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    Euchondrus adwani Eike Neubert & Zuhair Amr 2016, n. sp.

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    Euchondrus adwani n. sp. (Figure 1) Material: Holotype NMBE 539263; paratype NMBE 539264 /1. Type locality: Syria, surrounding of the monastery of Deir Moussa, 34.0219°N 36.8423°E, 1300 m a.s.l., 11.iii. 2010, leg. Adwan Shehab. *Corresponding author. Email: [email protected] © 2016 Taylor & Francis Measurements (holotype): Height = 11.04 mm; diameter = 4.13 mm; peristome height = 4.03 mm; peristome diameter = 2.97 mm; number of whorls = 8. Diagnosis. Euchondrus adwani n. sp. differs from the widespread E. septemdentatus by its conical shell (broadly oval in E. septemdentatus), its flat suture and teleoconch whorls (suture deeper, whorls much more rounded in E. septemdentatus), the heavy palatal labial callus (weaker in E. septemdentatus), the bar-like subangularis (weaker in E. septemdentatus), and the keeled last whorl (rounded in E. septemdentatus). Description: Shell solid, dextral, cylindrical, upper part cylindro-conical in outline; shell pale brownish to horny yellow coloured; 8 rather flat-sided teleoconch whorls, suture flat with a distinct white sutural thread; teleoconch smooth, glossy, last whorl with fine, straight and irregularly spaced striae; aperture subtriangular, peristome strongly thickened by a labial callus, moderately reflected, with a rich dentition (description clockwise): palatum with a small suturalis followed by a conical palatalis superior and a broad infrapalatalis with the latter two denticles placed on a thick callus; columellar side with a basalis and a straight columellaris; parietum with a strong and long parietalis, bordered by a small spiralis, subangularis large, bar-like, left side of the parietum with another small denticle at the attachment site of the peristome; last teleoconch whorl dorsally compressed forming a distinct blunt ridge (arrows); umbilicus slit-like open, periomphalum large, dish-like. Remarks: This species shows some superficial similarities with E. desertorum Rochanaburananda in Forcart, 1981 (Figure 2), which is endemic to the Negev Desert (Heller, 2009). Both species have a straight conical shell, but E. desertorum is considerably larger than E. adwani n. sp. and its aperture is rounded and not subtriangular. It also differs in the formation of the dentition: in E. desertorum, the infrapalatalis is bifid (simple in E. adwani), the spiralis is large and connected to the parietalis (small and disconnected in E. adwani), and the subangularis is weaker (very strong in E. adwani). The last whorl of E. adwani displays a distinct keel with an enlarged periomphalum, while in E. desertorum the dorsum is rounded, and the periomphalum is much smaller. Etymology: This species is named in honour of Dr. Adwan Shawabi, who was a keen collector of molluscs from Syria, and a personal friend, and who was killed in February 2015 in the Syrian civil war (Amr, 2015).Published as part of Eike Neubert & Zuhair Amr, 2016, On a new species of Euchondrus Boettger, 1883 from Syria (Pulmonata: Enidae), pp. 58-60 in Zoology in the Middle East 62 on pages 58-60, DOI: 10.1080/09397140.2015.1132564, http://zenodo.org/record/88703

    On a new species of Euchondrus Boettger, 1883 from Syria (Pulmonata: Enidae)

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    Eike Neubert, Zuhair Amr (2016): On a new species of Euchondrus Boettger, 1883 from Syria (Pulmonata: Enidae). Zoology in the Middle East 62: 58-60, DOI: 10.1080/09397140.2015.113256

    Succinea omanensis Neubert & Damme 2012, sp. nov.

