196,276 research outputs found

    Oedematopiella Naglis, 2011, gen. nov.

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    Key to males of Oedematopiella gen. nov. 1 It 1 ventrally with a tuft of long sinuate hairs on apical third (Fig. 6), length of hairs about twice the diameter of tarsomere; It 2 distinctly broadened and about 3 times as wide as It 3; sternum 4 ventrally with dense hairs of different lengths, some of them distinctly longer than hypopygium; cercus (Fig. 7) as long as Dsur, with yellow apical setae............... sarae sp. nov. - It 1 ventrally without tuft of sinuate hairs; It 2 weakly broadened and about 1.5 times as wide as It 3; sternum 4 ventrally with dense hairs of same lengths, not longer than hypopygium; cercus (Fig. 8) shorter than Dsur, with black apical setae............................................................................................... nathaliae sp. nov.Published as part of Naglis, Stefan M., 2011, Revision of the genus Oedematopus (Diptera, Dolichopodidae) with the description of a new genus and a key to the Neotropical genera of Hydrophorinae, pp. 40-52 in Zootaxa 2756 on page 48, DOI: 10.5281/zenodo.20101

    Oedematopiella sarae Naglis, 2011, sp. nov.

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    Oedematopiella sarae sp. nov. (Figs. 1 D, 5–7) Type material. HOLOTYPE ɗ: “ Costa Rica, San Jose, San Antonio de Escazu, 2000 m, ix-x 1996, C. Flores & P. Hanson” [INBio]. PARATYPES: 5 ɗ, same data as holotype [INBio]. Description. Male. Length (holotype): body 4.4 mm, wing 4.6 mm; thorax 1.6 mm, abdomen 2.3 mm. Head: Face and clypeus with dense silvery-white pruinosity, ratio narrowest distance between eyes to distance between ocellar setae = 4.5; ratio height of face and clypeus to narrowest distance between eyes = 3.8; palpus narrow and pointed, dark with silvery-white pruinosity and brown hairs, with 2–3 strong apical setae; antenna dark brown; scape about twice as long as pedicel; pedicel about as long as high; first flagellomere ovate and about as long as high; arista dorsal and bare; frons with white pruinosity above antennae; vertex dark metallic green; pair of strong ocellar, and pair of small postvertical setae present; upper postoculars short and uniseriate, lower long, pale and multiseriate. Thorax: Mesonotum metallic green with violet reflections, with brown pruinosity, and silvery-white pruinosity on area of postpronotal lobe; ac absent; 6 strong dc present decreasing in size anteriad; scutellum with pair of strong setae, and with 2–4 smaller additional setae; pleuron with dense grayish pruinosity; propleuron with fine pale hairs. Legs: Mostly dark brown, FII on basal 2 / 3, and FIII on basal 1 / 3 yellow; CI with pale hairs and some apical setae; FI with av row of short setae on basal 2 / 3; TI with ad setae at 1 / 4 and 1 / 2; with pd setae at 1 / 8 and 1 / 2; It 1 in apical third compressed and broadened, with ventral tuft of sinuous hairs on 2 / 3, twice as long as diameter of tarsomere (Fig. 6) (MSSC), and with 2–3 strong apical setae; It 2 strongly curved basally and U-shaped, compressed and strongly broadened, about 3 times as wide as It 3, ventrally with field of minute hairs (MSSC); It 3-5 with small ventral setae; FII with ventral row of 4–5 strong setae at middle, about 4 times as long as diameter of femur (MSSC); TII with ad/pd pair at about 1 / 8 and 1 / 2; FIII with ventral row of strong curved yellow setae along entire length, about twice as long as diameter of femur (MSSC); TIII with ad/pd pair at about 1 / 8 and 2 / 3. All claws well developed, pulvilli absent. Relative podomere ratios: I: 60: 66: 32: 17: 7: 7: 7; II: 55: 74: 45: 25: 6: 4: 6; III: 84: 97: 40: 44: 19: 11: 8. Wing: With brownish tinge; R 4 + 5 and M converging before wing apex; cross-vein dm-cu at acute angle to posterior wing margin (Fig. 1 D); RMx ratio: 2.5; CuAx ratio: 2.4; lower calypter pale yellow with brown rim, and fan of brown setae; halter pale yellow. Abdomen: Dark metallic green, with brown pruinosity; sternum 4 with dense apical hairs, lateral ones sinuate and twice as long as medians, segment forming hood for partially enclosing hypopygium. Male genitalia (Fig. 7): with 2 narrow epandrial lobes, lateral lobe with 2 small setae, median lobe bare and sinuate; Vsur very small and narrow, with 1 small apical seta; Dsur with two lobes, lateral lobe L-shaped and pointed apically; cercus with two arms, lateral arm triangular, median arm with yellow flattened setae. Female: Unknown. Etymology. The new species is named after my daughter Sarah.Published as part of Naglis, Stefan M., 2011, Revision of the genus Oedematopus (Diptera, Dolichopodidae) with the description of a new genus and a key to the Neotropical genera of Hydrophorinae, pp. 40-52 in Zootaxa 2756 on pages 48-50, DOI: 10.5281/zenodo.20101

