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Aulacophora opacipennis Chujo 1962
No full text<i>Aulacophora opacipennis</i> Chûjô, 1962 <p>(Figs 80–81, 100–107)</p> <p> <i>Aulacophora opacipennis</i> Chûjô, 1962: 98 (Taiwan).</p> <p> <b>Type material.</b> Holotype ♂ (TARI), labeled: “Kuraru [= Kenting, in Pingtung] / 31-VII-1931 / Col. T. Shiraki // Ho / Type [w, p, round label, red letters and border, but faded out] // Aulacophora / opacipennis / Chûjô [h] / DET. M. CHUJO [w, p] // 2312 [w, p]”. Paratypes: 1♀ (TARI), labeled: “Kuraru / 31-VII-1931 / Col. T. Shiraki [w, p] // ALLo / Type [w, p, round label, gray letters and border] // Aulacophora / opacipennis / Chûjô [h] / DET. M. CHUJO [w, p] // 2312 [w, p]”; 3♂, 2♀ (TARI), labeled: “Kuraru / 31-VII-1931 / Col. T. Shiraki [w, p] // Para / Type (w, p, round label, green letters and border) // Aulacophora / opacipennis / Chûjô [h] / DET. M. CHUJO [w, p] // 1463-1467 [w, p]”; 1♂, 1♀ (TARI), labeled: “ KUARU (sic!) [h] / FORMOSA [p] / 20.VI.1937 [h] / COL. M. CHUJO [w, p] // Para / Type [w, p, round label, green letters and border] // Aulacophora / opacipennis / Chûjô [h] / DET. M. CHUJO [w, p] // 1468 & 1470 [w, p]”; 1♀ (TARI), labeled: “Shiiago [= Maopu] Chikuto [= Chutung] / SHINCHIKU [= Hsinchu] / – 27–30.VI.1930 / Col. M. CHUJO [w, p] // Para / Type [w, p, round label, green letters and border] // Aulacophora / opacipennis / Chûjô [h] / DET. M. CHUJO [w, p] // 1469 [w, p]”; 1♂ (TARI), labeled: “Shizyukei (= Suchungchi, in Pingtung) / 1-VIII-1931 / Col. T. Shiraki [w, p] // Para / Type (w, p, round label, green letters and border) // Aulacophora / opacipennis / Chûjô [h] / DET. M. CHUJO [w, p] // 1462 [w, p]”; 1♀ (TARI), labeled: “HEITOU [= Pingtung City] / 22.V.1939 / Y. MIWA [w, p] // Para / Type (w, p, round label, green letters and border) // Aulacophora / opacipennis / Chûjô [h] / DET. M. CHUJO [w, p] // 1461 [w, p]”; 1♀ (TARI), labeled: “ Formosa / Shinchiku (= Hsinchu), - 18. / VII 1-30./ J. Sonan, [w, p] // Para / Type (w, p, round label, green letters and border) // Aulacophora / opacipennis / Chûjô [h] / DET. M. CHUJO [w, p] // 1460 [w, p]”; 1♂ (TARI), labeled: “ 7.VI.1914 / Taitô (= Taitung) [h] / Col. I. Nitobe [w, p] // Para / Type (w, p, round label, green letters and border) // Aulacophora / opacipennis / Chûjô [h] / DET. M. CHUJO [w, p] // 1460 [w, p]”.</p> <p> <b>Other specimens examined. TAIWAN</b>. Hsinchu: 2♂, 3♀, Wufeng, 14–16.VII.1982, leg. K. C. Chou & C. C. Pan (TARI, BMNH); Ilan: 1♂, Mt. Taiheizan (= Taipingshan), 23.VII.1940, leg. M. Chujo (TARI); Pingtung: 1♀, Sheting, 4.XI.2009, leg. M.-H. Tsou (RBCN); 1♂, Suchunghsi, 13.V.2013, leg. Y.-T. Chung (RBCN).</p> <p> <b>Diagnosis.</b> <i>Aulacophora opacipennis</i> is similar to <i>A. apicipes</i> with yellowish brown body, and black and opaque elytron but <i>A. opacipennis</i> is easily separated from <i>A. apicipes</i> by the yellowish brown legs and black elytron (black tibia and tarsi, yellow apical margin of elytron in <i>A. apicipes</i>).</p> <p> <b>Males.</b> Length 6.2 mm, width 3.1 mm. General color (Figs 80–81) yellowish brown except elytron black and opaque. Antenna (Fig. 100) filiform, antennomere I not enlarged, apico-lateral angles of antennomeres III–IV produced anteriorly; ratio of length of antennomeres III to XI about 1.0: 1.2: 1.1: 1.1: 1.1: 1.1: 1.1: 1.1: 1.5; ratio of length to width from antennomere III to XI about 2.7: 3.3: 2.7: 2.7: 2.9: 3.2: 3.5: 3.7: 5.4. Abdominal tergite forming notch (Fig. 106), weakly convex at middle; base weakly sclerotized. Median lobe of fifth abdominal ventrite rectangular, with median longitudinal ridge represented from middle extending beyong base, median longitudinal internal ridge well developed; disc depressed at apical area. Penis (Figs 102–103) wide, abruptly narrowed from apical 1/6 to apex, apex pointed, less widened at basal 2/5; curved in lateral view; tectum well sclerotized, apex pointed, abruptly widened at basal 1/3, lateral margin with triangular sclerite recurved at basal 1/3; endophallus with several stout setae scattered, and with one longitudinal sclerite, apex recurved and rounded, basally widened.</p> <p> <b>Females.</b> Length 7.5–7.9 mm, width 3.9–4.3 mm. Similar to male, but antenna more slender (Fig. 101), and apico-lateral angles of antennomeres III–IV less produced anterior; ratio of length of antennomeres III to XI about 1.0: 1.0: 1.0: 1.0: 1.0: 0.9: 0.9: 0.9: 1.0; ratio of length to width from antennomere III to XI about 3.7: 3.8: 4.2: 4.2: 4.2: 4.2: 4.4: 4.2: 5.5. Apical margin of abdominal ventrite V truncate. Gonocoxae (Fig. 104) slender, apex of each gonocoxa with eight setae from apical 1/6 to apex; gonocoxae connected at middle, base widened. Ventrite VIII (Fig. 105) weakly sclerotized; apex narrow, apical margin a little emarginate at middle, surface with dense short setae inside and along apical margin, and with extremely long setae near apical margin; spiculum short. Spermathecal receptaculum (Fig. 107) a little swollen, hardly separated from pump; pump strongly curved; spermathecal duct short, stout, and strongly curved, shallowly projecting into receptaculum.</p> <p> <b>Host plant.</b> Unknown.</p> <p> <b>Distribution.</b> Endemic to Taiwan.</p>Published as part of <i>Lee, Chi-Feng & Beenen, Ron, 2015, Revision of the genus Aulacophora from Taiwan (Coleoptera: Chrysomelidae: Galerucinae), pp. 151-190 in Zootaxa 3949 (2)</i> on pages 176-178, DOI: 10.11646/zootaxa.3949.2.1, <a href="http://zenodo.org/record/237252">http://zenodo.org/record/237252</a>
