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Nemastoma bidentatum subsp. gruberi Novak, Slana Novak, Kozel & Raspotnig 2021, ssp. nov.
No full textNemastoma bidentatum gruberi Novak, Slana Novak, Kozel & Raspotnig ssp. nov. urn:lsid:zoobank.org:act: 17EA28A1-5F36-4DFC-9191-9C8680496FAC Figs 1A, C, 2– 3, 4E, 5E, 6E, 7E, 8E, 9E, 10E, 11E, 12E, 13D; Table 8 Nemastoma bidentatum sparsum – Novak et al. 2002: 136 [partim: Brežice, Črnomelj, Kočevje, Krško, Litija, Metlika, Novo mesto, Ribnica, Sevnica,Trebnje]. — Novak 2004b: 245 [partim: Buje, Karojba, Vižinada]; 2005b: 105. Diagnosis Relatively long-legged subspecies of Nemastoma bidentatum, with strait Ch ventral margin, and Pa-Fe with either 4 subequidistantly positioned spines or pointed tubercles or long setae on low protrusions, or their combination in the distal Pa-Fe half, or alternatively, with 2–3 distal stout thorns. Rec sem with one tubular and 12 elongated balloon-like vesicles. Etymology The subspecies name ʻ gruberi ʼ is dedicated to Jürgen Gruber (Vienna), our teacher, colleague and friend, who generously introduced us to the taxonomy of harvestmen and helped in identifying problematic specimens. Material examined Holotype SLOVENIA – WL18 • 1 ♂; Dolenje Laknice, Mokronog; 45.93° N, 15.21° E; 237 m a.s.l.; 26 Oct. 2013; L. Slana Novak and T. Novak leg.; mixed forest litter sift; PMSL-Opiliones-GR 4661. Paratypes SLOVENIA – WL18 • 1 ♀; same collection data as for holotype; PMSL-Opiliones-GR 4664 • 5 ♂♂, 4 ♀♀; same collection data as for holotype; PMSL-Opiliones-GR4662, GR4666, GR4667, GR4669, GR4673, GR4676 to GR4679. Other material ITALY – VL05 • 1 ♀; Chiusa/Ricmanje; 29 Sep. 1999; L. Slana Novak and T. Novak leg. (275/1999, rev. 2015); PMSL. – VL15 • 1 ♀; Basovizza/Bazovica; 29 Sep. 1999; L. Slana Novak and T. Novak leg. (271/1999, rev. 2015); PMSL. CROATIA – UL92 • 3 ♂♂, 2 ♀♀ (paratypes of N. b. sparsum); vicinity of Buje, and VL02, Vižinada; Sep. 1963; R. Sturany leg.; NHMW 4700. – VL01 • 1 ♂; Karojba; 28 Feb. 1990; F. Potočnik leg. (1307/1998, rev. 2018); PMSL. – VL13 • 1 ♀; Mlini; 30 Sep. 1990; L. Slana Novak and T. Novak leg. (32/1990, rev. 2015); PMSL. – XL90 • 1 ♂, 2 ♀♀; Ponor Sušik, Drežnik; 8 Sep. 2009; A. Schönhofer leg. (Coll. ASc 334, ASc det., TN rev. 5/2012); PMSL. SLOVENIA – UL93 • 4 ♀♀, 1 juv.; Abrami; 19 Jul. 1999; L. Slana Novak and T. Novak leg. (148/1999, rev. 2018); PMSL • 2 ♀♀; ibid.; 20 Jul. 1999; L. Slana Novak and T. Novak leg. (188/1999, rev. 2015); PMSL • 1 ♀; Dragonja; 14 Sep. 2000; L. Slana Novak and T. Novak leg. (154/2001, rev. 2015); PMSL • 4 ♂♂, 4 ♀♀; Dragonja valley; 26 Jun. 2015; T. Novak leg. (78/2015 = GR 5123−5130); PMSL • 2 ♂♂, 3 ♀♀; Ivankovec; 23 Sep. 1998; L. Slana Novak and T. Novak leg. (1791/1998, rev. 2018); PMSL • 1 ♀; Lucija–Seča – Debernardo; 22 Sep. 1998; L. Slana Novak and T. Novak leg. (1817/1998, rev. 2019); PMSL • 1 ♂; Korte; 23 Sep. 1998; L. Slana Novak and T. Novak leg. (1821/1998, rev. 2018); PMSL • 33 ♂♂, 30 ♀♀; Medljan, Korte; 21 Aug. 1999; T. Novak leg. (239/1999, rev. 2018); PMSL • 1 juv.; ibid.; 22 Jun. 2000; B. Bertoncelj, B. Ogrizek and T. Novak leg. (217/2000, rev. 2018); PMSL • 1 ♀; ibid.; (218/2000, TN rev. 2019); PMSL • 1 ♂; Padna; 19 Jul. 1999; L. Slana Novak and T. Novak leg. (168/1999, rev. 2017); PMSL • 2 ♀♀; Parecag; 23 Sep. 1998; L. Slana Novak and T. Novak leg. (1788/1998, rev. 2018); PMSL • 1 ♀; Seča; 10 Oct. 1998; L. Slana Novak and T. Novak leg. (1844/1998, rev. 2008); PMSL • 1 ♂, 1 ♀; ibid.; 24 Oct. 1998, L. Slana Novak and T. Novak leg. (1881/1998, rev. 2007); PMSL • 3 ♀♀; ibid.; 21 Aug. 1999; M. Štangelj and T. Novak leg. (236/1999, rev. 2008); PMSL • 15 ♂♂, 21 ♀♀; ibid.; 21 Aug. 1999; M. Štangelj and T. Novak leg. (231/1999, rev. 2015); PMSL • 1 ♂, 6 ♀♀; Sečovlje; 1 Mar. 2009; L. Slana Novak and T. Novak leg. (1/2015); PMSL • 1 ♂; Sv. Peter; 11 Oct. 1998; L. Slana Novak and T. Novak leg. (1850/1998, rev. 2015); PMSL • 4 ♂♂, 2 ♀♀; Sv. Sab, Padna; 19 Jul. 1999; L. Slana Novak and T. Novak leg. (151/1999, rev. 2009); PMSL • 1 ♀; Škrline, Dragonja; 20 Jul. 1999; L. Slana Novak and T. Novak leg. (194/1999, rev. 2015); PMSL. – VL03 • 2 ♂♂, 1 ♀; Boršt; 14 Apr. 2011; T. Novak leg. (76/2011, rev. 2015); PMSL • 2 ♀♀; ibid.; (81/2011, rev. 2015); PMSL. – VL04 • 1 ♀; Ankaran; 29 Sep. 1999; L. Slana Novak and T. Novak leg. (276/1999, rev. 2015); PMSL • 1 ♂, 1 ♀; Kolombini, Spodnje Škofije; 26 Sep. 2007; L. Slana Novak and T. Novak leg. (94/2007, rev. 2018); PMSL • 5 ♂♂; Mlinarji – Osp; 26 Sep. 2007; L. Slana Novak and T. Novak leg. (97/2007, rev. 2017); PMSL • 1 ♂, 1 ♀; Rižana; 24 Sep. 2016; L. Slana Novak and T. Novak leg. (18/2018); PMSL • 1 ♀; Škocjanski zatok, Koper; 15 Oct. 2009; S. Polak leg. (PK 1/2015); NM • 4 ♂♂, 3 ♀♀; Zgornje Škofije – Tinjan; 26 Sep. 