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    Succinea omanensis sp. nov. (Figs. 11–12) Type specimens: Holotype NMBE 5018975; paratypes NMBE 5018976, 5019044; paratypes Thaytiniti NMBE 5019045–5019047. Type locality: Wadi Darbat, Dhofar, Oman. Stratum typicum: Paludal biomicritic limestones of the Zalumah Formation. Age: Late Priabonian. Material: Only known from the type locality. Diagnosis: A small shelled member of Succinea, with a rapidly growing teleoconch and the last whorl exceeding half of the complete height of the shell. Etymology: omanensis refers to the provenance of this new species. Description: Small shells with a rapidly growing teleoconch, shells probably reach a total shell length of about 10 mm; preserved paratype shell with almost 4 whorls, protoconch eroded; suture shallow, well-marked and somewhat reinforced, subsuturally slightly crenulate; teleoconch covered by a regular pattern of fine, axial growth riblets; last whorl amply open, constituting more than half of the complete height of the shell. Measurements: Holotype (Fig. 11): H = 4.94 mm; Wh 3.5. Remarks: It has to be stressed that the affiliation of this species to the extant genus Succinea is debatable. However, the shells are in fact quite similar to a number of Modern species within Succinea and related genera, which today are defined by using anatomical and molecular data, inapplicable to fossils. Creating a fossil genus to harbour these shells seems not to be advisable as long as material is so meagre and good arguments in favour of such a genus are available. Today, there are only two records of an extant succineid species from the Arabian Peninsula, i.e. Quickia concisa (MORELET, 1848). The shell of this species is smaller, and it has a deep and simple suture, and thus cannot be identified with the fossil specimens from Wadi Darbat (Neubert 1998: 370, fig. 63).Published as part of Neubert, Eike & Damme, Dirk van, 2012, Palaeogene continental molluscs of Oman, pp. 1-28 in Contributions to Natural History 20 on pages 12-13, DOI: 10.5169/seals-787080, http://zenodo.org/record/583854

    Figs 3–6. Lindholmiola lens. Fig. 3 in Revision of the genus Lindholmiola HESSE, 1931 (Gastropoda: Pulmonata: Helicodontidae)

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    Figs 3–6. Lindholmiola lens. Fig. 3: syntype Helix lentiformis, ZMZ 508656, "Thessalie et Attique", D = 11.2 mm; Fig. 4: NMBE 515542, Attika, Methana, 500 m before village, 12.07.1996, leg. E. Neubert, D = 12.77 mm; Fig. 5: syntype Helix lens var. callojuncta, NHMG 372, D = 12.0 mm; Fig. 6: syntype Helix lens var. elia, SMF 101961, Kumani, Elis, coll. Jetschin ex Boettger, 1883, D = 11.94 mm. — All photos Neubert/Bochud, × 3.Published as part of Subai, Peter & Neubert, Eike, 2014, Revision of the genus Lindholmiola HESSE, 1931 (Gastropoda: Pulmonata: Helicodontidae), pp. 1-94 in Contributions to Natural History 23 on page 11, DOI: 10.5169/seals-787037, http://zenodo.org/record/584234

    Cyclotopsis praecursor Neubert & Damme 2012, sp. nov.