Phygasia diluta Chujo 1963
No full textPhygasia diluta Chûjô, 1963, status resurrected (Figs 4, 5, 10, 11, 15, 18, 21, 24, 26–32) Phygasia ornata: Chûjô, 1936: 18 (part); Kimoto, 1991: 24 (part). Phygasia ornata diluta Chûjô, 1963: 400; Kimoto, 1966: 35 (part); Kimoto and Takizawa, 1997: 411 (synonymized with P. ornata). Phygasia diluta: Kimoto, 1971: 78 (raised to species rank). Phygasia taiwanensis Ge et al., 2010: 325. New synonym Description. Male. Length 5.8–6.4 mm, width 3.1–3.5 mm. General color (Fig. 4) yellowish-brown; antennomeres III-XI darkened, tibia basally darkened; elytron white; one transverse black spot at humerus and subapically, apex orange, with narrow transverse black band between white and orange areas. Pronotum about 1.8 times wider than long, disc smooth, with scattered fine punctures. Elytra 1.4 times longer than wide, lateral margin slightly rounded, widest just behind middle, disc with densely distributed small punctures and randomly occurring large punctures. Antennomeres III-VII widened (Fig. 10); ratio of length of antennomeres III to XI about 1.0: 0.8: 0.9: 0.9: 0.9: 0.9: 0.8: 0.8: 1.0, ratio of length to width of antennomeres III to XI about 1.7: 1.4: 1.5: 1.6: 1.8: 1.8: 2.1: 2.5: 3.2. Penis (Figs 15 a, b) elongate, about 4.3 X longer than wide, apex of tectum a little lower than that of penis, subparallel-sided, strongly narrowed subapically, apex rounded, moderately curved in lateral view; ventral side evenly convex. Female. Length 5.3 –7.0 mm, width 3.0–4.0 mm. Similar to males, but antennomeres III-VIII narrower (Fig. 11), ratio of length of antennomeres III to XI about 1.0: 1.0: 1.1: 1.0: 1.0: 1.0: 0.8: 0.8: 1.2, ratio of length to width of antennomeres III to XI about 2.1: 2.0: 2.2: 2.2: 2.3: 2.6: 2.4: 2.6: 4.0. Gonocoxa (Fig. 18) medially connected, apex widely rounded, base weakly sclerotized, three long setae at apex, three long setae at outer side. Spermatheca (Fig. 21) with receptacle strongly swollen; pump strongly curved, apex rounded; proximal spermathecal duct wide. Spiculum of sternite VIII (Fig. 24) longer then P. chengi but shorter than P. o r n a t a. Bursa-sclerites highly variable. Some individuals with only one pair of bursa-sclerites, each with single seta; others with two pairs of bursa sclerites, dorsal pair (Figs 27 a, 28 a) better developed, with three or four prominent setae, base well-sclerotized; ventral pair (Figs 27 b, 28 b) with one to three prominent setae but base weakly-sclerotized. Color variation. In some specimens, the transverse black spots on humerus and near elytral apex form transverse bands across elytral base and apex and the antennomeres III-XI, tibia, and tarsus are almost black (Fig. 5). Diagnosis. Phygasia diluta differs from P. ornata and P. chengi by its yellow-brown or dark brown antenna, tibia, and tarsus (antenna, tibia, and tarsus black in P. ornata and P. chengi); relatively wider penis (4.3 times longer than wide) compared to P. ornata (4.6 times longer than wide) and narrower penis compared to P. chengi (3.9 times longer than wide); relatively longer sternite VIII compared to P. chengi and shorter sternite VIII compared to P. ornata.. Type material examined. Holotype 3 (Fig. 29) of Phygasia ornata diluta Chûjô labeled (Fig. 30): “ Formosa Sauter / Takao (= Kaoshiung city) 1907. VIII. 1 / Holotypus Phygasia ornata ssp. diluta Chujo / Holotype / Phygasia ornata diluta CHUJO Det. M. CHUJO, 1961 ” (HNHM). Allotype Ƥ (Fig. 31) labeled (Fig. 32): “ Formosa Sauter / Kanshirei (= Kuangtyling, Tainan county) 908. VI. 7 -(19) 15. / Allotypus Phygasia ornata ssp. diluta Chujo / Allotype / Phygasia ornata diluta CHUJO Det. M. CHUJO, 1961 ” (HNHM). Material examined (27 specimens). 1 Ƥ, Pingtung, Kankau (= Changkou), 27.V. 1932, leg. R. Takahashi (TARI); 1 Ƥ, Pingtung, Kuaru (= Kenting), 15.VI. 1937, leg. M. Chujo (TARI); 1 Ƥ, same locality, 18–23.III. 1981, leg. K. S. Lin & T. Lin (TARI); 13, same locality, 22-26.III. 1982, leg. T. Lin & S. C. Lin (TARI); 1 Ƥ, same locality, 24.XI. 2009, leg. C.-F. Lee (TARI); 43, 4ƤƤ, Pingtung, Sheting, 15.VIII. 2009, leg. S.-F. Yu (TARI); 23, 5ƤƤ, same but with “leg. M.-H. Tsou” (TARI); 23, 2ƤƤ, Pingtung, Shuangliu, 19.VII. 2007, leg. S.-F. Yu (TARI); 13, same locality, 30.V. 1997, leg. C. W. & L. B. O’Brien (NMNS); 1 Ƥ, Pingtung, Tahanlintao, 22.VIII. 2011, leg. J.-C. Chen (TARI); 13, Taitung, 4km N. Chinglun, 30.V. 1997, leg. C. W. & L. B. O’Brien (NMNS). Host plant. Asclepiadaceae: Gymnema sylvestre (Retz.) Schultes. Distribution. Southern Taiwan (Fig. 26). This species is restricted to lowlands.Published as part of Lee, Chi-Feng, 2012, The genus Phygasia Chevrolat (Coleoptera: Chrysomelidae: Galerucinae: Alticini) in Taiwan, pp. 27-37 in Zootaxa 3256 on pages 30-32, DOI: 10.5281/zenodo.21050