    Thrypticus caeruleus Naglis & Negrobov 2020, sp. nov.

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    Thrypticus caeruleus Naglis & Negrobov sp. nov. (Figs 1–2) Type material. HOLOTYPE ♂: Turkey, 8 km S of Cine, river bank, 68 m, 10–12.ix.2014, 37°32′34″N, 28°03′46″E, Barták & Kubíc (CULSP). PARATYPE: 1 ♂, Turkey, Akyaka, pasture, 4 m, 37°03′08.9″N, 28°20′17.4″E, 16– 22.ix.2012, Barták & Kubíc (CULSP). Diagnosis. Antennae black, thorax and abdomen shiny blue-violet, with yellow setae; legs dark with yellow knees; mid tibia without ad seta; scutellum with only 2 strong marginal setae; cercus yellow; surstylus slender, yellow with black apex. Description. Male. Body length (holotype): 1.7 mm, wing length 1.3 mm. Head: frons shiny metallic green, with blue reflections and weak brown pruinosity; face shiny blue, with violet reflections and weak grey pruinosity; palpus dark brown, with small brown setae; proboscis brown; antennal segments black; postpedicel roundish, as long as high; arista apical, with microscopic pubescense; postocular setae yellowish-white. Thorax: mesonotum metallic bluish green, with violet reflections and weak grey pruinosity; thoracic setae light yellow; 5 pairs of strong dc; 6–7 pairs of short ac; scutellum with 2 strong marginal setae, without lateral setae; 1 small yellow ppl; pleura concolorous with mesonotum. Legs: coxae and femora shiny metallic green with violet reflections, apical 1/4 of femora and knees yellow, fore and mid tibiae and tarsomeres yellowish-brown, hind tibia and tarsomeres dark brown, setae and hairs yellow except as noted. Fore leg: coxa anteriorly with small hairs and strong apical setae; remaining leg without major setae; relative podomere ratios: 32/34/16:9:5:4:3. Mid leg: coxa with some small anterior and 2 strong anterolateral setae; femur with strong posterior preapical seta; tibia with 1 strong apical posterior seta; relative podomere ratios: 34/35/20:10:6:4:4. Hind leg: coxa with 2 strong lateral setae; tibia with 1 small posterior apical seta; relative podomere ratios: 39/47/14:16:9:5:5. Wing: hyaline, veins light brown; basal section of M shorter than distal section; R 4+5 and M parallel; M joining costa posteriad of apex; CuAx ratio: 0.4; RMx ratio: 2.6; lower calypter yellowish white, with pale yellow setae; halter stem brown, knob yellowish white. Abdomen: shiny metallic blue, with violet reflections, with pale yellow hairs and setae; tergite 8 dark brown. Hypopygium (Figs 2A, B, C): epandrium dark brown with blue reflections; cercus yellow, as long as surstylus, with small apical projection; surstylus yellow, with black apex, long and slender, parallel sided, with strong ventral and 3 small subapical setae, hypandrium black in basal half, yellow in apical half, with ventral notch; phallus slender; epandrial lobes fused, of different lengths, each bearing strong apical seta. Female: unknown. Etymology. The name of the new species refers to its blue shiny thorax and abdomen. Remarks. Using the key in Negrobov & Stackelberg (1971 –77), the new species runs to T. nigricauda Wood, but can be distinguished by the characters given in the key below. The male genitalia resemble that of T. divisus (Fig. 8B) but the new species is lacking an ad seta on the mid tibia and the apex of the surstylus is almost triangular and without notches.Published as part of Negrobov, Oleg P. & Naglis, Stefan, 2020, Two new species of Thrypticus Gerstäcker from Turkey (Diptera, Dolichopodidae) with checklist and key to Palaearctic species, pp. 111-125 in Zootaxa 4858 (1) on pages 112-114, DOI: 10.11646/zootaxa.4858.1.8, http://zenodo.org/record/441163

    Oedematopus moraviensis Naglis, 2011, sp. nov.

