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    On the generic placement of the narrow-range endemic ' Helix ' arguineguinensis Seddon & Aparicio, 1998 from Gran Canaria (Canary Islands)

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    Neiber, Marco T. (2015): On the generic placement of the narrow-range endemic ' Helix ' arguineguinensis Seddon & Aparicio, 1998 from Gran Canaria (Canary Islands). Zootaxa 3981 (2): 296-300, DOI: 10.11646/zootaxa.3981.2.1

    Monilearia Mousson 1872

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    Genus Monilearia Mousson, 1872 Type species: Helix phalerata Webb & Berthelot, 1833Published as part of Neiber, Marco T., 2015, On the generic placement of the narrow-range endemic ' Helix ' arguineguinensis Seddon & Aparicio, 1998 from Gran Canaria (Canary Islands), pp. 296-300 in Zootaxa 3981 (2) on page 297, DOI: 10.11646/zootaxa.3981.2.11, http://zenodo.org/record/24198

    Monilearia Neiber, 2015, s. str.

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    Subgenus Monilearia s. str. Monilearia (Monilearia) arguineguinensis (Seddon & Aparicio, 1998) Synonyms. Helix argonautula var. canariensis Mousson, 1874 (non Helix (Xerophila) canariensis Mousson, 1872 and Helix (Macularia) lactea var. canariensis Mousson, 1872). For further synonyms, see Seddon & Aparicio (1998). FIGURE 1. Schematic representations of the generalized penial appendage of orthurethran gastropods (A) and the vagianal appendage of genera in Cochlicellini: Obelus Hartmann, 1842 (B), Monilearia Mousson, 1872 (C), Cochlicella Férussac, 1821 (D), and Prietocella Schileyko & Menkhorst, 1997 (E) (after Ibáñez et al., 2003), and genital system (F) of Monilearia (Monilearia) arguineguinensis (Seddon & Aparicio, 1998). Abbreviations: A 1 –A 5, sections of generalized penial appendage of orthurethran gastropods and sections of vaginal appendage of Cochlicellini; ag, albumen gland; at, genital atrium; bc, bursa copulatrix; ep, epiphallus; f, flagellum; hd, hermaphroditic duct; p, penis; rm, retractor muscle; sov, spermoviduct; v, vagina; vd, vas deferens. Material examined. Spain, Canary Islands, Gran Canaria, settlement E of Argüineguin, 27 ° 45 ’ 38 ’’ N, 15 ° 40 ’04’’ W, 50 m a.s.l., February 2015, E. Schlegel leg. (1 shell, MN GEO-045 and 2 preserved specimens, ZMH 92264); Spain, Canary Islands, Gran Canaria, coll. Melvill-Tomlin, ex Wollaston (1 shell, NMW 1955.158.02113); Spain, Canary Islands, Gran Canaria, Argüineguin, syntype coll. Mousson, ex Wollaston (4 shells, syntypes of Helix argonautula var. canariensis Mousson, 1874 and of ‘ Helix ’ arguineguinensis Seddon & Aparicio, 1998, ZMZ 505021). Description. A detailed description of the shell (Fig. 2 A) is given in Wollaston (1878) or Seddon & Aparicio (1998) and therefore is not repeated here. Genitalia (Fig. 1 F): Right ommatophoran retractor independent of genitalia. Atrium short (0.9 mm), about as long as vagina (0.8 mm) and free oviduct (0.8 mm), the latter about as wide as vagina. Spermoviduct (4.0 mm) with a light touch of dark pigment, albumen gland withish, tongue-shaped (2.8 mm), hermaphroditic duct light-brownish, slightly undulating. Bursa copulatrix complex adhering to spermoviduct, with a long duct (3.2 mm) and an oval bursa (1.1 mm). Penis very short (0.6 mm), inflated, internally with a plug-shaped