Morphosphaera bimaculata Chujo
No full textMorphosphaera bimaculata Chûjô (Figs 3–6) Morphosphaera bimaculata Chûjô, 1938: 136 (Taiwan); Chûjô, 1962: 175 (redescription); Kimoto, 1969: 38 (additional records); Wilcox, 1971: 218 (list); Kimoto, 1989: 252 (additional records); Takizawa et al., 1995: 11 (additional records); Kimoto & Chu, 1996: 72 (list); Kimoto & Takizawa, 1997: 305 (key), 380 (list); Lee & Cheng, 2007: 115; Beenen, 2010: 462 (list); Yang et al., 2015: 189 (key, list). Type material. Lectotype Ƌ (TARI), here designated, labeled: “ARISAN / FORMOSA / 24.X.1933 [h] / COL. M. CHUJO [p, w] // CO / Type [p, w, yellow letters, circle label with yellow border] // Morphosphaera / bimaculata / CHÛJÔ [h] / DET. M. CHUJO [p, r] // 1371 [p, w]”. Paralectotypes: 1Ƌ (TARI), same as lectotype, but with “2350 [p, w]”; 1♀ (SDEI): “ Arisan, 1918. / X 2-23. / J. Sonan, [p, w] // Syntypus [p, r] // Morphosphaera / bimaculata / CHÛJÔ [h] / DET. M. CHUJO [p, g] // DEI Müncheberg / Col-05713 [p, g]”; 1Ƌ (TARI): “ Formosa / Karenko, - 19. / VII 20 -VIII 4. / T. Okuni, [p, w] // CO / Type [p, w, yellow letters, circle label with yellow border] // Morphosphaera / bimaculata / CHÛJÔ [h] / DET. M. CHUJO [p, r] // 1485 [p, w]”; 1♀ (TARI), same but with “1486 [p, w]”; 1Ƌ (TARI), same but with “1487 [p, w]”; 1Ƌ (SDEI): “ Formosa / Karenko, - 19. / VII 20 -VIII 4. / T. Okuni, [p, w] // Syntypus [p, r] // Morphosphaera / bimaculata / CHÛJÔ [h] / DET. M. CHUJO [p, g] // DEI Müncheberg / Col-05712 [p, g]”; 1Ƌ (TARI): “KARENKÔ [h] / FORMOSA [p] / 15.VII.1935 [h] / COL. M. CHUJO [p, w] // CO / Type [p, w, yellow letters, circle label with yellow border] // Morphosphaera / bimaculata / CHÛJÔ [h] / DET. M. CHUJO [p, W] // 1372 [p, w]”; 1♀ (TARI): “ Jujiro / 26-IV 1931 / Col. T. Shiraki [p, w] // CO / Type [p, w, yellow letters, circle label with yellow border] // Morphosphaera / bimaculata / CHÛJÔ [h] / DET. M. CHUJO [p, r] // 1484 [p, w]”. Description. Length 6.8–8.4 mm, width 4.4–5.6 mm. Head, scutellum, meso- and metathoracic, and abdominal ventrites reddish brown; antenna black except two basal antennomeres; abdominal ventrites medially darker; elytron bluish metallic; leg reddish brown but tibia and tarsus darkened (Figs 3 A–3C); prothorax yellow, pronotum with one pair of extremely large black spots at sides, and one median, wide, longitudinal brown band (Fig. 3 D). Antenna filiform (Fig. 4 A), 0.5x as long as body; length ratios of antennomeres II to XI about 1.0: 1.5: 1.8: 1.8: 1.8: 1.8: 1.7: 1.7: 1.6: 1.8, and length to width ratios of antennomeres II to XI about 1.6: 2.2: 2.6: 2.6: 2.6: 2.6: 2.6: 2.7: 2.5: 3.4. Aedeagus (Figs 4 B–4C) wide in dorsal view, about 6.5x longer than wide, parallelsided; apex tubelike with small rounded process at middle; ventral surface well sclerotized and smooth; narrow and slightly curved in lateral view; with four stout setae near apex of ventral surface of internal sac; apical processes of endophallic sclerite conjunct, apex rounded; apico-lateral process widely rounded; with one pair of processes erect and recurved outwards at apical 1/3; base tubelike. Gonocoxae reduced. Ventrite VIII (Fig. 4 D) with short spiculum; apex transverse, wide, with dense setae apically. Receptacle of spermatheca (Fig. 4 E) elongate, and slightly swollen, pump much narrower and longer, extremely curved; proximal spermathecal duct extremely short and wide. Diagnosis. Morphosphaera bimaculata is similar to M. montivaga in possessing one pair of black spots at sides and one median, wide, brown band on pronotum. But M. bimaculata is much smaller (M. montivaga is over 10 mm), with bluish or greenish metallic elytra (reddish brown elytra with purple reflections in M. montivaga). This species also lacks black stripes within the brown band on the pronotum, and the abdominal ventrites are reddish brown but medially darkened (abdominal ventrites blackish brown but sides yellowish brown in M. montivaga). Host plants. Ficus erecta Thunb. var. beecheyana (Hook. & Arn.) King, F. fistulosa Reinw. ex Blime, F. pumila L. var. awkeotsang (Makino) Corner, F. sarmentosa B. Ham. ex J. E. Sm. var. nipponica (Fr. & Sav.) Corner (Moraceae) (Lee and Cheng 2007). Biology. Morphosphaera bimaculata populations are presumably bivoltine. Adults have been documented as the overwintering stage. They hide between crevices of bark (Fig. 5 A) or leaves (Fig. 5 B). They become active during early or middle February. Based on laboratory rearing at 20–25ºC, females laid 29– 39 eggs in a single egg mass (Fig. 5 C). The eggs hatched in nine days. The larvae fed on leaves and the larval duration was 15 days (Figs 5 D–5F). Mature larvae (Fig. 5 G) crawled into the soil and built underground chambers for pupation. The pupal stage (Fig. 5 H) lasted approximately 18 days. The newly emerged adults went into summer dormancy during July, and became active during autumn. Larvae were again found during October. Other material examined. TAIWAN. Ilan: 2 exs., Fushan Botanical Park, 8.V.2008, leg. S.- F. Yu (RBCN); Kaoshiung: 17 exs., Erchituan, 31.III.2015, leg. B.- X. Guo (BPBM, CAS, MCSN, NHRS); 1 ex., Tengchih, 4.VII.2011, leg. M.- H. Tsou (TARI); 3 exs., same locality, 29.V.2013, leg. Y.