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    Oedematopus moraviensis sp. nov. (Figs. 1 B, 3) Type material. COSTA RICA: HOLOTYPE ɗ: “ Costa Rica, San Jose, Zurqui de Moravia, 1600 m, vi/ 1996, P. Hanson” [INBio]. Description. Male. Habitus very similar to Oedematopus crassipes. Head: Face with dense silvery-white pruinosity, ratio narrowest distance between eyes to distance between ocellar setae = 10; ratio height of face and clypeus to narrowest distance between eyes = 3.0; palpus dark with silvery-white pruinosity, with short brown hairs, and 1 long pale apical hair; antenna dark brown; scape about twice as long as pedicel; pedicel about as long as high; first flagellomere ovate and about as long as high; arista dorsal, bare; frons with dark brown pruinosity above antennae and with silvery-white maculation along eye margin; vertex black with metallic reflections; pair of relatively short ocellar setae present; upper postoculars short and uniseriate, lower long, white and multiseriate. Thorax: Metallic green ground color of mesonotum mostly covered with dense black pruinosity with purplish reflections, and with silvery-white pruinosity on area of anterior dc and on postpronotum; acrostichal setae absent; 6 dc present decreasing in size anteriad and posteriormost very weak; scutellum with dense brown pruinosity, with pair of strong marginal setae, but without smaller additional setae; pleuron with dense grayish pruinosity; propleuron with fine hairs. Legs: All legs including coxae dark, femora with metallic green reflections and with grey pruinosity, trochanters partially yellow; CI with vertical row of long pale hairs, and with black apical spine-like setae; FI anteriorly and posteriorly with dense long pale hairs on basal half (MSSC); TI and It 1-4 with ventral pile; TI with some very weak dorsal setae; FII with very weak spine-like av setae on basal 2 / 3 and with field of same spine-like setae apically (MSSC), and with row of short pale pv hairs along entire length (MSSC); TII with 2 small ad and pd setae; IIt 1 with ventral pile; FIII with ventral mulitseriate row of very short spine-like setae along entire length (MSSC); TIII strongly swollen, about 2.0 times as wide as FIII, with dense short hairs (MSSC). All claws well developed, pulvilli absent. Relative podomere ratios: I: 50: 47: 27: 7: 5: 4: 10; II: 62: 71: 58: 9: 5: 4: 9; III: 77: 102: 28: 20: 10: 6: 9. Wing: With brownish tinge; R 4 + 5 curved posteriad before wing apex; cross-vein dm-cu parallel to posterior wing margin and longer than distal section of M (Fig. 1 B); RMx ratio: 2.0; CuAx ratio: 7.2; lower calypter pale yellow, with brown rim and fan of brown setae; halter stem dark yellow, knob pale yellow. Abdomen: Male genitalia (Fig. 3): lateral epandrial lobe narrow, with 1 apical and 1 subapical seta; median epandrial lobe short and curved apically; Vsur with 3 spine-like setae; Dsur only slightly curved, with two arms, lateral arm with 1 strong flattened apical seta, median arm with 3 small setae subapically; cercus narrowed apically, with long curved dorsal setae. Female: Unknown. Etymology. The new species is named after the collecting site Zurqui de Moravia in Costa Rica. Remarks. Oedematopus moraviensis and O. crassitibia are very similar, show the same MSSCs, and probably are sister-species. It is also remarkable that both species were collected at the same collecting site and date.Published as part of Naglis, Stefan M., 2011, Revision of the genus Oedematopus (Diptera, Dolichopodidae) with the description of a new genus and a key to the Neotropical genera of Hydrophorinae, pp. 40-52 in Zootaxa 2756 on pages 44-45, DOI: 10.5281/zenodo.20101

    Medetera sakhalinensis Negrobov & Naglis, sp. nov.