penial papilla with apical pore; epiphallus (6.2 mm) about 10 times as long as penis, retractor muscle inserting in its distal fifth (in relation to the position of the hermaphroditic gland). Flagellum (2.1 mm), thin, about 1 / 3 the length of epiphallus. Vaginal appendage well developed, with a hook-shaped, broad A 1 section (2.9 mm); A 2 section short (0.4 mm), slightly bent and well muscularized; A 4 section 2.8 mm long and thin, the branched A 5 section (2.8 mm) split into three thin ducts that are about as long as the A 4 section. Internal wall of genital atrium equipped with several rugose, longitudinal pilasters that can be traced into the vagina and the vaginal appendage, in the latter two stronger and several weaker pilasters are discernible. Remarks. The structure of the vaginal appendage of M. arguineguinensis (Fig. 1) corresponds to that observed in other species assigned to the genus (Hesse, 1911; Odhner, 1932; Schileyko & Menkhorst, 1997; Ibáñez et al., 2006). Ibáñez et al. (2006) included three species in Monilearia (Lyrula) Wollaston, 1878 on the basis of a similar shell sculpture consisting of a decussate ornamentation of slightly undulating radial riblets and numerous (sometimes stronger) spiral lines. The shell of M. arguineguinensis (Fig. 2) has a well-developed radial ornamentation consisting of numerous, irregular riblets, but spiral lines are absent. Therefore, M. arguineguinensis is included in the nominate subgenus here. Seddon & Aparicio (1998, p. 19) clearly intended the name ‘ Helix ’ arguineguinensis as a new replacement name for the variety from the Canary Islands introduced by Mousson (1874), i.e. for Helix argonautula var. canariensis (not Helix canariensis Mousson, 1872 or Helix lactea var. canariensis Mousson, 1872). Although Seddon & Aparicion (1998, p. 21) attributed the name ‘ canariensis ’ erroneously to Wollaston (1878), the reference to the combination used by Wollaston (1878), Disculus argonautula var. canariensis (Mousson, 1874), in the synonymy of ‘ H. ’ arguineguinensis makes it clear, which name the authors replaced by the new name. According to the ICZN code Art. 72.7 both nominal taxa have the same name-bearing type despite any contrary designation. Seddon & Aparicio (1998) designated a shell from the Mellvill-Tomlin collection (NMW 1955.158.0 2113, see also Fig. 2 B) as lectotype of ‘ Helix ’ arguineguinensis. However, there are four shells labelled as H. argonautula var. canariensis in the Mousson collection at Zurich (ZMZ 505021, see also Fig. 2 C) which are syntypes of H. argonatula var. canariensis and according to ICZN Art. 72.7 also of ‘ H.’ arguineguinensis. There is no evidence that the specimen from the collection in Cardiff (NMW 1955.158.02113) was known to Mousson when introducing the name H. argonatula var. canariensis and therefore it cannot be regarded as belonging to the syntypes series. Hence, I regard its selection as lectotype as invalid (ICZN Art. 74.2) and only the shells in the Mousson collection at Zurich (ZMZ 505021) have type status.Published as part of Neiber, Marco T., 2015, On the generic placement of the narrow-range endemic ' Helix ' arguineguinensis Seddon & Aparicio, 1998 from Gran Canaria (Canary Islands), pp. 296-300 in Zootaxa 3981 (2) on pages 297-299, DOI: 10.11646/zootaxa.3981.2.11, http://zenodo.org/record/24198