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    Cyclotopsis praecursor sp. nov. (Figs. 8–9) Type specimens: Holotype NMBE 5018970; paratypes NMBE 5018971–5018973, MNHN/25, SMF 340184/25; paratypes Wadi Darbat, NMBE 5018974, 5019038– 5019043. Type locality: Thaytiniti, Dhofar, Oman. Stratum typicum: Paludal biomicritic limestones of the Zalumah Formation. Age: Late Priabonian. Material: This species is very abundant at Thaytiniti and Wadi Darbat. Etymology: praecursor from Latin for predecessor, a noun in apposition. Diagnosis: A fossil species of Cyclotopsis with a slightly elevated spire and an operculum with a lamella-like coiled spiral. Description: Shell much broader than high, spire slightly elevated; protoconch relatively large, consists of 2 whorls, surface sculpture not preserved; teleoconch of 3.5–4 rapidly increasing whorls; surface sculpture of 3–5 strong spirals (only preserved on the upper whorls); last whorl only slightly descending before the aperture; aperture almost circular, with a slightly thickened lip; umbilicus very large, broad dish-like, umbilical walls very probably smooth; operculum multispiral, spiral consisting of a broad raised lamella of at least 3 whorls. Measurements: Holotype (Fig. 8): H = 6.73 mm; W = 9.18 mm; h = 4.42 mm; w = 4.25 mm; Wh = 5.5. Remarks: Roger & al. (1994) probably refer to eroded specimens of this species as 'cf. valvata '. The preservation state of the specimens from the type locality in Thaytiniti is quite poor and mainly consists of internal casts. However, in a few cases, remains of the shells themselves are preserved, allowing reconstructing of some details of the surface sculpture. These specimens display the same teleoconch sculpture as the specimens from Wadi Darbat (Fig. 9), where the preservation state is much better (but no specimen with an operculum could be traced there). This fossil species is placed in the extant genus Cyclotopsis, because it is almost indistinguishable from the few species from India (for comparison see the syntype of Cyclostoma semistriatum SOWERBY, 1843 (Fig. 10), the type species of Cyclotopsis). It shares autapomorphic details of umbilicus and operculum together with the spiral sculpture of the teleoconch. It differs from most of the Soqotran species of Dioscopoma NEUBERT, 2009, which usually have a reticulate surface sculpture, a more narrow umbilicus, and an operculum with a flat sutural line and not a raised spiral; also, most of the species of Dioscopoma display a spiral sculpure on the inner umbilical wall, which is very probably smooth in C. praecursor sp. nov. Today, the Dhofar area is inhabited by two species of the genus Rochebrunia BOURGUIGNAT, 1881, but these have larger shells with a closed umbilicus and an almost smooth operculum. For a more detailed discussion of the pomatiid genera and species of the area see Neubert (2009), and for pomatiid operculum structure Wilmsmeier & Neubert (2012).Published as part of Neubert, Eike & Damme, Dirk van, 2012, Palaeogene continental molluscs of Oman, pp. 1-28 in Contributions to Natural History 20 on pages 10-12, DOI: 10.5169/seals-787080, http://zenodo.org/record/583854

    Comment on Psonis et al. (2015): 'Evaluation of the taxonomy of Helix cincta (Muller, 1774) and Helix nucula (Mousson, 1854); insights using mitochondrial DNA sequence data'

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    Neubert, Eike, Korábek, Ondřej (2015): Comment on Psonis et al. (2015): 'Evaluation of the taxonomy of Helix cincta (Muller, 1774) and Helix nucula (Mousson, 1854); insights using mitochondrial DNA sequence data'. Journal of Natural History 49 (37): 2257-2263, DOI: 10.1080/00222933.2015.102187

    Figs 52–54. Lindholmiola spectabilis. Fig. 52 in Revision of the genus Lindholmiola HESSE, 1931 (Gastropoda: Pulmonata: Helicodontidae)

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    Figs 52–54. Lindholmiola spectabilis. Fig. 52: paratype Lindholmiola lens spectabilis, SMF 158640, Greece, Makedonía, (peninsula) Athos, monastery Kireta, D = 17.34 mm; Fig. 53: NMBE 515645, Greece, Makedonía, mountain slope N of Podohóri, (tightly coiled shell, narrow umbilicus), D = 12.02 mm; Fig. 54: NMBE 515644, Greece, Makedonía, N boundary of Taxiárhis (small, strongly ribbed, slightly scalarid form), D = 11.09 mm. — All photos Neubert/Bochud, × 3.Published as part of Subai, Peter & Neubert, Eike, 2014, Revision of the genus Lindholmiola HESSE, 1931 (Gastropoda: Pulmonata: Helicodontidae), pp. 1-94 in Contributions to Natural History 23 on page 84, DOI: 10.5169/seals-787037, http://zenodo.org/record/584234

    Figs 55–56. Lindholmiola spectabilis. Fig. 55 in Revision of the genus Lindholmiola HESSE, 1931 (Gastropoda: Pulmonata: Helicodontidae)

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    Figs 55–56. Lindholmiola spectabilis. Fig. 55: NMBE 515646, Greece, Makedonía, Kalithia in direction to Anthohóri (coarsely ribbed, slightly scalarid form), D = 11.48 mm; Fig. 56: NMBE 515647, Greece, Angítis Canyon, well at the Mara cave (large, wide umbilicus), D = 14.2 mm. — All photos Neubert/ Bochud, × 3.Published as part of Subai, Peter & Neubert, Eike, 2014, Revision of the genus Lindholmiola HESSE, 1931 (Gastropoda: Pulmonata: Helicodontidae), pp. 1-94 in Contributions to Natural History 23 on page 85, DOI: 10.5169/seals-787037, http://zenodo.org/record/584234

    Cerastus praeinsularis Neubert & Damme 2012, sp. nov.

