as the female from Tunisia are generally somewhat narrower, and their frons is adorned with coarse elongate rugae in place of dense punctures that are present
Wolfgang Lackner, Codices Chrysostomici Graeci. IV. Codices Austriae.
Wolfgang Lackner, Codices Chrysostomici Graeci. IV. Codices Austriae.. In: Revue des études byzantines, tome 40, 1982. p. 283
Wolfgang Lackner, Codices Chrysostomici Graeci. IV. Codices Austriae.
Wolfgang Lackner, Codices Chrysostomici Graeci. IV. Codices Austriae.. In: Revue des études byzantines, tome 40, 1982. p. 283
Liopygus occidentalis Lackner & Vienna 2017, sp. nov.
Liopygus occidentalis sp. nov. (Figs 14–22) Type material. HOLOTYPE: J, side-mounted on a triangular mounting card, genitalia extracted, dismembered and glued to the same mounting card as the specimen, with the following labels: ‘ YEMEN SOCOTRA Island / Skant area, 1300-1500 m / N 12°34′33″, E 54°01′31″ / 31.i.-1.ii.2010, L. Purchart lgt.’ [printed]; followed by: ‘ Liopygus occidentalis / sp.nov. HOLOTYPUS / det.T.Lackner & P.Vienna / 2017’ [red label,handwritten] (NMPC). ALLOTYPE: ♀, glued on its back on a triangular mounting card, with the following labels: ‘ YEMEN, Socotra Island / Al Haghier Mts. / Scant Mt. env. / 12°34.6′N, 54°01.5′E, 1450 m / Jiří Hájek leg. 12-13.xi.2010 ’ [printed]; followed by: ‘ Liopygus occidentalis / sp. nov. ALLOTYPUS / det. T. Lackner & P. Vienna / 2017’ (red label, hand-written) (NMPC). PARATYPE: YEMEN: SOCOTRA: J, side-mounted on a triangular mounting point, genitalia extracted, dismembered and glued to the same mounting card as the specimen, with labels identical to those of the allotype (TLCM). Description. Body (Figs 14–15). PEL = 2.60–2.80 mm; APW = 0.80–0.90 mm; PPW = 1.30–1.50 mm; EW = 1.40–1.60 mm; EL = 1.60–1.80 mm, flattened, elongate, color light castaneous-brown. Head. Eyes strongly flattened, not visible from above. Frontoclypeal area finely punctate, slightly depressed, traces of supraorbital striae visible above eye, other striae absent. Labrum flattened, rectangular, each mandible with large triangular subapical tooth. Maxillary palpi very prominent, terminal maxillary palpomere longer than mandible itself, visible from dorsal view. Mentum large, quadrate, labial palpi thick, terminal labial palpomere rather thick, its width approximately half its length. Submentum considerably smaller than mentum, triangular. Antennal scape massive, its length approximately equals to that of antennal funicle. Pedicel thickened, approximately as long as antennomeres III and IV together; club large, oval, setose, intersegmental sutures visible. Pronotum almost quadrate, along midline slightly broader than long, anterior angles strongly projected, acute. Marginal pronotal stria very thin, present only on apical angles and behind head; lateral pronotal stria slightly distanced from pronotal margin, parallel to it, terminating anteriorly on anterior pronotal margin. Pronotal disc impunctate; pronotal hypomeron asetose. Elytra impunctate, approximately twice as long as broad, lateral epipleura bistriate, striae thin, complete; subhumeral elytral striae absent, humeral elytral stria very thin, bifurcate. Elytral disc only with scattered microscopic punctation, dorsal elytral striae 1–4 complete, stria 5 almost complete, shortened basally. Sutural elytral stria absent. Propygidium broad, approximately 2.5 times as broad as long, with scattered deep ocellate punctures, separated by approximately 1.5–2.0 times their diameter. Pygidium in basolateral angles with two shallow depressions, each with deep fossa; pygidial surface basally with several vague shallow punctures, otherwise impunctate. Prosternal lobe (Fig. 15) large, rounded, with scattered microscopic punctation; marginal prosternal stria present, weakened to absent on apex. Carinal prosternal striae present as very vague rudiments on prosternal apophysis; lateral prosternal striae double, deeply impressed, subparallel and bisinuate; surface of composite ventral plate laterad to it with large deep ocellate punctures. Mesoventrite (Fig. 15) anteromedially deeply inwardly arcuate, marginal mesoventral stria present, thin, disc glabrous. Metaventrite (Fig. 15) flattened, with scattered microscopic punctation, lateral metaventral stria long, almost straight, nearly reaching metacoxa. Lateral disc of metaventrite with several large deep ocellate punctures; metanepisternum covered by elytral epileuron, not visible. Abdominal ventrite I (Fig. 15) with scattered microscopic punctation, striate laterally. Legs. Protibial spur prominent; protarsal groove deep; outer protibial margin with 5 rather prominent teeth topped by tiny denticle; teeth 2–3 widely separated; mesotibia on outer margin with two widely spaced short denticles, two additional longer denticles situated near tarsal insertion; outer margin ventrally with sparse row of tiny widely spaced denticles. Metatibia slightly longer than mesotibia, otherwise similar to it, except for there is only a single denticle on outer margin medially instead of two. Male genitalia. Sternite VIII (Figs 16–17) divided into two parts; tergite VIII and sternite VIII not fused (Fig. 18). Tergite IX (Fig. 19) divided into two parts; tergite X (Figs 19–20) small, its basal part wedged between the divided parts of tergite IX; sternite IX (Figs 19–20) or spiculum gastrale