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    Medetera sakhalinensis Negrobov & Naglis sp. nov. (Figs 2 A–E) Type material. HOLOTYPE ♂: RUSSIA, Sakhalin, 20 km S Yuzhno-Sakhalinsk, Dachnoe, 25.VII. 1962, leg. Shamyshev. PARATYPES: 15 ♂, same data as holotype; 1 ♂, Sakhalin, 29 km SW Yuzhno-Sakhalinsk, Urozhayny, 13.VII. 1962, leg. Shamyshev; 3 ♂, Sakhalin, 38 km W Yuzhno-Sakhalinsk, Pyatireche, 28.VII. 1962, leg. Shamyshev. Types deposited in the Zoological Institute of the Russian Academy of Sciences. Description. Male. Frons black with a greenish tint, brilliant, with gray pollinosity on top of face, width of face at middle twice width of postpedicel (1.1: 0.6). Proboscis black. Palpus black, with black hairs. Antenna black. Postpedicel triangular, about as long as wide, with yellow hairs. Arista black, inserted near apex of postpedicel. Ratio length to width of postpedicel to length of arista 0.5: 0.6: 3.1. Postocular bristles below white, proboscis with some black bristles. Mesonotum greenish-black, with gray pollinosity. Propleuron with 2 short, black hairs. More than 5 small additional setae between sr and anteriormost dc; 1 npl, 1 sa, 7–8 pairs of dc, which increase in length from front to back; ac arranged in 2 rows, well developed, posteriormost bristles longer than distance between rows; 1 pair of strong scutellars with 1 pair of short lateral bristles. Legs black, knees yellow. Fore coxa with black hairs. Fore femur in apical quarter with some long av hairs. Fore tibia without strong bristles, with short hairs. Fore tarsomere 1 with short ventral hairs. Fore leg length ratio (from tibia to tarsomere 5) 3.7: 1.7: 0.9: 0.7: 0.4: 0.7. Mid femur with long ad hairs, longer than width of femur. Mid tibia in basal third with 1 ad and 1 pd bristle, apically with 1 long bristle. Mid leg length ratio (from tibia to tarsomere 5) 4.1: 2.5: 1.1: 0.8: 0.5: 0.6. Hind femur with long black bristles, about as long as width of femur. Hind tibia and tarsomeres with black short hairs. Hind leg length ratio (from tibia to tarsomere 5) 5.7: 1.4: 2.1: 1.2: 0.5: 0.6. Wing transparent, with yellow veins. Basal section of CuA without swelling. R 4 + 5 and M 1 + 2 convergent. M 1 + 2 in apical part slightly curved. Ratio costal section between R 2 + 3 and R 4 + 5 and between R 4 + 5 and M 1 + 2 2.4: 0.4. Apical segment of M 1 + 2 longer than basal segment: 5.8: 7.4. Apical part of M 3 + 4 longer than crossvein dm–cu: 1.1: 2.4. Anal lobe small, anal angle obtuse. Calypter yellow with dark yellow hairs. Halter yellow. Abdomen black, with black hairs, and gray pollinosity. Hypopygium black, surstylus and cercus dark yellow. Epandrium (Fig. 2 A) oval, tapering in apical half, length greater than width. Surstylus (Fig. 2 E) narrow, curved, not divided into dorsal and ventral arms; length greater than width; with finger–like dorsoapical projection bearing 2 setae, with ventroapical conical process, and with 1 strong apical and 1 subapical seta. Hypandrium (Fig. 2 D) parallel–sided, with rounded apex. Cercus (Fig. 2 C) elongate-oval, with 2 strong, curved apical spines. Female. Unknown. Length (mm): body 1.8–1.9, wing 2.0– 2.1. Etymology. The species is named after the collecting site Sakhalin. Remarks. In the Palaearctic key (Negrobov & Stackelberg 1971 – 77), the new species runs to Medetera borealis Thuneberg, 1955, but can be distinguished by the following characters: 1 Ventral arm of surstylus with conical process ventroapically (Fig. 2 A); hypandrium with round apex (Fig. 2 D)....................................................................................... Medetera sakhalinensis sp. nov . - Ventral arm of surstylus without conical process ventroapically; hypandrium with ovate apex........... Medetera borealisPublished as part of Negrobov, Oleg P. & Naglis, Stefan, 2015, Two new species of Medetera Fischer von Waldheim (Diptera, Dolichopodidae) from Russia and Mongolia, pp. 386-390 in Zootaxa 3964 (3) on pages 387-389, DOI: 10.11646/zootaxa.3964.3.8, http://zenodo.org/record/24004

    Thrypticus kechevi Naglis & Negrobov 2020, sp. nov.