- T. Chung (RMNH); 13 exs., same locality, 15.VI.2013, leg. B.- X. Guo (BMNH, NME, NHMB); 2 exs., same locality, 5.X.2013, leg. W.- C. Liao (ZSM); Nantou: 1 ex., Wanfengtsun, 12.IV.2007, leg. W.- T. Liu (TARI); 1 ex., same locality, 9.I.2008, leg. W.- T. Liu (TARI); Pingtung: 1 ex., Shuangliu, 4.V.2005, leg. J.- F. Tsai (TARI); 1 ex., Tahanshan, 18.VII.2007, leg. C.- F. Lee (MTD); 1 ex., same locality, 14.III.2011, leg. J.- C. Chen (MTD); 1 ex., same locality, 26.III.2013, leg. C.- F. Lee (TARI); 1 ex., 3.VI.2013, leg. J.-C. Chen (TARI); 2 exs., Wutai, 7.X.2012, leg. S.- F. Yu (MTD); Taipei: 1 ex., Fushan, 18.III.2014, leg. H.- J. Chen (SMNS); 1 ex., Kuanyinshan, 18.V.2011, leg. H. Lee (TARI); 1 ex., Manyuehyuan, 11.XI.2007, leg. M.- H. Tsou (IRSB); Taitung: 1 ex., Guanshan, 31.X.2009, leg. P.- F. Wang (TARI); Taoyuan: 1 ex., Hsuanyuan, 18.VI.2015, leg. H. Lee (IRSB); 1 ex., Tungyangshan, 12.IV.2007, leg. S.- F. Yu (TARI); 1 ex., 10.V.2009, leg. M.-H. Tsou (IRSB). Distribution. Endemic to Taiwan. Morphosphaera bimaculata is parapatric with M. chrysomeloides. This species inhabits high elevations (500–1000 m) and M. chrysomeloides occurs in lowlands (below 1000 m) (Fig. 6).Published as part of Lee, Chi-Feng & Bezdĕk, Jan, 2016, Revision of the genus Morphosphaera Baly (Coleoptera: Chrysomelidae: Galerucinae), pp. 1-41 in Zootaxa 4179 (1) on pages 6-11, DOI: 10.11646/zootaxa.4179.1.1, http://zenodo.org/record/26223
Aulacophora kotoensis Chujo
No full text<i>Aulacophora kotoensis</i> Chûjô <p>(Figs 4–7, 62–65, 67, 69–72)</p> <p> <i>Aulacophora kotoensis</i> Chûjô, 1962: 79 (Taiwan).</p> <p> <b>Type material.</b> <i>Aulacophora kotoensis</i>: holotype ♂ (TARI), labeled: “KOTOSHO / (BOTEL-TOBAGO IS.) (= Lanyu island) / FORMOSA / 20.VI–10.VII.1938 / COLL. M. CHUJO [w, p] // Ho / Type [w, p, round label, red letters and border, but faded out] // Aulacophora / kotoensis / Chûjô [h] / DET. M. CHUJO [w, p] // 1813 [w, p]”. Paratypes: 1♀ (TARI), labeled: “KOTOSHO / (BOTEL-TOBAGO IS.) (= Lanyu island) / FORMOSA / 20.VI– 10.VII.1938 / COLL. M. CHUJO [w, p] // ALLo / Type (w, p, round label, gray letters and border) // Aulacophora / kotoensis / Chûjô [h] / DET. M. CHUJO [w, p] // 1811 [w, p]”; 1♂, 1♀ (TARI), labeled: “KOTOSHO / (BOTEL- TOBAGO IS.) (= Lanyu island) / FORMOSA / 20.VI–10.VII.1938 / COLL. M. CHUJO [w, p] // Para / Type (w, p, round label, green letters and border) // Aulacophora / kotoensis / Chûjô [h] / DET. M. CHUJO [w, p] // 1820 & 676 [w, p]”; 1♂, 1♀ (TARI), labeled: “KOTOSHO / FORMOSA / IV.1936 / COLL. Y. CHUJO [w, p] // Para / Type (w, p, round label, green letters and border) // Aulacophora / kotoensis / Chûjô [h] / DET. M. CHUJO [w, p] // 1823 & 684 [w, p]”; 2♂, 2♀ (TARI), labeled: “Kotosho / 10 III–14.IV.1920 / Sonan [w, h] // Para / Type (w, p, round label, green letters and border) // Aulacophora / kotoensis / Chûjô [h] / DET. M. CHUJO [w, p] // 1422-1424, 2599 [w, p]”; 2♂ (TARI), labeled: “Kotosho / 20.IX.1923 / Col. T. Okuni. [w, p] // Para / Type (w, p, round label, green letters and border) // Aulacophora / kotoensis / Chûjô [h] / DET. M. CHUJO [w, p] // 1425 & 1426 [w, p]”; 1♂, 2♀ (TARI), labeled: “ Formosa / Kotosho / III–IV.1932 / S. Hirayama [w, p] // Para / Type (w, p, round label, green letters and border) // Aulacophora / kotoensis / Chûjô [h] / DET. M. CHUJO [w, p] // 681-683 [w, p]”; 2♂, 2♀ (TARI), labeled: “[Koutousyo] / FORMOSA / XII-1931 - II-1932 / S. Hirayama // Para / Type (w, p, round label, green letters and border) // Aulacophora / kotoensis / Chûjô [h] / DET. M. CHUJO [w, p] // 677-680 [w, p]”.</p> <p> <b>Other specimens examined. TAIWAN</b>. Taitung: 1♂, Lanyu Is., 22–26.III.1998, leg. C.-F. Lee (TARI); 3♂, same locality, 29.III.2009, leg. U. Ong (RBCN); 1♂, 1♀, same locality, 26.IV.2009, leg. U. Ong (BMNH); 1♂, same locality, 17.III.2012, leg. T.-H. Lee (MNHUB); 2♂, same locality, 17.III.2012, leg. M.-H. Tsou (MNHUB); 2♂, same locality, 5.V.2012, leg. S.-F. Yu (TARI); 2♂, same locality, 14.IV.2013, leg. B.-X. Guo (TARI); 2♂, 14.IV.2013, leg. B.-X. Guo (TARI); 1♂, 22.XI.2013, leg. Y.-T. Chung (TARI); 1♂, 2♀, Hsiaolanyu Is., 26.V.2009, leg. Y.-T. Chung (TARI); 2♂, 1♀, same locality, 25.VI.2009, leg. Y.-T. Chung (TARI); 1♀, same locality, 27.VIII.2009, leg. Y.-T. Chung (TARI).</p> <p> <b>Diagnosis.</b> <i>Aulacophora kotoensis</i> is similar to <i>A. indica</i> with the enlarged first antennomere in male and conelike pygidium projecting from elytral apices in female, but <i>A. kotoensis</i> has a pair of tubercles on the pronotum in male (without tubercles in <i>A. indica</i>), slender and black pygidium in female (wide and yellowish brown pygidium in <i>A. indica</i>).</p> <p> <b>Males.