apically divided into two ‘tails’, pointed basally. Aedeagus (Fig. 21) tube-like, median lobe protruding from tegmen; phallobase (Fig. 22) approximately 1.5 times shorter than fused parameres. Differential diagnosis. The Socotran species strongly resembles the species Liopygus decemstriatus (Motschulsky, 1863) from Sri Lanka. It can, however, be readily distinguished from it based on the following characters: both dorsal elytral striae 2 and 4 are complete in the Socotran species (incomplete in L. decemstriatus); dorsal elytral stria 5 reaches even further basally in the Socotran species; propygidium of the Socotran species is entirely punctate (only partly punctate in L. decemstriatus); pygidium of the Socotran species is punctate only in its proximal part between the two fossa (pygidium of L. decemstriatus is punctate entirely). Comments. The genus Liopygus Lewis, 1891 contains 16 described species distributed exclusively in southeast Asia (MAZUR 2011).All Liopygus species, as far as known, live under bark, preying on small arthropods (DESBORDES 1919). Although DESBORDES (1919) published a key to the species of the genus, difficulties using it for species determination were voiced by GOMY (2006), who described the only species of Liopygus after the World War II. The discovery of a member of Liopygus in Socotra Archipelago, considerably far from the distribution area of the genus is highly interesting, and we advocate a revision of the genus. Etymology. Specific epithet, the Latin adjective ‘ occidentalis ’ (= western), refers to the disjunct geographical location of the new species. Distribution. Endemic to Socotra Island, so far known only from the highest part of Hagher mountains.Published as part of Lackner, Tomáš & Vienna, Pierpaolo, 2017, Histeridae of Socotra (Coleoptera: Histeroidea), pp. 55-76 in Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) (Acta. Ent. Mus. Natl. Pragae) 57 on pages 61-64, DOI: 10.1515/aemnp-2017-0107, http://zenodo.org/record/532469
Codices Chrysostomici Graeci. IV : Codices Austriae
Lackner Wolfgang. Codices Chrysostomici Graeci. IV : Codices Austriae. Aubervilliers : Institut de Recherche et d'Histoire des Textes (IRHT), 1981. 152 p. (Documents, études et répertoires de l'Institut de Recherche et d'Histoire des Textes, 15
Ctenophilothis Kryzhanovskij 1987
Ctenophilothis Kryzhanovskij, 1987 Ctenophilothis Kryzhanovskij, 1987: 25. Type species: Xenonychus chobauti Théry, 1900, original designation. Ctenophilothis: Olexa (1990): 143; Mazur (1997): 268; Mazur (2004): 91; Lackner (2010): 64, 90; Mazur (2011): 213. Diagnosis. Ctenophilothis was recently diagnosed by Lackner (2010), to which the reader is referred for a complete diagnosis of the genus. Differential diagnosis. Ctenophilothis is most similar to the species of the genus Philothis, differing from them chiefly by the shape of protibia that is adorned with two distal teeth topped by denticles followed by numerous long denticles in Ctenophilothis, whereas the species of the genus Philothis have their outer margin of protibia with two large triangular teeth topped with tiny denticle without any subsequent denticles (Lackner 2010). See the key to the Palaearctic genera of the Saprininae by the author for correct generic placement (Lackner 2010). Biology. Ctenophilothis is a psammophilous genus with two described species. The type specimen of the genus, C. chobauti, has been collected in sand, near rotten stems of Orobanchaceae. The rest of the known specimens, as far as their biologies are known, have been collected in sand near decaying plants. Nothing is known about the biology of C. altus. According to Kovarik & Caterino (2005) Ctenophilothis “inhabits sandy areas where shifting sand can bury and kill vegetation occurring there. Histerids will prey upon larvae of Diptera and Coleoptera that colonize the decaying plant material”. Ctenophilothis belongs to the strict psammophiles and its species are morphologically well adapted to life in sand. Most of the examined specimens have been taken in spring, with some specimens of C. chobauti collected also in October. In the southern Palaearctic, there are two Saprininae taxa associated with Orobanchaceae: Paravolvulus syphax (Reitter, 1904) known from Algeria, Tunisia, Morocco, and Saudi Arabia (Mazur 2011) recently collected also from Syria (Gomy 2013) and Chalcionellus hauseri (Schmidt, 1894) known from Turkmenistan, Iran, Kazakhstan and Mongolia (Mazur 2011). Both these taxa develop in the dry and decaying stalks of Cystanche (Orobanchaceae) (Kovarik & Caterino 2005). C. hauseri can be occasionally found together with its larvae within the stalks of Cystanche flava as well as in the surrounding sand in large numbers, where they prey upon flies of the genus Eumerus (Kryzhanovskij & Reichardt, 1976). Distribution. Ctenophilothis is an element of the Sahara desert: C. chobauti is known from Algeria and has recently been discovered in Morocco (Lackner 2010, Gomy 2011) and C. altus is known exclusively from Egypt (Mazur 2011), see also Fig. 27.Published as part of Lackner, Tomáš, 2013, Revision of the genus Ctenophilothis Kryzhanovskij, 1987 (Coleoptera: Histeridae: Saprininae), pp. 273-282 in Zootaxa 3691 (2) on page 274, DOI: 10.11646/zootaxa.3691.2.6, http://zenodo.org/record/25450
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