2007; L. Slana Novak and T. Novak leg. (88/2007); PMSL. – VL07 • 4 ♀♀; Hrastje, Šmarje; 17 Aug. 2000 (674/2000, rev. 2015); PMSL. – VL13 • 1 ♂; Poletiči, Kubed; 4 Sep. 2016; T. Marinko leg. (27/2016); PMSL • 1 ♂, 1 ♀; Sočerga; 28 Sep. 1990; L. Slana Novak and T. Novak leg. (31/1990, rev. 2015); PMSL. – VL14 • 4 ♂♂, 4 ♀♀; Bezovica; 29 Sep. 1990; L. Slana Novak and T. Novak leg. (LSN 1/1990, TN rev. 2018); PMSL • 1 ♂; Hrpelje, Kozina; 2 Apr. 1993; S. Brelih leg. (651/1998, rev. 2018); PMSL • 2 ♂♂; Predloka; 29 Sep. 1990; L. Slana Novak and T. Novak leg. (35/1990, rev. 2015); PMSL • 2 ♂♂, 1 ♀; Rižana; 29 Sep. 1990; L. Slana Novak and T. Novak leg. (33/1990, rev. 2015); PMSL. – VL15 • 1 ♀; Botač; 22 Mar. 1990; M. Černila, B. Horvat and I. Sivec leg. (1246/1997); PMSL. – VL16 • 1 ♂; Divača; 2 Apr. 1993; S. Brelih leg. (174/2001, rev. 2015); PMSL. – VL17 • 1 ♂; Vrabče, Podnanos; 10 Mar. 1994; S. Brelih leg. (632/1998, rev. 2017); PMSL. – VL18 • 1 ♂; Uhanje; 3 Sep. 2001; T. Novak leg. (329/2001, rev. 2018); PMSL. – VL23 • 1 ♂, 1 ♀; Poljane pri Podgradu; 18 Aug. 2011; L. Slana Novak and T. Novak leg. (166/2012, rev. 2019); PMSL. – VL25 • 3 ♀♀; Buje; 10 Mar. 1994; S. Brelih leg. (163/2001, rev. 2018); PMSL • 1 ♂, 2 ♀♀; Goriče pri Framljah; L. Slana Novak and T. Novak leg. (343/2011); PMSL • 1 ♀; ibid.; (344/2011); PMSL. – VL28 • 1 ♂, 1 ♀; Podvrh, Raka; 18 Jul. 2012; P. Kozel, T. Kozel and T. Novak leg. (55/2012); PMSL • 9 ♂♂, 7 ♀♀; ibid.; (GR 3734–3746); PMSL • 3 ♀♀; ibid.; (GR 3791); PMSL. – VL29 • 6 ♂♂, 12 ♀♀; Dolnje Brezovo, Sevnica; 30 Jun. 2012; (GR 3670–3683); PMSL. – VL36 • 1 ♂, 1 ♀; Palčje – Mt. Javorniki; 1 Oct. 2011; L. Slana Novak and T. Novak leg. (361/2011, rev. 2019); PMSL. – VL45 • 1 ♀; Koritnice − Mašun; 2 Oct. 2011; L. Slana Novak and T. Novak leg. (329/2011, rev. 2014); PMSL. – VL47 • 1 ♀; Suha reber, Koritnice; 21 Jan. 2007; S. Polak leg. (107/2007); NM. – VL54 • 1 ♀; Sviščaki; 20 May 1989; T. Trilar leg. (271/2000, rev. 2017); PMSL • 1 ♀; ibid.; 25 May 1989; T. Trilar leg. (275/2000, rev. 2017); PMSL • 1 ♀; ibid.; 21 Jul. 2001; B. Bertoncelj, U. Bertoncelj and T. Novak leg. (29/2001, rev. 2018); PMSL • 1 ♀; ibid.; (34/2001, rev. 2011); PMSL • 1 ♂, 3 ♀♀; Travni Dolci, Mt. Snežnik; 12 Aug. 2001; L. Slana Novak and T. Novak leg. (195a/2001, rev. 2020); PMSL • 2 ♂♂, 4 ♀♀; Mt. Veliki Snežnik; 20 Jul. 1999; L. Slana Novak and T. Novak leg. (185a/1999, rev. 2020); PMSL • 1 ♂; ibid.; 12 Sep. 2018; L. Slana Novak and T. Novak leg. (116/2018); PMSL. – VL36 • 1 ♂, 1 ♀; Palčje − Javorniki; L. Slana Novak and T. Novak leg. (361/2011, rev. 2015); PMSL. – VL66 • 1 ♀; Benete, Runarsko; 31 May 2004; L. Slana Novak and T. Novak leg. (33/2004, rev. 2015); PMSL. – VL67 • 5 ♂♂, 3 ♀♀; Nova vas – Volčje; 26 Sep. 2006; L. Slana Novak and T. Novak leg. (25/2006, rev. 2019); PMSL • 1 ♂; Runarsko; 26 Sep. 2006; L. Slana Novak and T. Novak leg. (38/2006, rev. 2015); PMSL • 1 ♂, 1 ♀; Veliki vrh, Runarsko; 26 Sep. 2006; L. Slana Novak and T. Novak leg. (36/2006, rev. 2015); PMSL • 7 ♂♂, 5 ♀♀; Volčje; 26 Sep. 2006; L. Slana Novak and T. Novak leg. (28/2006, rev. 2015); PMSL. – VL69 • 1 ♂; Veliki vrh pri Šmarju; 10 May 2008; L. Slana Novak and T. Novak leg. (156/2011); PMSL. – VL74 • 1 ♂, 1 ♀; Mt. Goteniški Snežnik; 28 May 2000; L. Slana Novak and T. Novak leg. (247/2000, rev. 2007); PMSL • 1 ♀; Strma reber, Zgornji Čačiči; 27 Sep. 1996; L. Slana Novak and T. Novak leg. (LSN 126/1996, TN rev. 2008); PMSL. – VL75 • 2 ♂♂; Draga; 17 Aug. 2011; L. Slana Novak and T. Novak leg. (94/2011); PMSL • 1 ♂, 2 ♀♀; Mt. Goteniški Snežnik; 28 May 2000; L. Slana Novak and T. Novak leg. (253/2000, rev. 2007); PMSL. – VL76 • 1 ♂; Bukovica, Ribnica; Jul.1994; B. Oven leg. (LSN 25/1996, TN rev. 2009); PMSL • 2 ♂♂, 1 ♀; Lipovšica, Sodražica; 22 Aug. 2011; L. Slana Novak and T. Novak leg. (100/2011, rev. 2018); PMSL • 1 ♂; Nova Štifta; 22 Aug. 2011; L. Slana Novak and T. Novak leg. (105/2011); PMSL • 2 ♂♂, 3 ♀♀; Ribnica; 6 Sep. 2000; M. Mlakar leg. (828/2000, rev. 2008); PMSL • 1 ♂, 2 ♀♀; ibid.; (831/2000, rev. 2019); PMSL • 1 ♀; Mt. Turn; 27 Jun. 1980; B. Drovenik leg. (383/2002, rev. 2018); PMSL • 1 ♂; ibid.; 28 Aug. 1980; B. Drovenik leg. (394/2002, rev. 2009); PMSL. – VL77 • 1 ♂; Dolnje Retje; 3 May 2008; L. Slana Novak and T. Novak leg. (161/2011); PMSL • 1 ♀; Rašica, Velike Lašče; 16 Jun. 1986 (261/2002, rev. 2015); PMSL • 1 ♂, 1 ♀; Velike Poljane, Ortnek; 3 May 2008; L. Slana Novak and T. Novak leg. (142/2011, rev. 2019); PMSL. – VL78 • 1 ♂; Laporje; 3 May 2008; L. Slana Novak and T. Novak leg. (151/2011); PMSL • 1 ♀; Zagradec pri Grosupljem; 7 Jul. 1986; L. Slana Novak and T. Novak leg. (500/2002, rev. 2015); PMSL. – VL79 • 1 ♀; Veliki Lipoglav − Dobrinje; 8 Jul. 1986; L. Slana Novak and T. Novak leg. (467/2002, rev. 2015); PMSL. – VL83 • 2 ♂♂; Vršič, Banjaloka; 24 Apr. 2001; S. Brelih leg. (121/2001, rev. 2019); PMSL. – VL84 • 1 ♂; Jelenja vas; 3 May 2008; L. Slana Novak and T. Novak leg. (137a/2011); PMSL • 1 ♂; ibid.; (153/2011); PMSL • 1 ♂; Pleš, Borovec pri Kočevski Reki; 19 Sep. 2013; P. Kozel and T. Novak leg. (71/2013, rev. 2018); PMSL. – VL85 • 1 ♀; Grčarice; 18 Oct. 2013; T. Novak leg. (63/2013, rev. 2015); PMSL • 1 ♂, 2 ♀♀; Grčarske Ravne; 18 Oct. 2013; T. Novak leg. (66/2013, TN rev. 2019); PMSL • 3 ♀♀; Kočevje; 6 Sep. 2000; M. Mlakar leg. (822/2000, rev. 2008); PMSL • 1 ♂, 1 ♀; ibid.; (825/2000, rev. 2007); PMSL • 2 ♂♂, 1 ♀; Mt. Stojna, Kočevje; 25 Oct. 1996; S. Brelih leg. (92/2001, rev. 2008); PMSL. – VL86 • 1 ♀; Dolenja vas; 3 May 2008; L. Slana Novak and T. Novak leg. (110/2011); PMSL • 1 ♂, 1 ♀; ibid.; (154/2012, rev. 2018); PMSL. – VL87 • 1 ♂; Breg pri Zagradcu; 4 Nov. 1993; S. Brelih leg. (805/1998, rev. 2007); PMSL. – VL89 • 1 ♀; Potok, Stična; 4 Mar. 1994; S. Brelih leg. (658/1998, rev. 2015); PMSL. – VL93 • 1 ♂; Brsnik, Fara; 3 May 2008; L. Slana Novak and T. Novak leg. (157/2011, rev. 2018); PMSL. – VL94 • 1 ♂, 1 ♀; Dolenja Žaga; 29 Apr. 2001; S. Brelih leg. (131/2001, rev. 2015); PMSL • 1 ♀; Dolenja Žaga – Podstene; 26 Jun. 1999; L. Slana Novak and T. Novak leg. (90/1999, rev. 2008); PMSL • 1 ♂, 1 ♀; Gorenja Žaga; 3 May 2008; L. Slana Novak and T. Novak leg. (111a/2011); PMSL • 1 ♂, 1 ♀; ibid.; (159/2011); PMSL • 1 ♀; Ograja; 3 May 2008; L. Slana Novak and T. Novak leg. (109a/2011); PMSL • 1 ♀; ibid.; (121a/2011); PMSL • 1 ♀; ibid.; (144/2011); PMSL • 1 ♀; ibid.; (148/2011); PMSL • 1 ♂, 1 ♀; Ograja – Suhor − Morava; 3 May 2008; L. Slana Novak and T. Novak leg. (120a/2011); PMSL • 1 ♂, 1 ♀; ibid.; (138a/2011); PMSL • 1 ♂, 1 ♀; ibid.; (152/2011); PMSL • 1 ♂; Ograja – Zdihovo; 26 Jun. 1999; L. Slana Novak and T. Novak leg. (76/1999, rev. 2008); PMSL • 3 juvs; ibid.; 12 Jun. 1999; L. Slana Novak and T. Novak leg. (32/1999, rev. 2008); PMSL • 2 juvs; Podstene pri Kostelu; 12 Jun. 1999; L. Slana Novak and T. Novak leg. (24/1999, rev. 2008); PMSL • 1 ♂; Rajndol; 21 Mar. 1994; S. Brelih leg. (821/1998, rev. 2008); PMSL • 5 ♂♂, 3 ♀♀; Srednji Potok, Banja Loka; 8 Oct. 2012; P. Kozel and T. Novak leg. (68/2015); PMSL • 1 ♂, 1 ♀, 2 juvs; Štalcerski preval; 12 Jun. 1999; L. Slana Novak and T. Novak leg. (33/1999, rev. 2008); PMSL. – VL95 • 1 ♂, 2 ♀♀; Dolga vas; 17 Aug. 1985; T. Novak, M. Slana Novak and L. Slana Novak leg. (LSN 97/1986, TN rev. 2007); PMSL • 1 ♂, 1 ♀; Livold; 3 May 2008; L. Slana Novak and T. Novak leg. (123a/2011); PMSL • 1 ♂, 1 ♀; ibid.; (158/2011); PMSL. – VL97 • 1 ♀; Dvor, Lašče; 10 May 2008; L. Slana Novak and T. Novak leg. (112/2011); PMSL • 1 ♂, 2 ♀♀; Železno, Dobrnič; 4 Nov. 1993; S. Brelih leg. (812/1998, rev. 2007); PMSL. – VL99 • 3 ♀♀; Nova gora, Čatež; 15 Aug. 2007; L. Slana Novak and T. Novak leg. (61/2007, rev. 2015); PMSL. – VM80 • 1 ♂; Litija; 10 Sep. 2000; M. Mlakar leg. (809/2000, rev. 2008); PMSL • 1 ♂, 1 ♀; ibid.; (834/2000, rev. 2015); PMSL. – WL03 • 2 ♂♂; Dolenje Kozice; 18 Apr.−13 May 2002; B. Drovenik and A. Gergeli leg. (509/2018); PMSL • 1 ♂; ibid.; 18 Apr.−13 May 2002; B. Drovenik and A. Gergeli leg. (550/2018); PMSL • 1 ♂; ibid.; 5 Jul. 2002; B. Drovenik and A. Pirnat leg. (1135/2018); PMSL • 1 ♀; ibid.; 6 Aug. 2002; B. Drovenik and A. Pirnat leg. (1090/2018); PMSL • 1 ♀; ibid.; 30 Sep. –28 Oct. 2002; B. Drovenik and A. Gergeli leg. (PK 64/2012); PMSL • 1 ♀; ibid.; 25 Nov. 2002; B. Drovenik and A. Pirnat leg. (270/2018); PMSL • 2 ♀♀; ibid.; 25 Nov. −23 Dec. 2002; B. Drovenik and A. Gergeli leg. (1027/2018); PMSL • 4 ♂♂, 5 ♀♀; Srednji Radenci; 17 Aug. 1985; L. Slana Novak, M. Slana Novak and T. Novak leg. (LSN 51/1986, rev. 2008); PMSL • 3 ♂♂, 4 ♀♀; Stari trg ob Kolpi; 21 Mar. 1994; S. Brelih leg. (851/1998, rev. 2008); PMSL • 5 ♂♂, 5 ♀♀; Dolenji Radenci – Breg pri Sinjem vrhu; 3 May 2008; L. Slana Novak and T. Novak leg. (135a/2011); PMSL • 5 ♂♂, 2 ♀♀; ibid.; (149/2011); PMSL. – WL05 • 1 ♂, 3 ♀♀, 1 juv.; Črmošnjice; 16 Aug. 1985; L. Slana Novak, M. Slana Novak and T. Novak leg. (160/1997); PMSL • 1 ♀; Koprivnik, Kočevski Rog; 21 Mar. 1994; S. Brelih leg. (867/1998, rev. 2008); PMSL • 1 ♂, 2 ♀♀; Mt. Mirna gora; 8 Oct. 2012; P. Kozel and T. Novak leg. (59/2015); PMSL • 4 ♂♂, 4 ♀♀; ibid.; 8 Oct. 2012; P. Kozel and T. Novak leg. (66/2015); PMSL. – WL06 • 1 ♂; Cerovec − Selišče, Podturn pri Dolenjskih toplicah; 10 May 2008; L. Slana Novak and T. Novak leg. (145/2011, rev. 2017); PMSL. – WL07 • 1 ♂; Boršt pri Dvoru; 10 May 2008; L. Slana Novak and T. Novak leg. (113/2012); PMSL • 1 ♀; Prečna, Mirna; Jun. 2000; V. Jaćimović leg. (PK and TN 100/2020); PMSL. – WL08 • 1 ♂; Rdeči kal, Dobrnič; 4 Nov. 1993; S. Brelih leg. (872/1998, rev. 2007); PMSL • 1 ♂, 1 ♀; Trebnje; 29 Aug. 2000; M. Mlakar