</b> Length 6.5–7.1 mm, width 3.5–3.8 mm. General color (Figs 4–5) yellowish brown but metathoracic and abdominal ventrites, middle and hind legs black; outer margins of femur and tibia, and tarsi of front legs black; labrum dark brown; antenna dark brown except three basal antennomeres yellowish brown. Antenna (Fig. 62) filiform and slender, antennomere I enlarged; ratio of length of antennomeres III to XI about 1.0: 1.1: 1.1: 1.1: 1.1: 1.0: 1.0: 1.0: 1.2; ratio of length to width from antennomere III to XI about 3.9: 3.4: 3.5: 3.5: 3.4: 3.2: 3.6: 3.6: 4.7. Pronotum with deep, transverse groove; one pair of tubercles behind groove. Elytra with cluster of erect hairs behind humerus. Abdominal tergite VIII well sclerotized (Fig. 67), apical margin with wide triangular incision and apices acute. Median lobe of abdominal ventrite V rectangular, with median, longitudinal, wide groove and abbreviated near apex. Penis (Figs 64–65) slender, parallel-sided, abruptly narrowed at apical 1/7 and apically tapering, apex recurved; slightly curved from middle to apex in lateral view; tectum laterally sclerotized; endophallus with clusters of short setae, and with one longitudinal sclerite, apically tapering, apex strongly curved in lateral view, basally widened.</p> <p> <b>Females.</b> Length 7.2 mm, width 3.9 mm. Similar to male (Figs 6–7), but pronotum with transverse groove shallow, and without tubercles; cluster of erect hairs on elytra absent; antennomere I not enlarged (Fig. 63); ratio of length of antennomeres III to XI about 1.0: 1.1: 1.0: 1.0: 1.0: 1.0: 1.0: 1.0: 1.2; ratio of length to width from antennomere III to XI about 3.0: 3.2: 3.2: 3.3: 3.3: 3.2: 3.2: 3.2: 4.4. Pygidium projecting from elytral apex, parallel-sided, slender, and apex rounded. Middle of apical margin of abdominal ventrite V (Fig. 69) emarginate, weakly convex at middle of emargination. Gonocoxae (Fig. 70) slender, apex of each gonocoxa with seven or eight setae from apical 1/6 to apex; gonocoxae connected at middle, base slender. Ventrite VIII (Fig. 71) weakly sclerotized; apex narrow, apical margin emarginate at middle, with dense short setae along apex; spiculum short. Spermathecal receptaculum (Fig. 72) a little swollen; pump strongly curved; spermathecal duct short, stout, shallowly projecting into receptaculum.</p> <p> <b>Host plant.</b> Cucurbitaceae: <i>Trichosanthes quinquangulata</i> A. Gray (Chûjô 1962; present study).</p> <p> <b>Remarks.</b> To exclude conspecifity between <i>Aulacophora kotoensis</i> and <i>A. abdominalis</i> (Fabricius, 1781) it was necessary to study the type specimens of <i>Crioceris abdominalis</i> in the Fabricius-collection. The results of the study of these specimens are treated extensively in the section on <i>Aulacophora abdominalis</i> and partly in the section on <i>A. indica</i>. For the status of <i>A. kotoensis</i> it is relevant that we have found no arguments to treat this species as conspecific with <i>A. abdominalis</i>.</p> <p> <b>Distribution.</b> Lanyu and Hsiaolanyu islands (Taiwan). Hsiaolanyu is very small and deserted island which is very close to Lanyu Island. The size of island is about 1.75 km 2.</p>Published as part of <i>Lee, Chi-Feng & Beenen, Ron, 2015, Revision of the genus Aulacophora from Taiwan (Coleoptera: Chrysomelidae: Galerucinae), pp. 151-190 in Zootaxa 3949 (2)</i> on pages 167-169, DOI: 10.11646/zootaxa.3949.2.1, <a href="http://zenodo.org/record/237252">http://zenodo.org/record/237252</a>
Tenebrionid Beetles of the Genus Promefhis Pascoe from Nansei Islands, Japan (CoIeoptera)
No full textSeven species of the Promethis Pascoe, 1869 from Nansei Islands are enumerated with a key to the species based on male. P. tokunoshimana sp. nov. is described. The records of P. oshimana from Takarajima Is. by Kaszab and Tokunoshima by M. T. Chujo are obliterated from the distribution
The Theme of “ASADI-GA-TUYU” : Linking the Story of NII-NO-CHUJO
No full textThe context in this story that NII-NO-CHUJO looks for the substitute steady for his first lover appears to be far from the theme of the story, proposed by the author of this report, “the revival of the SOCHI-NO-MIYA family.” This report proposes to bring to light how NII-NO-CHUJO's love affair highlights the superiority of GEN-NO-CHUJO's daughter, a descendant of the SOCHI-NO-MIYA family, by comparing her with other girlfriends, and demonstrates that his association with her is of great importance to the SOCHINO-MIYA family. This study proves more clearly that the theme of the story is “ the revival of the SOCHI-NO-MIYA family.” In addition, this report shows that the main characters are managed and manipulated by GEN-NOCHUJO who is told that he had flourished before the age of the story, and proposes that the revival of the family seems to have been orchestrated by GEN-NO-CHUJO behind the scenes, in pursuit of the happiness and glory of his daughter
Siemssenius metallipennis Chujo