    A misinterpreted disjunction: the phylogenetic relationships of the North African land snail Gyrostomella (Gastropoda: Stylommatophora: Helicidae)

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    Neiber, Marco T., Korábek, Ondřej, Glaubrecht, Matthias, Hausdorf, Bernhard (2022): A misinterpreted disjunction: the phylogenetic relationships of the North African land snail Gyrostomella (Gastropoda: Stylommatophora: Helicidae). Zoological Journal of the Linnean Society 194 (4): 1236-1251, DOI: 10.1093/zoolinnean/zlab059, URL: https://doi.org/10.1093/zoolinnean/zlab05

    Sermyla kupaensis Lentge-MAAss & Neiber & Gimnich & Glaubrecht 2021, SP. NOV.

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    SERMYLA KUPAENSIS SP. NOV. (FIGS 1P–T, 4, 11A, B) Zoobank registration: urn: lsid: zoobank. org:act: 14715EED-3C76-4F47-AD05-9DE6F6F8127D Etymology: For the type locality, the township of Kupa in south-western Sulawesi. Type material: Holotype, Indonesia, Sulawesi, Kupa, stream E of road Makassar-Parepare, 4°7.567´S 119°37.426´E (MZB Gst. 12.191 w); 16 paratypes from the type locality (ZMB 190982); 88 paratypes from the type locality (ZMB 191388 w); 80 paratypes from the type locality (MZB Gst. 12.192 w); five paratypes from the type locality (ZMH 94335 w). Additional material examined: Indonesia: Sulawesi: Makassar, 5°8′ S 119°25′ E (ZMB 210079, 210082). Diagnosis: Shell as in S. riquetii. Veliger larvae (> 1000) are released from the subhaemocoelic brood pouch of the female. Description Shell (Fig. 1P–T): The shell of S. kupaensis cannot be distinguished from that of S. riquetii. Reproduction: Female S. kupaensis individuals were found to contain only veliger larvae (> 1000) in their brood pouches, i.e. the species has an ovoviviparous reproductive strategy. Juvenile shell (Fig. 5B): Since we did not observe any shelled juveniles in the brood pouch, we display the image of a veliger larvae with an already clearly visible operculum. We are not able to make any further statements concerning the juvenile shell of S. kupaensis. Radula (Fig. 11A, B): Typical, as in other Thiaridae (see: Glaubrecht et al., 2009). Rachidian with a subtriangular central denticle with broadly rounded tip and three smaller denticles on either side of central denticle, also triangular in shape with a rounded tip. Central denticles of lateralia broader and rounder than seen in other thiarids, flanked by two relatively large denticles forming a pronounced wing-shaped plate on the inner side projecting towards the central teeth. Marginal teeth with six to seven denticles. Ecology: This species lives in water bodies above tidal influence, although empty shells may be washed downstream. Near Kupa it was found exclusively at elevations of several dozen metres above the sea level on an elevated plateau with muddy pools of stagnant water adjacent to the river course, but not in the running stream itself. The species seems to prefer these pools with stagnant water over running water, apparently due to the richer muddy substrate it lives in and feeds from. Distribution: It is known only from the type locality. Shells found in Makassar further south on Sulawesi along the same coast line are probably also attributable to this species. Remarks Shells of S. kupaensis cannot be distinguished from S. riquetii but are, on average, slightly larger than samples from Thailand, Vietnam and Bali, for which an euviviparous reproductive strategy could be confirmed here. The different reproductive strategy, i.e. ovoviviparity, and the divergent mitochondrial haplotypes, suggest that the specimens from Sulawesi represent a distinct evolutionary lineage. Specimens of S. carbonata from Australia, which has a euviviparous reproductive strategy, usually have a higher spire than S. kupaensis. Moreover, some populations of S. carbonata can readily be distinguished by their weaker shell sculpture.Published as part of Lentge-MAAss, Nora, Neiber, Marco T., Gimnich, France & Glaubrecht, Matthias, 2021, Evolutionary systematics of the viviparous gastropod Sermyla (Gastropoda: Cerithioidea: Thiaridae), with the description of a new species, pp. 736-762 in Zoological Journal of the Linnean Society 192 (3) on pages 753-754, DOI: 10.1093/zoolinnean/zlaa120, http://zenodo.org/record/701735

    Figs 81–84 in Revision of the land snail genus Landouria Godwin-Austen, 1918 (Gastropoda, Camaenidae) from Java

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    Figs 81–84 (opposite page). Shells of species of Landouria from Java. 81. L. leucochila (Gude, 1905), Java (holotype, NHM 1922.8.29.87). 82. L. zonifera sp. nov., Yogyakarta, Kulonprogo, Sibolong (holotype, MZB 17727). 83. L. ciliocincta (Möllendorff, 1897), East Java, Zuider Mountains (lectotype, SMF 9176a). 84. L. ciliocincta, East Java, Malang, Kondangmerak, protected forest (MZB 19211). Scale bar: 5 mm.Published as part of Nurinsiyah, Ayu Savitri, Neiber, Marco T. & Hausdorf, Bernhard, 2019, Revision of the land snail genus Landouria Godwin-Austen, 1918 (Gastropoda, Camaenidae) from Java, pp. 1-73 in European Journal of Taxonomy 526 on page 58, DOI: 10.5852/ejt.2019.526, http://zenodo.org/record/315221

    Landouria menorehensis Nurinsiyah & Neiber & Hausdorf 2019, sp. nov.