leg. (787/2000, rev. 2018); PMSL • 1 ♂; ibid.; (787a/2000, rev. 2018); PMSL • 1 ♀; ibid.; (788/2000, rev. 2008); PMSL. – WL13 • 4 ♀♀; Balkovci, Vinica; 15 Aug. 1999; M. Štangelj and T. Novak leg. (212/1999, rev. 2008); PMSL • 2 ♂♂, 2 ♀♀; Dolenji Radenci – Breg pri Sinjem vrhu; 3 May 2008; L. Slana Novak and T. Novak leg. (146/2011); PMSL • 4 ♂♂, 1 ♀; Kot pri Dramlju; 16 Aug. 1999; M. Štangelj and T. Novak leg. (207/1999, rev. 2015); PMSL • 3 ♂♂, 2 ♀♀; Špeharji, Vinica; 16 Aug. 1999; M. Štangelj and T. Novak leg. (201/1999, rev. 2006); PMSL • 6 ♂♂, 5 ♀♀; Vinica – Sinji vrh; 17 Aug. 1985; L. Slana Novak, M. Slana Novak and T. Novak leg. (LSN 59/1986, TN rev. 2007); PMSL • 2 ♂♂; Vukovci; 18 Apr. 1996; S. Brelih leg. (764/1998, rev. 2007); PMSL. – WL14 • 1 ♂, 1 ♀; Bistrica, Črnomelj; 26 Jun. 1999; L. Slana Novak and T. Novak leg. (65/1999, rev. 2017); PMSL • 1 ♀; Črnomelj; 15 Aug. 1999; M. Štangelj and T. Novak leg. (221/1999, rev. 2008); PMSL • 3 ♂♂, 3 ♀♀, 2 juvs; Desinec – Črnomelj; 17 Aug. 1985; L. Slana Novak, M. Slana Novak and T. Novak leg. (385/1998, rev. 2007); PMSL • 2 ♂♂, 2 ♀♀; Tanča gora; 21 Mar. 1994; S. Brelih leg. (861/1998, rev. 2015); PMSL • 5 ♂♂, 2 ♀♀; Sinji vrh, Vinica; 16 Aug. 1999; M. Štangelj and T. Novak leg. (205/1999, rev. 2007); PMSL. – WL15 • 1 ♀; Krupa spring; 10 Mar. 1994; S. Brelih leg. (808/1998, rev. 2007); PMSL • 6 ♂♂, 3 ♀♀, 3 juvs; Lokve, Črnomelj; 2 Jul. 1995; L. Slana Novak and T. Novak leg. (105/1997, rev. 2009); PMSL • 3 ♂♂, 2 ♀♀; Mihelja vas; 18 Aug. 1985; L. Slana Novak, M. Slana Novak and T. Novak leg. (LSN 55/1986, TN rev. 2017); PMSL • 1 ♀; Praprot; 6 Aug. 1997; T. Novak leg. (418/1998, rev. 2008); PMSL • 1 ♀; ibid.; (432/1998, rev. 2008); PMSL • 1 ♀; ibid.; 22 Jul. 1999; L. Slana Novak and T. Novak leg. (37/1999, rev. 2008); PMSL • 1 ♀; ibid.; (110/1999, rev. 2008); PMSL • 2 ♂♂, 2 ♀♀; Stranska vas; 22 Jul. 1999; L. Slana Novak and T. Novak leg. (108/1999, rev. 2007); PMSL • 2 ♂♂, 1 ♀; Vrčice; 10 May 2008; L. Slana Novak and T. Novak leg. (110/2012); PMSL • 1 ♂, 3 ♀♀; Vrčice – Semič; 8 Oct. 2012; P. Kozel and T. Novak leg. (64/2015); PMSL. – WL16 • 2 ♀♀; Laze, Uršna Sela; 10 May 2008; L. Slana Novak and T. Novak leg. (103/2012, rev. 2015); PMSL • 1 ♀; ibid.; (108/2012, rev. 2018); PMSL. – WL17 • 1 ♂, 1 ♀; Brod, Novo Mesto; 31 Aug. 2000; M. Mlakar leg. (811/2000, rev. 2007); PMSL • 1 ♂; Novo mesto; Karaman leg. (87/1983 microscopic preparation, JH det., TN rev. 2011); PMSL • 1 ♂, 1 ♀; ibid.; Karaman leg. (108/1983 microscopic preparation, JH det., TN rev. 2011); PMSL • 1 ♀; ibid.; (814/2000); PMSL • 1 ♂; ibid.; (153/2018 microscopic preparation, JH det., TN rev. 2018); PMSL • 1 ♀, 1 ♀; ibid.; 31 Aug. 2000; M. Mlakar leg. (811/2000, rev. 2007); PMSL • 1 ♂, 2 ♀♀; Ratež; 29 Apr. 1995; S. Brelih leg. (750/1998, rev. 2007); PMSL. – WL18 • 6 ♂♂, 5 ♀♀; Dolenje Laknice; 26 Oct. 2013; T. Novak leg. (GR 4661, 4662, 4664, 4666, 4667, 4669, 4673, 4676, 4677−4679); PMSL. – WL19 • 1 ♀; Boštanj – Mokronog – Križ; L. Slana Novak and T. Novak leg. (70/2007, rev. 2011); PMSL • 1 ♂; Jelovec; 15 Aug. 2007; L. Slana Novak and T. Novak leg. (56/2007); PMSL. – WL23 • 1 ♂; Zilje; 18 Apr. 1996; S. Brelih leg. (767/1998, rev. 2007); PMSL • 1 ♂, 2 ♀♀; Zilje – Vinica; 17 Aug. 1985; L. Slana Novak, M. Slana Novak and T. Novak leg. (364/1998, rev. 2007); PMSL. – WL24 • 5 ♂♂, 1 ♀, 1 juv.; Gorenjci pri Adlešičih; 15 Aug. 1999; M. Štangelj and T. Novak leg. (218/1999, rev. 2007); PMSL • 1 ♂; Dragoši, Griblje; 18 Apr. 1994; S. Brelih leg. (733/1998, rev. 2007); PMSL • 4 ♂♂, 2 ♀♀; Griblje − Desinec; 16 Aug. 1985; L. Slana Novak, M. Slana Novak and T. Novak leg. (161/1997, rev. 2009); PMSL. – WL25 • 1 ♂; Bečka jama cave, Cad. No. 6275, Boldraž; 9 Jun. 2004; V. Germovšek, T. Tkavc, A. Kocuvan and T. Novak leg. (21/2004, rev. 2007); PMSL • 1 ♂; Božakovo; 3 May 2004; L. Slana Novak and T. Novak leg. (48/2004, rev. 2007); PMSL • 2 ♂♂, 3 ♀♀; Bušinja vas; 18 Apr. 1996; S. Brelih leg. (743/1998, rev. 2007); PMSL • 1 ♂, 4 ♀♀; Kapljišče; 16 Aug. 1985; L. Slana Novak, M. Slana Novak and T. Novak leg. (LSN 113/1986, TN rev. 2007); PMSL • 2 ♂♂, 1 juv.; ibid. (LSN 119/1986, TN rev. 2007); PMSL • 3 ♀♀; Metlika; 31 Aug. 2000; M. Mlakar leg. (816/2000, rev. 2007); PMSL • 1 ♂; ibid.; (818/2000, rev. 2015); PMSL • 7 ♂♂, 9 ♀♀; Trnovec, Metlika; 23 Jul. 1999; L. Slana Novak and T. Novak leg. (141/1999, rev. 2008); PMSL • 1 ♂, 2 ♀♀; Sv. Urban, Metlika; 18 Apr. 1996; S. Brelih leg. (784/1998, rev. 2007); PMSL. – WL26 • 1 ♂; Krašnji vrh; 9 Jun. 2004; V. Germovšek, T. Tkavc, A. Kocuvan and T. Novak leg. (37/2004, rev. 2007); PMSL • 1 ♂; Markučev Mlin, Krašnji vrh; 1 Jul. 2007; L. Slana Novak and T. Novak leg. (21/2007); PMSL • 4 &fema
Nemastoma bidentatum subsp. martensi Novak, Slana Novak & Raspotnig 2021, ssp. nov.