No full textSiemssenius metallipennis (Chûjô) (Figs 1 C–1D, 5A– 5I) Pseudoliroetis metallipennis Chûjô, 1962: 181; Kimoto, 1969: 36 (additional records); Takizawa et al., 1995: 12; Kimoto & Chu, 1996: 70 (catalogue). Siemssenius metallipennis: Kimoto & Takizawa, 1997: 181; Zhang et al., 2008: 127 (key). Type locality. Chiayi county, Alishan (阿里山), 23°30’32”N, 120°48’07”E, 2200 m. Type material (n= 2). Holotype ♂ (head and prothorax lost but glued back with those of Basilepta varians Chûjô, 1956; TARI), labeled: “ Arisan (= Alishan, 阿里山) / FORMOSA / 24–25.V.1933 / Col. M. CHUJO [p, w] / / Pseudoliroetis / metallipennis / CHÛJÔ [h] // DET M. CHUJO [p, w] // Ho / Type [p, w, circle label with red letters and border but faded out] // 2366 [p, w]”. Paratype: 1♀ (TARI), labeled: “Mt. ARI- SAN [h] / FORMOSA [p] / 7.VII.1929 [h] / COL. [p] M. CHÛJÔ [h, w] // Pseudoliroetis / metallipennis / CHÛJÔ [h] // DET M. CHUJO [p, w] / / ALLo / Type [p, w, circle label with gray letters and border] // 2365 [p, w]”. Additional specimens (n= 19). Chiayi: 2♂♂ (TARI), Alishan (阿里山), 20–23.VI.1956, leg. S. C. Chiu; 2♀♀ (SEHU), same locality, 4–5.VII.1975, leg. H. Takizawa; 1♂, 4♀♀ (SEHU), same locality, 9–10.VII.1981, leg. H. Takizwa; Nantou: 4♂♂, 2♀♀ (TARI), Tatachia (塔塔加), 18.V.2010, leg. M.- H. Tsou; 1♀ (TARI), same locality, 10.V.2011, leg. C.- F. Lee; 1♀ (TARI), same locality, 9.VII.2014, leg. C.- F. Lee; 1♂, 1♀ (TARI), same locality, 1.VII.2015, leg. J.- C. Chen. Description. Length 7.9–9.4 mm, width 4.1–5.5 mm. General color yellow, but head, scutellum, femora, tarsi, and apices of tibiae black, antennae apically paler; elytra greenish or purplish bronze (Figs 1 C–1D). Antenna filiform, relatively longer in male (Fig. 5 A), as long as body, antennomeres III–V apically widened, length ratios of antennomeres III to XI about 1.0: 1.3: 1.3: 1.4: 1.4: 1.3: 1.3: 1.3: 1.6, and length to width ratios of antennomeres III to XI about 2.5: 3.5: 3.5: 4.0: 4.2: 4.5: 4.6: 5.2: 7.7; relatively shorter in female (Fig. 5 B), about 0.9x as long as body, antennomeres III–V apically widened, length ratios of antennomeres III to XI 1.0: 1.1: 1.2: 1.0: 1.1: 1.1: 1.0: 1.0: 1.4, and length to width ratios of antennomeres III to XI 3.2: 3.6: 3.8: 3.6: 4.0: 4.2: 4.3: 4.4: 5.9. Apical margin of abdominal ventrite V with two well developed incisions in male, surface with deep, wide, longitudinal groove, apically expanding into most of area between incisions; apical margin of ventrite IV with very small incision midway (Fig. 5 E). Aedeagus concave and basally widened, weakly curved in lateral view; apico-lateral angles obtusely angulate, narrow in lateral view, base obtusely angulate; middle of apical margin with a long, broad tube-like process, wide in lateral view, apex with median notch; dorsal process subapically and strongly swollen in lateral view but weakly swollen in dorsal view, apex pointed and triangulate in lateral view (Figs 5 C–5D). Abdominal ventrite V not modified in female, apical margin truncate, disc without depression (Fig. 5 F); apical of margin of ventrite IV straight. Gonocoxae reduced. Ventrite VIII (Fig. 5 H) heartshaped, with short spiculum, with scarce elongate setae along apical margins of triangulate processes. Tergite VIII (Fig. 5 G) with apex rounded and dense setae on disc. Receptacle of spermatheca (Fig. 5 I) weakly swollen, barely separated from pump, pump much narrower and longer, sharply curved; proximal spermathecal duct long and narrow. Diagnosis. See diagnosis of Siemssenius cheni sp. nov. Host plant. Lonicera acuminata Wall., 1832 (Caprifoliaceae) (Figs 3 B–3C). Distribution. Two localities at mountain areas in south Taiwan (Nantou and Chiayi counties) where it is sympatric with S. rufipennis (Chûjô, 1962) (Fig. 4).Published as part of Lee, Chi-Feng, 2016, Review of the genus Siemssenius Weise, 1922 (Coleoptera: Chrysomelidae: Galerucinae) from Taiwan, with descriptions of five new species, pp. 367-384 in Zootaxa 4158 (3) on pages 371-374, DOI: 10.11646/zootaxa.4158.3.4, http://zenodo.org/record/25832
Tenebrionid Beetles of the Genus Promefhis Pascoe from Nansei Islands, Japan (CoIeoptera)
No full textSeven species of the Promethis Pascoe, 1869 from Nansei Islands are enumerated with a key to the species based on male. P. tokunoshimana sp. nov. is described. The records of P. oshimana from Takarajima Is. by Kaszab and Tokunoshima by M. T. Chujo are obliterated from the distribution
Monolepta gracilipes Chujo 1938
No full textMonolepta gracilipes Chûjô, 1938 reinstated Luperodes pallidulus Chûjô, 1935: 162. Monolepta pallidula Chûjô, 1938: 144; Chûjô, 1962: 123; Chûjô, 1963 a: 25; Chûjô, 1963 b: 393; Chûjô, 1965: 97; Kimoto, 1965: 488; Kimoto, 1966: 32; Kimoto, 1969: 47; Kimoto, 1986: 58; Kimoto, 1987: 189; Kimoto, 1989: 256; Kimoto, 1991: 14; Takizawa et al., 1995: 11. Monolepta gracilipes Chûjô, 1938: 145; Chûjô, 1962: 132; Chûjô, 1965: 97; Kimoto, 1969: 47 (synonymized with M. pallidula); Kimoto, 1986: 58. Description. Total length: males: 5.1–6.4 mm (O: 5.63 mm; n= 6); females: 4.9–6.2 mm (O: 5.73 mm; n= 6). General color yellowish-brown (Fig. 1); in some specimens antenna, apex of tibia, and tarsi dark brown. Antennomeres 4–10 about 6.2 times longer than broad at apex (Fig. 13); length of second to third antennomere 0.79 –1.00 (O: 0.87); length of third to fourth antennomere 0.36–0.50 (O: 0.59) (Fig. 12). Pronotal length to width 0.65–0.73 (O: 0.69). Maximal width of elytra to length of elytra 0.64–0.69 (O: 0.66). Length of basi-metatarsus to metatibia 0.50–0.57 (O: 0.55). Male genitalia (Fig. 11). Median lobe broad, sub-parallel, curved apically; apex emarginated. Tectum wide and short, about 0.5 x as long as median lobe. Ventral surface well sclerotized, with a ridge from apex to apical 1 / 7. Endophallus with apical spiculae horn-like; with very long median spiculae; four pairs of