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    Thrypticus kechevi Naglis & Negrobov sp. nov. (Figs 3–4) Type material. HOLOTYPE ♂: Turkey, Akyaka, pasture, 4 m, 37°03′09″N, 28°20′17″E, 23–27.ix.2012, Barták & Kubíc (CULSP). PARATYPE: 1 ♂, same data as holotype (CULSP). Diagnosis. Antennal scape and pedicel yellow, postpedicel black, thorax and abdomen shiny metallic green, with yellow setae; legs entirely yellow; mid tibia with strong ad seta; scutellum with 2 strong marginal setae and 2 smaller lateral setae; surstylus and cercus yellow. Description. Male. Body length (holotype): 1.8 mm, wing length 1.7 mm. Head: frons shiny metallic green, with blue reflections and weak grey pruinosity; face shiny blue, with violet reflections and dense grey pruinosity; palpus dark brown, with strong yellow seta; proboscis brown; antennal scape and pedicel yellow, postpedicel black, yellow on lateral side, roundish, as long as high; arista apical, with microscopic pubescense; postocular setae yellowish white. Thorax: mesonotum metallic green, with blue reflections and dense grey pruinosity; thoracic setae light yellow; 5 pairs of strong dc; 6–7 pairs of short ac; scutellum with 2 strong marginal setae, with 2 small lateral setae; 1 strong yellow ppl; pleura concolorous with mesonotum. Legs: coxae yellow, basal half brownish, remaining legs yellow, except tarsomere 5 brown, setae and hairs yellow except as noted. Fore leg: coxa anteriorly with small hairs and strong apical setae; remaining leg without major setae; relative podomere ratios: 32/31/16:7:5:3:4. Mid leg: coxa with some small anterior and 2 strong anterolateral setae; femur with 1 strong posterior preapical seta; tibia with 1 strong brown ad seta at 1/4 and 1 strong brown apical posterior seta; relative podomere ratios: 40/38/20:13:8:5:4. Hind leg: coxa with 2 strong lateral setae; tibia with ventral row of short erect setae and 1 small posterior apical seta; relative podomere ratios: 42/48/10:16:10:5:4. Wing: hyaline, veins light brown; basal section of M shorter than distal section; R 4+5 and M parallel; M joining costa at apex; CuAx ratio: 0.3; RMx ratio: 2.0; lower calypter yellowish white, with pale yellow setae; halter stem brown infuscated, knob yellowish white. Abdomen: shiny metallic green, with blue reflections, with pale yellow hairs and setae; tergite 8 light brown. Hypopygium (Figs 4A, B, C): epandrium dark brown with green reflections; cercus yellow, longer than surstylus, with long, slender apical projection; surstylus yellow, long and slender, parallel-sided, with 1 strong, yellow inwards curved ventral seta and 4 smaller apical setae; hypandrium black in basal half, yellow in apical half, asymmetric; phallus with broadened, flattened apex; epandrial lobes fused, of different length, each bearing strong apical seta. Female: unknown. Etymology. The new species is dedicated to the Bulgarian dipterist, Mihail Kechev. Remarks. Using the key in Negrobov & Stackelberg (1971 –77), the new species runs to T. viridis Parent, but can be distinguished by the characters given in the key below. The new species shows some similarity in the hypopygial characters with T. pruinosus (Fig. 9C) (see Remarks under that species) having a cercus with a strong apical projection, but the shape and chaetotaxy of the surstylus is quite different.Published as part of Negrobov, Oleg P. & Naglis, Stefan, 2020, Two new species of Thrypticus Gerstäcker from Turkey (Diptera, Dolichopodidae) with checklist and key to Palaearctic species, pp. 111-125 in Zootaxa 4858 (1) on pages 115-116, DOI: 10.11646/zootaxa.4858.1.8, http://zenodo.org/record/441163

    Medetera colombensis Naglis & Bickel, 2012, sp. nov.