No full text<i>Nemastoma bidentatum martensi</i> Novak, Slana Novak & Raspotnig ssp. nov. <p>urn:lsid:zoobank.org:act: CCDC54EF-D41A-4547-A4D4-50370212C243</p> <p>Figs 2–3, 4G, 5G, 6G, 7G, 8G, 9G, 10G, 11G, 12G, 13F; Table 10</p> Diagnosis <p> Subspecies of <i>Nemastoma bidentatum</i> with Ch basal article with a saw-like series of 1–3 μm high denticles on anterior margin of ventral hump, and a row of 5–11 denticles and tubercles in the distal half of Pa-Fe. Rec sem of 12–14 slightly elongated balloon-like vesicles.</p> Etymology <p> The subspecies name ʻ <i>martensi</i> ʼ is dedicated to Jochen Martens (Mainz), our teacher, colleague and friend, who provided the first modern revision of harvestmen in Slovenia.</p> Material examined <p> <b>Holotype</b> SLOVENIA – <b>VL23</b> • 1 ♂; Poljane pri Podgradu; 45.50° N, 14.10° E; 597 m a.s.l.; 25 Sep. 2011; L. Slana Novak and T. Novak leg.; thermophile scrub and mixed forest litter sift; PMSL-Opiliones-TN 287/2011. <b>Paratypes</b> SLOVENIA – <b>VL23</b> • 8 ♂♂, 5 ♀♀; same collection data as for holotype; PMSL-Opiliones-TN 287/2011.</p> <p> <b>Other material</b></p> <p> SLOVENIA – <b>VL15</b> • 3 ♂♂, 8 ♀♀; Buje; 27 Sep. 2011; L. Slana Novak and T. Novak leg. (316/2011, rev. 2015); PMSL • 2 ♂♂; ibid.; (316a/2011); PMSL • 1 ♀; Mt. Ostrič; 29 Aug. 2014; L. Slana Novak and T. Novak leg. (32/2014); PMSL. – <b>VL23</b> • 4 ♂♂, 1 ♀; Poljane pri Podgradu; 15 May 2011; L. Slana Novak and T. Novak leg. (40/2012); PMSL • 1 ♂; ibid.; 18 Aug. 2011; (165/2012); PMSL • 3 ♂♂, 2 ♀♀; ibid.; 20 May 2012; L. Slana Novak and T. Novak leg. (GR 3597−3601, TN det.); PMSL • 1 ♂, 2 ♀♀; ibid.; 24 Oct. 2013; L. Slana Novak and T. Novak leg. (GR 6017, 6018, 6020, TN det.); PMSL • 2 ♂♂, 4 ♀♀; ibid.; 28 Oct. 2013; L. Slana Novak and T. Novak leg. (GR 4651, 4653, 4654, 4656, 4657, 4659, TN det.); PMSL • 1 ♂, 1 ♀; Mt. Velika Medvižica; 18 Jul. 2014; L. Slana Novak and T. Novak leg. (15/2014); PMSL. – <b>VL24</b> • 1 ♂; Mala Plešivica, Golac; 20 Sep. 2012; T. Novak leg. (169/2012); PMSL • 1 ♂; ibid.; 5 Oct. 2012; L. Slana Novak and T. Novak leg. (PK 20/2012, TN rev. 2018); PMSL • 1 ♀; ibid.; 21 May 2014; P. Kozel and T. Novak leg. (PK 18/2014); PMSL • 1 ♀; ibid.; (PK 18/2014); PMSL • 1 ♂; Mt. Mala Pleševica – Mt. Lipica; 19 Oct. 2014; T. Novak leg. (26/2015); PMSL • 1 ♂; Obrov – Golac; 19 Sep. 2011; L. Slana Novak and T. Novak leg. (253/2011); PMSL • 1 ♂; Mt. Velika Pleševica; 19 Oct. 2014; T. Novak leg. (14/2015); PMSL • 1 ♂, 1 ♀; ibid.; (16/2015); PMSL • 1 ♂; Velika vrata, Mt. Velika Pleševica; 6 Oct. 2012; L. Slana Novak and T. Novak leg. (186/2012); PMSL • 4 ♂♂, 2 ♀♀; ibid.; (PK 16/2012, TN rev. 2018); PMSL. – <b>VL25</b> • 1 ♀; Buje – Suhorje; 27 Sep. 2011; L. Slana Novak and T. Novak leg. (298/2011); PMSL. – <b>VL26</b> • 3 ♂♂; Fameljska loza, Senožeče; 1 Oct. 2011; L. Slana Novak and T. Novak leg. (351/2011); PMSL. – <b>VL33</b>; PMSL • 3 ♂♂; Starod; 25 Sep. 2011; L. Slana Novak and T. Novak leg. (269/2011); PMSL. – <b>VL34</b> • 1 ♂, 1 ♀; Harije; 19 Sep. 2011; L. Slana Novak and T. Novak leg. (224a/2011); PMSL. – <b>VL43</b> • 2 ♀♀; Novokrajska jama cave, Cad. No. 810, Novokračine; 21 Jul. 2006; P. Trontelj and M. Zagmajster leg. (PK and TN 96/2020); PMSL • Novokračine; 2 ♂♂, 2 ♀♀; 27 Sep. 2011; L. Slana Novak and T. Novak leg. (307/2011, rev. 2015); PMSL. – <b>VL44</b> • 1 ♂, 1 ♀; Jablanica – Trpčane; 9 Sep. 2011; L. Slana Novak and T. Novak leg. (252/2011, rev. 2015); PMSL. – <b>VL53</b> • 1 ♂, 1 ♀; Mt. Šešnovica; 27 Sep. 2011; L. Slana Novak and T. Novak leg. (300/2011); PMSL. – <b>VL54</b> • 1 ♀; Bezgovec, Zabiče; 13 Nov.–10 Dec. 2002; B. Drovenik and A. Gergeli leg. (PK 50/2012; TN rev. 2018); PMSL • 1 ♀; ibid.; 16 Oct.–13 Nov. 2002; B. Drovenik and A. Gergeli leg. (PK 58/2012; TN rev. 2018); PMSL • 1 ♂; Mt. Snežnik; 12 Sep. 2018; L. Slana Novak and T. Novak leg. (116b/2018); PMSL. – <b>VL84</b> • 1 ♂, 1 ♀; Pleš, Borovec pri Kočevski Reki; 19 Sep. 2013; T. Novak leg. (60/2013, rev. 2015); PMSL. – <b>VL87</b> • 1 ♂; Veliki Grintovec, Zagradec, Žužemberk; 10 May 2008; L. Slana Novak and T. Novak leg. (117/2011, rev. 2015); PMSL.</p> Description <p> <b>Male</b> (holotype)</p> <p>BODY. Length 1.63, width 1.26.</p> <p>CHELICERAE. Ch basal article (without Apo) length 0.57, 2.5 times as long as wide at Ch max width at dorsal hump, and with a saw-like series of 1–3 μm high denticles on anterior margin of ventral hump (not visible on Figs). Ch-Apo small, 0.31 times as high as basal article length, rounded-trapezoid, widest distally, highest posteriorly; Ch-Apo height 0.18, max width 0.15, min width 0.09 (at the base), frontal margin quarter moon-like. Secretion field medially. Distal article length 0.58, max width 0.19, movable finger length 0.23 (Figs 4G, 5G, 6G).</p> <p> PEDIPALPS. Pa-Tr oval-trapezoid, moderately high (Pa-Tr height:length <i>~</i> 0.6:1), Pa-Fe moderately robust (Pa-Fe min:max width ~1:3.0), with 7 subequidistant low, stout, irregularly shaped tubercles and denticles in the distal Pa-Fe half. Pa-Pt moderately robust (length: max width ~ 1:4.0) (Figs 5G, 6G). Pa article lengths in Table 10.</p> <p>PENIS. Pe length 1.44, base 0.45, glans 0.12, stylus 0.07 (Figs 7G, 8G).</p> <p>LEGS. Pseudoarticle leg-Fe formula I−IV: 0−1−1−2. Tarsomere leg-Fe formula I−IV: 8−17−8−9. Leg article lengths in Table 10.