long, curved, lateral spiculae, apex acute; ventral spiculae consisting of a pair of long spiculae, longitudinal row of short rounded sclerites, and cluster of slender, curved spiculae at middle. Female genitalia (Figs. 14–16). Spermathecal cornu (Fig. 14) slender, apically coiled; middle part long, apically and strongly curved, slightly widened near cornu; nodulus spherical, large; proximal spermathecal duct long, largest near nodulus, tightly coiled. Dorsal part of bursa-sclerites (Fig. 15) with teeth at outer margin of apical 1 / 3, extending into middle of dorsal surface, basally widened; ventral part (Fig. 16) slender, with teeth at inner margin. Color variation. Some individuals have a black-margined elytra and a black metasternum and metepisternum (Fig. 2). Type materials examined. The lectotype 3 of Monolepta gracilipes Chûjô, here designated to preserve stability and to make more universal use of this name, is labeled: “ Formosa. Musha (= Wushe, Nantou), 1919. V. 18 – VI. 15, T. Okuni / CoType / Monolepta gracilipes CHÛJÔ DET. M. Chujo / 1475 ” (TARI); paralectotypes: 33 (1472, 1474, 1480), 2 ƤƤ (1473, 1488), with same data as lectotype (TARI), one paralectotype with same data deposited at the DEI; 13 (1471), 1 Ƥ (2582): “ Formosa Arisan (= Alishan, Chiai), 1918. X. 2 – 23, J. Sonan (TARI); 13 (1481): “Ritozan (= Litungshan, Taoyuan), 31.VII. 1928, S. Issiki, 13 (1353), 1 Ƥ (1354): “ INOUE (= Chingchuan, Hsinchu), FORMOSA, 21.VII. 1935 COL. M. Chujo” (TARI); 1 Ƥ (2583): “ Formosa, Shinchiku (= Hsinchu), 1918. VII. 1–30, J. Sonan” (TARI); 1 Ƥ (1476): “Taiheizan (= Taipingshan, Ilan), 25.VIII. 1923, Col. T. Shiraki” (TARI); 1 Ƥ (1489): “Kobayashi (= Hiaolin, Kaohsiung), 25.VII. 1929, Formosa, Y. Miwa” (TARI); 13 (1478): “Shikiku (= Sunchitsun, Ilan), 3.XI. 2928, Col. J. Sonan” (TARI); 1 Ƥ (1479): “Nokosan (= Nengkaoshan, Nantou), 11.V. 1919, Okuni, Sonan” (TARI); 1 Ƥ (2584): “Nishimura (= Hsitsun, Miaoli), 24.VII. 1929, Formosa, Y. Miwa” (TARI); 1 Ƥ (1477): “Nankotaizan (= Nanhutashan, Taichung), 31 -X- 1928, Col. J. Sonan” (TARI). Specimens examined. 53, 5ƤƤ (# 6095-6104), Taipei, Lengshuikeng, 22.VI. 2008, leg. S.-F. Yu; 1 Ƥ (# 6436), Taipei, Sanchih, 30.VI. 2008, W.-T. Liu; 1 Ƥ (# 432), Taipei, Tienmu, 08.XII. 2006, leg. S.-F. Yu; 1 Ƥ (# 2322), Taoyuan, Tungyanshan, 08.VII. 2007, leg. S.-F. Yu; 1 Ƥ (# 6945), Taoyuan, Tamanshan, 03.VIII. 2008, leg. M.-H. Tsao; 13 (# 6142), Hsinchu, Lupi, 24.VI. 2008, leg. H. Lee; 23 (# 6733-6734), 2 ƤƤ (#6732, 6736), same locality, 19.VII. 2008, leg. M.-H. Tsao; 23, 1Ƥ, (# 6737-6739), same data but with 20.VII. 2008; 4 ƤƤ (# 6814 -6815, 6817- 6818), same data but with 26.VII. 2008; 1 Ƥ (# 2759), Pingtung, Tahanshan, 18.VII. 2007, leg. M.-H. Tsao; 13, same locality, 20.VII. 2007, leg. C.-F. Lee; 33, 7ƤƤ, Taitung, Tajen, 17.IX. 2008, leg. W.-C. Wang; 1 Ƥ (# 4508), Taitung, Taimali, 20.III. 2008, leg. P.-F. Wang. (all in TARI) Notes. The Taiwanese population was recorded firstly as Luperodes pallidulus Baly, 1874 by Chûjô (1935). It was transferred to the genus Monolepta Chevrolat by Chûjô (1938). At the meantime, he described those specimens with black-margined elytra as a new species: M. gracilipes Chûjô, 1938. Kimoto (1969) regarded these as a color variation of M. pallidula. Actually, the Taiwanese population is a distinct species with color variation which is different from the uniform M. pallidula (see below). Since M. gracilipes is an available name, it should be reinstated. Diagnosis. This species is close to Monolepta pallidula with the yellowish-brown body and similar endophallus, but differing by the wide and truncate apex of median lobe, relatively wider dorso-ventrally median lobe (Fig. 17), the narrowed anterior area of middle part of spermatheca (Fig. 19), the more prominent-toothed and narrow-based dorsal bursa sclerites (Fig. 20), and the ventral bursa sclerites with more and fine teeth. (Fig. 18). In addition, Monolepta gracilipes displays color variation on the elytra and metasternum in contrast with no color variation in M. pallidula. Distribution. Taiwan. This species seems to be the most common species of Monolepta in Taiwan based on lots of records and field experiences of the senior author.Published as part of Lee, Chi-Feng, 2009, A taxonomic revision of Monolepta pallidula species group in Taiwan (Coleoptera: Chrysomelidae: Galerucinae), pp. 15-27 in Zootaxa 2170 on pages 16-23, DOI: 10.5281/zenodo.18919
Monolepta gracilipes Chujo 1938