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    Medetera colombensis sp. nov. (Fig. 5) Type material examined. HOLOTYPE: SRI LANKA. 3: “ Sri Lanka Col. Dist. Colombo Museum Gardens 50 ft. 17-23.II. 1977 Karl. V. Krombein” (USNM). Description. Male. Length (holotype): Body 2.4 mm, wing 2.3 mm. Head: Frons, face and clypeus metallic olive green with grey pruinosity; palpus metallic green; proboscis dark brown; scape and pedicel yellow, first flagellomere black; lower postoculars white. Thorax: Dorsum metallic olive-green, with dense greyish-white pruinosity; thoracic setae black, except as noted; 2 strong dc bordering mesonotal depression, anterior setae hair-like; ac small, shorter than distance between ac rows; 2 strong scutellars and 2 lateral setae one-third length of medians; pleura metallic olive-green, with grayish-white pruinosity; proepisternum with 2 white setae below. Legs: Coxae and basal 3 / 4 of femora dark brown, reminder of legs yellow, but tarsomeres slightly infuscated. Fore leg: without major setae; relative podomere ratios: 57 / 53 / 29 / 19 / 15 / 9 / 7. Mid leg: coxa with 2 white anterolateral setae; tibia with small black ad/pd setal pair at 1 / 4, and 1 black apicoventral seta; relative podomere ratios: 58 / 62 / 40 / 23 / 17 / 8 / 6. Hind leg: coxa with white lateral seta; femur with some white dorsal setae on basal half; tibia with 1 strong white subapical seta dorsally and small black apical tooth (MSSC); basitarsus without basal tubercle; relative podomere ratios: 65 / 78 / 16 / 39 / 20 / 7 / 6. Wing: Veins brown; R 4 + 5 and M distinctly converging towards wing margin; CuAx ratio: 0.4; RMx ratio: 2.8; lower calypter pale yellow, with fan of yellow setae; haltere yellow. Abdomen: Dark metallic olive-green; setae white; epandrium black, cercus brown. Male genitalia (Fig. 5): epandrium subovate; hypandrium with acute apex; epandrial seta small, positioned near epandrial lobes; epandrial lobes very close, lateral lobe bearing long, sinuous seta, median lobe with normal seta; surstylus sickle-shaped, with basoventral, short tooth-like seta and 3 long midventral setae, apex with 4-5 short setae; cercus short, with short lateral arm. Female: Unknown. Etymology. The species is named after the type locality. Remarks. Medetera colombensis belongs to the flaviscutellum species-group which includes a single Oriental species, M. philippinensis Bickel. Using Bickel’s (1987) key, the species runs to the Australian species M. gingra Bickel and M. bunyensis Bickel which do not have a yellow colored scutellum. It can be separated from all members of this species-group by the striking sickle-shaped surstylus and small cercus.Published as part of Naglis, Stefan & Bickel, Daniel J., 2012, Medetera (Diptera, Dolichopodidae) of Sri Lanka, pp. 55-63 in Zootaxa 3188 on pages 60-61, DOI: 10.5281/zenodo.21056

    Revision of the genus Oedematopus (Diptera, Dolichopodidae) with the description of a new genus and a key to the Neotropical genera of Hydrophorinae

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    FIGURE 6. Oedematopiella sarae sp. nov., male fore tarsomeres, left lateral view (scale bare = 0.1 mm).Published as part of Naglis, Stefan M., 2011, Revision of the genus Oedematopus (Diptera, Dolichopodidae) with the description of a new genus and a key to the Neotropical genera of Hydrophorinae, pp. 40-52 in Zootaxa 2756 on page 49, DOI: 10.5281/zenodo.20101

    Phoomyia Naglis & Grootaert, gen. nov.