</p> <p> <b>Female</b></p> <p>BODY. Length 2.23, width 1.60.</p> <p>CHELICERAE. Ch basal article length 0.60, 2.9 times as long as wide at dorsal hump, with slightly, evenly arched dorsal margin in front dorsal indentation, and medial straight portion in middle of slightly concave ventral margin (Fig. 13F). Distal article length 0.63, max width 0.20, movable finger length 0.23.</p> <p>PEDIPALPS. Pa-Tr with moderately high, arched, with straight posterior portion of dorsal margin, Pa-Fe slightly widened distally; Pa-Pt, Pa-Ti and Pa-Ta elongated (Fig. 13F). Pa article lengths in Table 10.</p> <p>OVIPOSITOR. Ovipositor length 0.94, Rec sem of 12–14 slightly elongated vesicles (Fig. 11G).</p> <p>LEGS. Leg article lengths in Table 10.</p> Distribution <p>Croatia, Slovenia. Vertical distribution in Slovenia: 320−1030 m a.s.l. Type locality: Poljane pri Podgradu (45.50° N, 14.10° E, 597 m a.s.l.), Slovenia.</p> Ecology <p> <i>Nemastoma b. martensi</i> ssp. nov. is native to lowland to submontane habitats with thermophile scrub and <i>Quercetalia pubescenti–petraeae</i> and <i>Tilia platyphyllos</i> Scop. forest communities, with partly substantial admixture of <i>Fagus sylvatica</i> and <i>Corylus avellana</i> L. in southwestern Slovenia. Phenology: adults probably eurychronous.</p>Published as part of <i>Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor & Raspotnig, Günther, 2021, Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence, pp. 1-67 in European Journal of Taxonomy 777</i> on pages 28-30, DOI: 10.5852/ejt.2021.777.1561, <a href="http://zenodo.org/record/5637112">http://zenodo.org/record/5637112</a>
Nemastoma bidentatum subsp. martensi Novak & Novak & Kozel & Schaider & Komposch & Lipovšek & Podlesnik & Paušič & Raspotnig 2021, ssp. nov.
No full textNemastoma bidentatum martensi ssp. nov. × N. bidentatum schmidti ssp. nov. Figs 3, 9O Nemastoma (Stridulostoma) seliskari Hadži, 1973a: 48, figs 33c, 37a−e. Nemastoma bidentatum bidentatum – Martens 1978: 107 [partim: Nemastoma (Stridulostoma) seliskari Hadži, 1973]. Nemastoma bidentatum bidentatum × N. bidentatum sparsum – Novak & Gruber 2000: 286 [partim: Novo mesto, Mt. Mirna gora]. Nemastoma bidentatum pluridentatum – Schönhofer 2013: 36 [partim: Nemastoma (Stridulostoma) seliskari Hadži, 1973, and Nemastoma seliskari Hadži, 1973]. Diagnosis Typical male hybrids with Ch basal article with large hump in front of dorsal indentation, Ch-Apo with knee-like frontal flexion and narrowly drawn out upper portion, Pa-Fe club-shaped, but thinner than in N. b. schmidti ssp. nov., with four simple, equidistant ventral spines in distal Pa-Fe half, and Pa-Ti with minute hump. Material examined SLOVENIA – VL15 • 1 ♂; Prelože pri Lokvi; 27 Sep. 2011; L. Slana Novak and T. Novak leg. (310/2011); PMSL. – VL35 • 3 ♂♂, 5 ♀♀; Ribnica – Ostrožno Brdo; 27 Sep. 2011; L. Slana Novak and T. Novak leg. (324/2011, rev. 2015); PMSL. – VL53 • 1 ♂; Zabiče; 27 Sep. 2011; L. Slana Novak and T. Novak leg. (301/2011, rev. 2015); PMSL. – WL05 • 1 ♂; Mt. Mirna gora; 28 Jul. 1948; A. Seliškar leg; PMSL. – WL17 • 1 ♂; Novo Mesto; Karaman leg. (107/1983 microscopic preparation, JH det., TN rev. 2017); PMSL. Remarks In Hadži’s original preparation, labelled “ Nemastoma bidentatum Roewer pulli”, with two cover glasses on the same microscope slide, there are two males and a female. One male corresponds to the type specimen of Nemastoma (Stridulostoma) seliskari Hadži, 1973 (Hadži 1973a). For this species, Hadži (ibid.) founded a new subgenus Stridulostoma based on the existence of an alleged “stridulatory organ” on the Fe III (Hadži 1973a: figs 37č–d). Gruber (1976: 797) explained that the stridulatory organ alone does not justify establishing of a new subgenus, especially because this new species differs from N. (Lugubrostoma) triste pluridentatum Hadži, 1973 exclusively by possession of this organ, and accordingly, synonymized Stridulostoma with Nemastoma. Following these remarks, Schönhofer (2013: 36) synonymized N. seliskari with N. b. pluridentatum. We found that the portion of the Fe III, identified by Hadži (1973a) as a stridulatory organ in the type specimen, is in fact a depression-damage, caused by desiccation. Moreover, such damage is present in nearly all (sic!) the other femora of this freshly moulted specimen, but nothing resembles the fine lamellated structure shown in Hadži’s drawing (1973a: figs 37č–d). No similar structure was found in the specimens of Nemastoma collected later on Mt. Mirna gora. One might imagine that a careless inspection under a microscope at low magnification could result in the illusion of a ʻstridulatory organʼ. However, the drawing of its lamellae cannot be considered anything but a pure fabrication. Moreover, Pa-Fe in Hadži (1973a: fig. 33c), being under the same cover glass in the preparation, belongs to the same specimen, although Hadži (ibid.) ascribed it to N. (Lugubrostoma) triste pluridentatum. See remark under N. pluridentatum stat. nov.Published as part of Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor & Raspotnig, Günther, 2021, Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence, pp. 1-67 in European Journal of Taxonomy 777 on pages 45-46, DOI: 10.5852/ejt.2021.777.1561, http://zenodo.org/record/563711
Nemastoma kozari Novak, Kozel, Podlesnik & Raspotnig 2021, sp. nov.