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    Landouria menorehensis sp. nov. urn:lsid:zoobank.org:act: E09F6255-8F7A-49E7-A3BC-E5B795EE2015 Figs 47, 55, 71, 88; Tables 3–4 Diagnosis Landouria menorehensis sp. nov. is characterized by a whitish, globose shell with a distinct brown band above the rounded periphery and a moderately narrow umbilicus. Etymology Landouria menorehensis sp. nov. is named after its occurrence in the Menoreh karst area. Material examined Holotype INDONESIA • holotype (det. anat.); Yogyakarta, Kulonprogo, Sibolong, agroforestry; 7°44′52″ S, 110°08′12″ E; 687 m a.s.l.; 8 Jan. 2013; A. Nurinsiyah leg.; measurements: D = 13.4 mm, H = 8.6 mm; MZB 19201. Paratypes INDONESIA – Central Java, Yogyakarta • 1 spec.; Kulonprogo, Sibolong, agroforestry along the way to Sibolong Hill; 7°45′00″ S, 110°07′00″ E; 721 m a.s.l.; ZMH 133441 • 2 spec.; same data as for preceding; ZMH 133442 • 1 spec.; same data as for preceding; ZMH 133443 • 12 spec. (det. anat.); Kulonprogo, Sibolong, agroforestry; 7°44′44″ S, 110°08′02″ E; 716 m a.s.l.; MZB 17686 • 9 spec.; same data as for preceding; MZB 17702 • 5 spec.; same data as for preceding; ZMH 133311 • 1 spec. (det. anat.); Kulonprogo, Sibolong, snakefruit plantation; 7°44′44″ S, 110°08′02″ E; 716 m a.s.l.; MZB 17724 • 1 spec.; same data as for preceding; MZB 17725 • 2 spec. (det. anat.); Kulonprogo, Sibolong, near Kiskendo Cave, shrubs; 7°44′50″ S, 110°08′09″ E; 721 m a.s.l.; MZB 17644 • 1 spec.; same data as for preceding; ZMH 133444 • 6 spec.; same data as for holotype; MZB 19202 • 3 spec.; same data as for holotype; ZMH 133439 • 1 spec.; same data as for holotype; ZMH 133440 • 2 spec.; Kulonprogo, Sibolong, agroforestry; 7°44′51″ S, 110°08′12″ E; 673 m a.s.l.; MZB 18039 • 1 spec.; same data as for preceding; ZMH 133445 • 2 spec.; Kulonprogo, Jonggrangan; 7°44′20″ S, 110°08′32″ E; 730 m a.s.l.; ZMH 140034. Description SHELL (Fig. 88; Tables 3–4). Slightly depressed conical-globose, with 5.25–5.75 convex whorls; protoconch smooth; teleoconch with fine, irregular wrinkles; apical side with indistinct incised spiral lines, umbilical side with more distinct spiral lines; without scaly processes; whitish with a distinct brown band above periphery; body whorl rounded; aperture almost circular; upper insertion of peristome distinctly descending; peristome expanded, reflexed and thickened; umbilicus slightly eccentric, moderately narrow, comprising 16–24% of shell diameter, hardly obscured by columellar edge. GENITALIA (Figs 47, 71; Table 4). Atrium short; penis short, distal part cylindrical, proximal part slightly dilated, without penis coecum; membranaceous sheath around penis connected with proximal part of epiphallus by tissue; penial retractor runs from diaphragm to distal part of short epiphallus; flagellum long, broadest in its middle part, almost smooth, without node, tapering towards its proximal end; vas deferens narrow and long; vagina long; oviduct short; peduncle of bursa copulatrix subdivided into a short, broad distal part and a narrow and long proximal part, ending in an elongate oval bursa beside albumen gland. Right ommatophoral retractor runs between penis and vagina. Remarks Landouria menorehensis sp. nov. shares with L. pakidulan sp. nov. and L. ciliocincta the whitish shell with a brown band at the periphery. For differences, see remarks under L. pakidulan sp. nov. Distribution Landouria menorehensis sp. nov. is endemic to the Menoreh karst area (Fig. 55).Published as part of Nurinsiyah, Ayu Savitri, Neiber, Marco T. & Hausdorf, Bernhard, 2019, Revision of the land snail genus Landouria Godwin-Austen, 1918 (Gastropoda, Camaenidae) from Java, pp. 1-73 in European Journal of Taxonomy 526 on pages 66-67, DOI: 10.5852/ejt.2019.526, http://zenodo.org/record/315221

    (Gastropoda: Cerithioidea: Thiaridae), with the description of a new species

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    Figure 9. Structure analysis of AFLP data based on five primer combinations and a matrix corrected by AMARE for the reduced dataset of S. carbonata in Australia. Results have been assigned to their geographical localities.Published as part of Lentge-MAAss, Nora, Neiber, Marco T., Gimnich, France & Glaubrecht, Matthias, 2021, Evolutionary systematics of the viviparous gastropod Sermyla (Gastropoda: Cerithioidea: Thiaridae), with the description of a new species, pp. 736-762 in Zoological Journal of the Linnean Society 192 (3) on page 750, DOI: 10.1093/zoolinnean/zlaa120, http://zenodo.org/record/701735

    Figs 72–75 in Revision of the land snail genus Landouria Godwin-Austen, 1918 (Gastropoda, Camaenidae) from Java

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    Figs 72–75. Shells of species of Landouria from Java. 72. L. intumescens (Martens, 1867), East Java, Surabaya (lectotype, ZMB 5160a). 73. L. intumescens, East Java, Guwoterus (MZB 19207). 74. L. intumescens?, East Java, Wonosari (paralectotype, ZMB 230704). 75. L. moussoniana (Martens, 1867), East Java, Wonosari (syntype, ZMB 109902). Scale bar: 5 mm.Published as part of Nurinsiyah, Ayu Savitri, Neiber, Marco T. & Hausdorf, Bernhard, 2019, Revision of the land snail genus Landouria Godwin-Austen, 1918 (Gastropoda, Camaenidae) from Java, pp. 1-73 in European Journal of Taxonomy 526 on page 52, DOI: 10.5852/ejt.2019.526, http://zenodo.org/record/315221
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