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    <i>Phoomyia</i> Naglis & Grootaert gen. nov. <p> Type species: <i>Phoomyia srilankensis</i> Naglis & Brooks <b>sp. nov.</b></p> <p> <b>Diagnosis.</b> <i>Phoomyia</i> is distinguished from other genera of Dolichopodidae by the following combination of characters: body non-metallic, head and thorax grey, abdomen mostly yellow or brown with grey pruinosity (Fig. 3A); frons and face broad in both sexes; scape with dorsal hairs; vein M beyond crossvein dm-cu with strong anterior bend, strongly convergent with R4+5 (Fig. 3B); hind tibia of males with bifurcate posteroapical projection (Fig. 3C); hind basitarsus with 1 strong dorsal seta (Fig. 3C); male cercus extremely reduced (Figs 1 A, 1B, 2A, 2B); female terminalia with tergite 10 undivided and acutely V-shaped with acanthophorite spine-like setae encircling apex (Fig. 1 E).</p> <p> <b>Description.</b> Body length 2.0– 3.5 mm, non-metallic flies with dense grey pruinosity, legs yellow (Fig. 3A). <b>Head:</b> Face broad; eyes distinctly separated, almost parallel-sided in males and females; frons broad; scape with dorsal hairs; first flagellomere subovate, about as long as basal height; arista dorsal to apicodorsal; occiput concave; postvertical setae present. <b>Thorax:</b> 6 pairs dorsocentral setae, fifth pair strongly offset medially; acrostichal setae biseriate; 1 pair strong scutellar setae, and 1 hair-like seta laterad. <b>Legs:</b> Hind coxa with 1 strong lateral seta; mid and hind femur with strong ad preapical seta; male hind tibia with bifurcate posteroapical projection (MSSC) (Fig. 3C); hind basitarsus (Fig. 3C) with 1 strong dorsal seta; claws and pulvilli simple. <b>Wing</b> (Fig. 3B): Vein R4+5 curved posteriad in distal part; vein M with distinct bend at 1/2 beyond crossvein dm-cu, converging towards R4+5, joining costa anteriad of wing apex. <b>Abdomen:</b> Male: hypopygium (Figs 1 A-D, 2A, 2B) with epandrium distinctly longer than high; epandrial lobes weakly developed or absent; ventral lobe of surstylus shorter than dorsal lobe; postgonite with short blunt apex; hypandrium narrow; phallus slender; cercus very reduced, not longer than surstylus. Female: terminalia (Figs 1 E, 1F) with sternite 8 lacking rod-like apicolateral projections; tergite 10 undivided, acutely V-shaped in dorsal view with acanthophorite spine-like setae encircling apex.</p> <p> <b>Etymology.</b> The name is composed of the Thai word <i>Phoo</i> which means crab, and <i>myia</i> from the Greek for fly. Adults of the new genus are often found inside the entrance burrows of ghost crabs (<i>Ocypode</i> Weber) on sandy beaches. The gender is feminine.</p> <p> <b>Remarks.</b> <i>Phoomyia</i> is most closely related to <i>Argyrochlamys</i> and <i>Pseudargyrochlamys</i>, sharing non-metallic body coloration and female terminalia with tergite 10 undivided and acutely V-shaped with acanthophorite spinelike setae encircling apex (Figs 1 E, 1F). In other female dolichopodines, tergite 10 is divided medially into broad hemitergites each bearing a transverse row of acanthophorite spine-like setae (see figures in Brooks 2005). Females of <i>Phoomyia</i> lack the distinctive rod-like apicolateral projections of tergite 8 that are present in species of <i>Argyrochlamys</i> (see Brooks 2005, figs 6D, 6E, 7D, 7E). We have also observed these rod-like projections in an undescribed species of <i>Pseudargyrochlamys</i> from South Africa (Grichanov, unpublished photograph); however, their presence in the other species of this genus remains to be confirmed. The three genera can be distinguished from each other and the remaining Dolichopodinae in the key provided below.</p> <p> <b> <i>Phoomyia srilankensis</i> Naglis & Brooks sp. nov.</b> (Fig. 1)</p> <p> <b>Type material. HOLOTYPE</b> 3: <b>SRI LANKA</b>: S. Prov., Kosgoda, 44 mls SSE Colombo, 25.I.62, Loc. 21, swept on herbs on sandy beach (ZMLU). <b>PARATYPES</b>: <b>SRI LANKA</b>: 3Ƥ: same data as holotype; 23: E. Prov., Kuchchaveli, 20 mls NW Trincomalee, 9–10.II.1962, Loc. 60, swept on herbs on sandy beach; 1 3: W. Prov., Ratmalana, 9 mls S Colombo, 7–13.I.1962, Loc. 6, swept on dry grassland; 2 Ƥ: N. Prov., 2 mls W Point Pedro, 13.II.1962, Loc. 69, swept on herbs on sandy beach (all ZMLU).