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    Cerastus praeinsularis sp. nov. (Figs. 18–19) Type specimens: Holotype NMBE 5018986, paratype NMBE 5019050. Type locality: Wadi Darbat, Dhofar, Oman. Stratum typicum: Paludal biomicritic limestones of the Zalumah Formation. Age: Late Priabonian. Material: This species is only represented by the holotype specimen. Etymology: praeinsularis to pinpoint the early presence of this species in the area, which later gave rise to the Soqotra Archipelago, Yemen. Diagnosis: Species characterised by the large periomphalum. Description: Shell elongate turreted, with regularly increasing whorls (protoconch and first teleoconch whorl not preserved); upper teleoconch whorls covered by a fine axial sculpture of ribs; aperture broadly oval, columellar part triangular, partly covering the umbilicus; umbilicus broadly open, with a large periomphalum, delimited by a blunt crest (see arrows in Fig. 18b). Measurements: Holotype (Fig. 18a): H = 11.81 mm; Wh > 5. Remarks: For the familial and generic affiliation, the same rationale is used as in C. pseudoena sp. nov. However, the peculiar form of the umbilicus of C. praeinsularis sp. nov. clearly differs from that of its fossil congener, but is quite similar to species from the Soqotran cerastid radiation. In this respect, it particularly resembles shells of the Modern endemic Soqotran genera Achatinelloides NEVILL, 1878, and Passamaella PFEIFFER, 1877 (Neubert 2005a, 2005b). This is here shown by a comparison with Achatinelloides hadibuensis (GODWIN-AUSTEN, 1881) (Fig. 20), which shows a very similar umbilicus formation with a large periomphalum delimited by a blunt crest (see arrow in Fig. 20). In our material, we also found a small shell consisting of a small protoconch and almost four teleoconch whorls, which are also covered by a dense pattern of axial ribs (Fig. 19). The imperfect formation of the aperture makes it very probable that this is a juvenile shell. Superimposing this fossil with the holotype of C. praeinsularis sp. nov. reveals that the diameter and growing increment of both shells match almost perfectly. For this reason we interpret this shell as a juvenile of C. praeinsularis sp. nov. (NMBE 5019050).Published as part of Neubert, Eike & Damme, Dirk van, 2012, Palaeogene continental molluscs of Oman, pp. 1-28 in Contributions to Natural History 20 on pages 18-19, DOI: 10.5169/seals-787080, http://zenodo.org/record/583854

    Figs 30–32. Lindholmiola girva. Fig. 30 in Revision of the genus Lindholmiola HESSE, 1931 (Gastropoda: Pulmonata: Helicodontidae)

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    Figs 30–32. Lindholmiola girva. Fig. 30: syntype Helix girva SMF 7047, "Albanien", D = 9.22 mm; Fig. 31: NMBE 515656, Greece, Makedonía, N boundary of Taxiárhis (small rounded form), D = 8.66 mm; Fig. 32: NMBE 515655, Greece, Makedonía, 400 m S of Kehrókambos (broad form, wide umbilicus), D = 9.75 mm. — All photos Neubert/Bochud, × 3.Published as part of Subai, Peter & Neubert, Eike, 2014, Revision of the genus Lindholmiola HESSE, 1931 (Gastropoda: Pulmonata: Helicodontidae), pp. 1-94 in Contributions to Natural History 23 on page 59, DOI: 10.5169/seals-787037, http://zenodo.org/record/584234
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