No full textNemastoma kozari Novak, Kozel, Podlesnik & Raspotnig sp. nov. urn:lsid:zoobank.org:act: 7F207FF7-9241-4854-A4C3-B7DED507F740 Figs 2, 4F, 5F, 6F, 7F, 8F, 9F, 10F, 11F, 12F, 13E; Table 9 Diagnosis Species of the N. b. complex with highly arched, symmetrical Pa-Tr dorsal margin, concave dorsal margin of Pa-Ti and equilateral triangular glans, and Rec sem of 5 elongated vesicles attached to a bladder. Etymology The species name ʻ kozari ʼ is dedicated to Senad Kozar (Travnik), whose friendship and hospitality helped us in a searching expedition on the Vlašić planina Mts. Material examined Holotype BOSNIA AND HERZEGOVINA – YK10 • 1 ♂; Mrazišče, the Vlašić planina Mts, Travnik; 44.28° E, 17.63° N; 1550 m a.s.l.; 1 Oct. 2019; S. Kozar, J. Podlesnik, P. Kozel and T. Novak leg.; light submontane beech and fir forest litter; PMSL-Opiliones-TN 137/2019. Paratypes BOSNIA AND HERZEGOVINA – YK10 • 2 ♂♂, 1 ♀; same collection data as for holotype; PMSL- Opiliones-TN 137/2019. Other material BOSNIA AND HERZEGOVINA – YK10 • 4 ♂♂, 10 ♀♀; Oštrike, Vlašić Planina Mts; 1 Oct. 2019, S. Kozar, J. Podlesnik, P. Kozel and T. Novak leg. (GR5534–5543, GR5547–5550); PMSL. Description Male (holotype) BODY. Length 1.90, body width: 1.41. CHELICERAE. Ch basal article (without Apo) length 0.61, 2.0 times as long as wide at Ch max width at dorsal hump. Ch-Apo small, 0.24 times as high as basal article length, rounded-trapezoid, widest distally; Ch-Apo height 0.14, max width 0.12, min width 0.09 (at the base), ventral margin slightly concave. Secretion field apically. Distal article length 0.60, max width 0.19, movable finger length 0.23 (Figs 4F, 6F, 9F). PEDIPALPS. Pa-Tr relatively very high (Pa-Tr height:length ~ 0.7:1), Pa-Fe moderately robust (Pa-Fe min:max width ~1:2.7), with 4 subequidistant stout spines in the distal Pa-Fe third. Pa-Pt relatively thin (length:max width ~1:4.5), Pa-Ti dorsally conspicuously concave (Figs 5F, 6F). Pa article lengths in Table 9. PENIS. Pe length 1.55, base 0.60, glans 0.13, stylus 0.04 (Figs 7F, 8F). LEGS. Pseudoarticle leg-Fe formula I−IV: 0−1−1−2. Tarsomere leg formula I−IV: 8−14−8−9. Leg article lengths in Table 9. Female BODY. Length 2.03, body width 1.67. CHELICERAE. Ch basal article length 0.63, 2.4 times as long as wide at dorsal hump, with slightly arched dorsal margin in front of indentation, and slightly concave ventral margin (Fig. 13E). Distal article length 0.66, max width 0.23, movable finger length 0.28. PEDIPALPS. Pa-Tr with moderately high, arched dorsal margin, Pa-Fe with conspicuous distal bonce, Pa-Pt and Pa-Ti elongated (Fig. 13E). Pa article lengths in Table 9. OVIPOSITOR. Ovipositor length 0.93, Rec sem consisted of bladder and 5 elongated sac-like vesicles (Fig. 11F). LEGS. Leg lengths in Table 9. Distribution Bosnia and Herzegovina, probably endemic to the Vlašić planina Mts. Vertical distribution: 1550−1630 m a.s.l. Type locality: Mrazišče, Mt. Paljenik in the Vlašić planina Mts, Travnik (44.28° N, 17.63° E, 1550 m a.s.l.), Bosnia. Ecology Nemastoma kozari sp. nov. individuals have been found in light submontane beech and fir forests, but may inhabit also other vegetation types. Phenology: adults probably eurychronous.Published as part of Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor & Raspotnig, Günther, 2021, Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence, pp. 1-67 in European Journal of Taxonomy 777 on pages 27-28, DOI: 10.5852/ejt.2021.777.1561, http://zenodo.org/record/563711
Nemastoma pluridentatum Novak & Novak & Kozel & Schaider & Komposch & Lipovšek & Podlesnik & Paušič & Raspotnig 2021, stat. nov.
No full textNemastoma pluridentatum (Hadži, 1973) stat. nov. Novak Figs 2, 4D, 5D, 6D, 9D, 10D, 12D; Table 7 Nemastoma (Lugubrostoma) triste pluridentatum Hadži, 1973a: 43, fig 33a (original description) [partim; the description also includes N. bidentatum schmidti ssp. nov., N. bidentatum gruberi ssp. nov. × N. bidentatum schmidti ssp. nov. and N. bidentatum martensi ssp. nov. × N. bidentatum schmidti ssp. nov.]. Nemastoma bidentatum bidentatum – Martens 1978: 107. Nemastoma ? bidentatum pluridentatum – Novak 2005a: 314, fig. 34. Nemastoma bidentatum pluridentatum – Schönhofer 2013: 36. Diagnosis (partly according to Hadži 1973a) Robust, short-legged member of the Nemastoma b. complex, with extremely stout Ch and Pa. Ch-Apo small, slightly trapezoid button-like, wider than high, with secretion field in frontal third. Pa-Tr nearly round, Pa-Fe and Pa-Pt very stout, Pa-Fe club-shaped, with 4 large, wide humps. Material examined Holotype BOSNIA AND HERZEGOVINA – YK10 • 1 ♂; Mt. Paljenik, Vlašić planina Mts; 1933 m a.s.l.; 18 Jun. 1962; E. Pretner leg. (4/2005, JH’s microscopic preparation); PMSL. (Note: UTM code in Novak (2005a) was erroneously cited as YJ19.) Partial redescription Note: male body and penis missing; female unknown. Male (holotype) CHELICERAE. Ch basal article length (without Apo) 0.57, stout, 1.8 times as long as wide at Ch max width at dorsal hump, distally rounded. Ch-Apo rounded low trapezoid, with longest margin distally, secretion field frontally. Distal article length 0.57, max width 0.26, movable finger length 0.28 (Figs 4D, 6D, 9D). PEDIPALPS. Pa stout, Pa-Fe extremely stout, with 4 large, bulky humps (instead of spines), club-shaped (Pa-Fe min:max width ~ 1:3.6), Pa-Pt extremely stout (length:max width ~1:2.8) (Figs 5D, 6D). Pa article lengths in Table 7. LEGS. Pseudoarticle leg-Fe formula I−IV: 0−2−1−2. Tarsomere formula I−IV: 8−13−5−8. Leg article lengths in Table 7. Remarks Under the name Nemastoma (Lugubrostoma) triste pluridentatum, Hadži (1973a) described four taxa: a) N. pluridentatum (Hadži, 1973) stat. nov. (Hadži 1973a: fig. 33a) from Mt. Paljenik (1943 m a.s.l.) in the Vlašić Mts, Bosnia, b) a hybrid N. b. gruberi ssp. nov. × N. b. schmidti ssp. nov. (Hadži 1973a: fig. 33b, č) from the surroundings of Novo mesto, Slovenia, c) a hybrid N. b. martensi ssp. nov. × N. b. schmidti ssp. nov. (Hadži 1973a: fig. 33c) from the surroundings of Novo mesto, which Hadži (ibid.) erroneously cited for Mt. Vlašić, and d) Nemastoma b. schmidti ssp. nov. from Mt. Šmarna gora, Slovenia (Hadži 1973a: fig. 34a), from the surroundings of Ljubljana (Hadži 1973a: fig. 34b), and from the cave Jama Sv. Janeza pri Prestranku near Postojna, Slovenia (Hadži 1973a: fig. 34c). Figs 33 and 34 were drawn freehand; consequently, shapes and dimensions only partly correspond to the real measurements. Hadži’s original microscopic preparation labeled “ Nemastoma bidentatum Roewer pulli, A-1065 (Novo Mesto), legit Karaman” includes two males and a female, but no juveniles. Hadži (ibid.) drew some details of the males, including damaged legs; see remarks on Nemastoma (Stridulostoma) seliskari Hadži, 1973 under N. b. martensi ssp. nov. × N. b. schmidti ssp. nov. Given that some important parts, e.g., Pa of one male, are missing, Hadži probably made this incomplete preparation after observation of the specimens under a microscope. In his original paper (Hadži 1973a), the type specimen and the type locality of N. pluridentatum stat. nov. were not designated, but he did that in the catalogue (Hadži 1973b: 13) under a mistyped name: “ N. (L.) triste pleuridentatum Hadži, 1973 ”. There he wrote: “Loc. typ.: Novo mesto ”. Since the specimens from this locality belong to the above-mentioned hybrids, Novo mesto cannot be the type locality of N. pluridentatum stat. nov.; that is the first-mentioned location in Bosnia, as fig. 33a (Hadži 1973a) is the only drawing that refers to this taxon. Unfortunately, only the preparation of one Ch, one Pa and four legs of the male holotype are preserved, while the body, as well as a further male and female mentioned by Hadži (1973a: 44) are missing. Because of the lack of any information on the Pe, the saw on Fe IV in preserved legs is the only available character allowing us to place this taxon into