No full textMonolepta gracilipes Chûjô, 1938 reinstated Luperodes pallidulus Chûjô, 1935: 162. Monolepta pallidula Chûjô, 1938: 144; Chûjô, 1962: 123; Chûjô, 1963 a: 25; Chûjô, 1963 b: 393; Chûjô, 1965: 97; Kimoto, 1965: 488; Kimoto, 1966: 32; Kimoto, 1969: 47; Kimoto, 1986: 58; Kimoto, 1987: 189; Kimoto, 1989: 256; Kimoto, 1991: 14; Takizawa et al., 1995: 11. Monolepta gracilipes Chûjô, 1938: 145; Chûjô, 1962: 132; Chûjô, 1965: 97; Kimoto, 1969: 47 (synonymized with M. pallidula); Kimoto, 1986: 58. Description. Total length: males: 5.1–6.4 mm (O: 5.63 mm; n= 6); females: 4.9–6.2 mm (O: 5.73 mm; n= 6). General color yellowish-brown (Fig. 1); in some specimens antenna, apex of tibia, and tarsi dark brown. Antennomeres 4–10 about 6.2 times longer than broad at apex (Fig. 13); length of second to third antennomere 0.79 –1.00 (O: 0.87); length of third to fourth antennomere 0.36–0.50 (O: 0.59) (Fig. 12). Pronotal length to width 0.65–0.73 (O: 0.69). Maximal width of elytra to length of elytra 0.64–0.69 (O: 0.66). Length of basi-metatarsus to metatibia 0.50–0.57 (O: 0.55). Male genitalia (Fig. 11). Median lobe broad, sub-parallel, curved apically; apex emarginated. Tectum wide and short, about 0.5 x as long as median lobe. Ventral surface well sclerotized, with a ridge from apex to apical 1 / 7. Endophallus with apical spiculae horn-like; with very long median spiculae; four pairs of long, curved, lateral spiculae, apex acute; ventral spiculae consisting of a pair of long spiculae, longitudinal row of short rounded sclerites, and cluster of slender, curved spiculae at middle. Female genitalia (Figs. 14–16). Spermathecal cornu (Fig. 14) slender, apically coiled; middle part long, apically and strongly curved, slightly widened near cornu; nodulus spherical, large; proximal spermathecal duct long, largest near nodulus, tightly coiled. Dorsal part of bursa-sclerites (Fig. 15) with teeth at outer margin of apical 1 / 3, extending into middle of dorsal surface, basally widened; ventral part (Fig. 16) slender, with teeth at inner margin. Color variation. Some individuals have a black-margined elytra and a black metasternum and metepisternum (Fig. 2). Type materials examined. The lectotype 3 of Monolepta gracilipes Chûjô, here designated to preserve stability and to make more universal use of this name, is labeled: “ Formosa. Musha (= Wushe, Nantou), 1919. V. 18 – VI. 15, T. Okuni / CoType / Monolepta gracilipes CHÛJÔ DET. M. Chujo / 1475 ” (TARI); paralectotypes: 33 (1472, 1474, 1480), 2 ƤƤ (1473, 1488), with same data as lectotype (TARI), one paralectotype with same data deposited at the DEI; 13 (1471), 1 Ƥ (2582): “ Formosa Arisan (= Alishan, Chiai), 1918. X. 2 – 23, J. Sonan (TARI); 13 (1481): “Ritozan (= Litungshan, Taoyuan), 31.VII. 1928, S. Issiki, 13 (1353), 1 Ƥ (1354): “ INOUE (= Chingchuan, Hsinchu), FORMOSA, 21.VII. 1935 COL. M. Chujo” (TARI); 1 Ƥ (2583): “ Formosa, Shinchiku (= Hsinchu), 1918. VII. 1–30, J. Sonan” (TARI); 1 Ƥ (1476): “Taiheizan (= Taipingshan, Ilan), 25.VIII. 1923, Col. T. Shiraki” (TARI); 1 Ƥ (1489): “Kobayashi (= Hiaolin, Kaohsiung), 25.VII. 1929, Formosa, Y. Miwa” (TARI); 13 (1478): “Shikiku (= Sunchitsun, Ilan), 3.XI. 2928, Col. J. Sonan” (TARI); 1 Ƥ (1479): “Nokosan (= Nengkaoshan, Nantou), 11.V. 1919, Okuni, Sonan” (TARI); 1 Ƥ (2584): “Nishimura (= Hsitsun, Miaoli), 24.VII. 1929, Formosa, Y. Miwa” (TARI); 1 Ƥ (1477): “Nankotaizan (= Nanhutashan, Taichung), 31 -X- 1928, Col. J. Sonan” (TARI). Specimens examined. 53, 5ƤƤ (# 6095-6104), Taipei, Lengshuikeng, 22.VI. 2008, leg. S.-F. Yu; 1 Ƥ (# 6436), Taipei, Sanchih, 30.VI. 2008, W.-T. Liu; 1 Ƥ (# 432), Taipei, Tienmu, 08.XII. 2006, leg. S.-F. Yu; 1 Ƥ (# 2322), Taoyuan, Tungyanshan, 08.VII. 2007, leg. S.-F. Yu; 1 Ƥ (# 6945), Taoyuan, Tamanshan, 03.VIII. 2008, leg. M.-H. Tsao; 13 (# 6142), Hsinchu, Lupi, 24.VI. 2008, leg. H. Lee; 23 (# 6733-6734), 2 ƤƤ (#6732, 6736), same locality, 19.VII. 2008, leg. M.-H. Tsao; 23, 1Ƥ, (# 6737-6739), same data but with 20.VII. 2008; 4 ƤƤ (# 6814 -6815, 6817- 6818), same data but with 26.VII. 2008; 1 Ƥ (# 2759), Pingtung, Tahanshan, 18.VII. 2007, leg. M.-H. Tsao; 13, same locality, 20.VII. 2007, leg. C.-F. Lee; 33, 7ƤƤ, Taitung, Tajen, 17.IX. 2008, leg. W.-C. Wang; 1 Ƥ (# 4508), Taitung, Taimali, 20.III. 2008, leg. P.-F. Wang. (all in TARI) Notes. The Taiwanese population was recorded firstly as Luperodes pallidulus Baly, 1874 by Chûjô (1935). It was transferred to the genus Monolepta Chevrolat by Chûjô (1938). At the meantime, he described those specimens with black-margined elytra as a new species: M. gracilipes Chûjô, 1938. Kimoto (1969) regarded these as a color variation of M. pallidula. Actually, the Taiwanese population is a distinct species with color variation which is different from the uniform M. pallidula (see below). Since M. gracilipes is an available name, it should be reinstated. Diagnosis. This species is close to Monolepta pallidula with the yellowish-brown body and similar endophallus, but differing by the wide and truncate apex of median lobe, relatively wider dorso-ventrally median lobe (Fig. 17), the narrowed anterior area of middle part of spermatheca (Fig. 19), the more prominent-toothed and narrow-based dorsal bursa sclerites (Fig. 20), and the ventral bursa sclerites with more and fine teeth. (Fig. 18). In addition, Monolepta gracilipes displays color variation on the elytra and metasternum in contrast with no color variation in M. pallidula. Distribution. Taiwan. This species seems to be the most common species of Monolepta in Taiwan based on lots of records and field experiences of the senior author.Published as part of Lee, Chi-Feng, 2009, A taxonomic revision of Monolepta pallidula species group in Taiwan (Coleoptera: Chrysomelidae: Galerucinae), pp. 15-27 in Zootaxa 2170 on pages 16-23, DOI: 10.5281/zenodo.18919
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