the N. b. complex, interpreted as N. ? bidentatum pluridentatum (Novak 2005a: fig. 34) or N. b. pluridentatum (Schönhofer 2013). In 2019, we visited the type locality Mt. Paljenik, but we did not find further specimens of N. pluridentatum. Instead we found a new, much different N. kozari sp. nov. there, which unlikely hybridize with N. pluridentatum stat. nov. Consequently, we ascribed the species status to N. pluridentatum. See remarks under N. relictum stat. nov., N. kozari sp. nov. and hybrids N. b. gruberi ssp. nov. × N. b. schmidti ssp. nov. and N. b. martensi ssp. nov. × N. b. schmidti ssp. nov. Distribution Bosnia and Herzegovina, endemic to central Bosnia, probably endemic to the Vlašić Mts. Type locality: Mt. Paljenik in the Vlašić planina Mts, Bosnia, (44.29° N, 17.64° E, 1933 m a.s.l.), Bosnia and Herzegovina. Ecology Nemastoma pluridentatum stat. nov. inhabit the highest peak areas with the Mastgrass Gentianello crispatae-Nardetum strictae Redžić 1990 communities. Phenology: adults probably eurychronous.Published as part of Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor & Raspotnig, Günther, 2021, Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence, pp. 1-67 in European Journal of Taxonomy 777 on pages 18-20, DOI: 10.5852/ejt.2021.777.1561, http://zenodo.org/record/563711
Fig. 5. Nemastoma bidentatum Roewer, 1914 in Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence
No full textFig. 5. Nemastoma bidentatum Roewer, 1914 complex, ♂♂. Pa (medial view).Published as part of Novak, Tone, Novak, Ljuba Slana, Kozel, Peter, Schaider, Miriam Gudrun, Komposch, Christian, Lipovšek, Saška, Podlesnik, Jan, Paušič, Igor & Raspotnig, Günther, 2021, Hidden diversity within the Nemastoma bidentatum Roewer, 1914 complex (Opiliones: Nemastomatidae) Part I: Morphological evidence, pp. 1-67 in European Journal of Taxonomy 777 on page 53, DOI: 10.5852/ejt.2021.777.1561, http://zenodo.org/record/563711
Overcoming obstacles in nonequilibrium holography
No full textWe study universal spatial features of certain non-equilibrium steady states corresponding to flows of strongly correlated fluids over obstacles. This allows us to predict universal spatial features of far-from-equilibrium systems, which in certain interesting cases depend cleanly on the hydrodynamic transport coefficients of the underlying theory, such as , the shear viscosity to entropy density ratio. In this work we give a purely field-theoretical definition of the spatial collective modes identified earlier and proceed to demonstrate their usefulness in a set of examples, drawing on hydrodynamic theory as well as holographic duality. We extend our earlier treatment by adding a finite chemical potential, which introduces a qualitatively new feature, namely damped oscillatory behavior in space. We find interesting transitions between oscillatory and damped regimes and we consider critical exponents associated with these. We explain in detail the numerical method and add a host of new examples, including fully analytical ones. Such a treatment is possible in the large-dimension limit of the bulk theory, as well as in three dimensions, where we also exhibit a fully analytic non-linear example that beautifully illustrates the original proposal of spatial universality. This allows us to explicitly demonstrate how an infinite tower of discrete modes condenses into a branch cut in the zero-temperature limit, converting exponential decay into a power law tail
Hydrodynamics without boosts
No full textWe construct the general first-order hydrodynamic theory invariant under time translations, the Euclidean group of spatial transformations and preserving particle number, that is with symmetry group ISOU. Such theories are important in a number of distinct situations, ranging from the hydrodynamics of graphene to flocking behaviour and the coarse-grained motion of self-propelled organisms. Furthermore, given the generality of this construction, we are are able to deduce special cases with higher symmetry by taking the appropriate limits. In this way we write the complete first-order theory of Lifshitz-invariant hydrodynamics. Among other results we present a class of non-dissipative first order theories which preserve parity
What can we deduce from facial expressions?
No full textŽe Aristotel je zapisal, da obstajajo obrazni premiki, ki spremljajo točno določeno čustvo, preučevanje obraznih izrazov pa je doživelo razcvet z raziskavami in zapisi Charlesa Darwina in nevrologa Guillauma Duchenne de Boulogne. Vprašanje, ki se pojavi ob preučevanju obraznih izrazov, je univerzalnost čustev, kar pomeni, da je izražanje le-teh enako pri vseh ljudeh in izrazi niso priučeni. Znanstveniki so identificirali šest osnovnih čustev in sicer presenečenje, strah, gnus, jeza, veselje in žalost. V diplomskem delu spoznamo delo Paula Ekmana, ki je z raziskavo izoliranega plemena v Papua Novi Gvineji predstavil močne dokaze, da so osnovna čustva univerzalna. Se pa med znanstveniki pojavlja razdvojenost, saj so metode, ki jih je uporabil Ekman, vprašljive. Carlos Crivelli in Lisa Feldman Barrett v ločenih raziskavah izoliranih plemen namreč nista našla dokazov, s katerimi bi podprla Ekmanovo teorijo. Nasprotno, njune raziskave niso pokazale univerzalnosti, temveč medkulturne razlike pri izražanju čustev. Diplomsko delo zajema tudi uporabnost poznavanja obraznih izrazov pri ugotavljanju laži, kjer problem predstavlja vpliv serij in filmov na splošno predstavo o mikro obraznih izrazih. Ljudje pogosto kontroliramo obrazne izraze, zato spoznamo razloge zakaj jih kontroliramo, tehnike upravljanja obraznih izrazov in namige, ki nakazujejo na to, da oseba kontrolira obrazno mimiko.
Empirični del diplomske naloge je sestavljen iz spletne ankete, v kateri je bilo 12 fotografij različnih obraznih izrazov, anketiranci pa so morali označiti čustvo, ki pripada izrazu na fotografiji. Na podlagi 278 odgovorov smo ugotovili, da ljudje ne prepoznajo enako obraznih izrazov, saj so se njihovi odgovori pri nekaterih čustvih močno razlikovali.Aristotle already wrote that there are facial movements that accompany a specific emotion. The study of facial expressions flourished with the research and writings of Charles Darwin and the neurologist Guillaume Duchenne de Boulogne. A question that arises when studying facial expressions is the universality of emotions, which means that their expression is the same in all people and expressions are not learned. Scientists have identified six basic emotions, namely surprise, fear, disgust, anger, joy and sadness. In the thesis, we get to know the work of Paul Ekman, who presented strong evidence that basic emotions are universal through research on an isolated tribe in Papua New Guinea. However, there is a split among scientists, as the methods used by Ekman are questionable. In separate studies of isolated tribes, Carlos Crivelli and Lisa Feldman Barrett found no evidence to support Ekman\u27s theory. On the contrary, their research did not show universality, but rather cross-cultural differences in the expression of emotions. The thesis also covers the usefulness of knowing facial expressions in detecting lies, where the problem is the influence of series and movies on the general idea of micro facial expressions. Humans often control facial expressions, so learn about the reasons why we control them, techniques for managing facial expressions and clues that indicate that a person is controlling facial expressions.
The empirical part of the thesis consists of an online survey in which there were 12 photos of different facial expressions, and the respondents had to mark the emotion that belongs to the expression in the photo. Based on 278 answers, we found out that people do not recognize facial expressions in the same way, as their answers for some of the emotions differed greatly
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