</p> <p> <b>Description. Male.</b> Length (holotype): body 2.2 mm, wing 2.3 mm. All setae whitish yellow except as noted. <b>Head:</b> Face broad, at narrowest part 2.5 times as wide as distance between ocellar setae, with dense silvery-white pruinosity; clypeus rounded apically, not projecting beyond lower eye margin; palp short, yellow; antennal segments yellow, with yellow setae, apical segment of arista black; scape with dorsal hairs, about as long as pedicel; pedicel short, with apical ring of short setulae; first flagellomere ovate, rounded apically, as long as high; arista dorsal to dorsoapical, bare, 2.5 times as long as first flagellomere, first segment short, second segment 6 times longer than first segment; frons broad, with dense grey pruinosity; pair of strong vertical setae on level of ocellar setae; pair of strong ocellar setae, as long as vertical setae; pair of strong postvertical setae; postocular setae uniseriate. <b>Thorax:</b> Mesonotum and scutellum covered with dense grey pruinosity; 6 pairs of strong dorsocentral setae, fifth pair offset medially; 7 pairs of acrostichal setae; scutellum with pair of strong marginal setae, and with small, hair-like lateral seta; pleura with dense grey pruinosity; propleura with small hairs in front of anterior spiracle, and with 1 strong seta ventrally. <b>Legs:</b> Entirely light yellow (including coxae). Fore leg: coxa with some strong anterior setae; femur without strong setae; tibia with strong ad setae at 1/4 and 1/2, two small pd setae at same level, and 2–3 apical setae; tarsomeres simple; relative podomere ratios: 40, 44, 22/7/6 /5/5. Mid leg: coxa bare; femur with 1 strong ad seta at 4/5; tibia with small ad seta at 1/5, strong ad setae at 1/4 and 2/3, small pd seta at 1/5, strong pd setae at 1/3 and 2/3, with strong av setae at 1/3 and 1/2, and with ring of strong apical setae; tarsomeres simple; relative podomere ratios: 52, 54, 31/ 22/15/10 /7. Hind leg: coxa with 1 strong lateral seta at 1/3; femur with 4–5 strong dorsal setae on basal 1/3, and with strong ad seta at 2/3; tibia with a row of 3 ad/pd setal pairs on basal half, 1 strong ad seta subapically, 1 long and thin, sinuous dorsoapical seta, and with 1 pale and 1 black small apically curved posteroapical projections (MSSC); basitarsus with 1 strong dorsal seta; relative podomere ratios: 53, 55, 20/ 25/13/10 /7. All claws and pulvilli simple. <b>Wing:</b> Hyaline, veins whitish yellow; R3+4 slightly curved posteriad in distal half; M with strong anterior bend at 1/2 beyond dm-cu, converging with R4+5 and joining costa anteriad of wing apex; crossvein dm-cu at 1/2 from base of CuA; CuAx ratio: 0.25; RMx ratio: 4.5; lower calypter white with white setae; halter yellowish white. <b>Abdomen:</b> Mostly dark brown, with grey pruinosity, segments 1 and 2 yellow; segments 3–5 with narrow yellow apical rim; epandrium basally dark brown, apically yellow, cercus and surstylus yellow. Hypopygium (Figs 1 A–D): epandrium about 2 times longer than wide; basiventral epandrial lobe not developed, basiventral epandrial seta positioned subapically on infolded ventral epandrial margin; apicoventral epandrial lobe very reduced, bearing modified sinuous dorsal seta and 2 closely approximated ventral setae; ventral lobe of surstylus short with 1 seta near base and several closely approximated apical setae; dorsal lobe of surstylus digitiform with thick peg-like dorsal seta and several apical setae; cercus setose, with blunt-tipped thumb-like apicolateral projection; hypandrium elongate, narrow in ventral view; phallus slender, finely wrinkled. <b>Female:</b> Similar to male, but face slightly broader; hind tibia without posteroapical projections; terminalia (Figs 1 E, 1F) as in generic description.</p> <p> <b>Etymology.</b> The name refers to the country where the species was found.</p> <p> <b>Remarks.</b> This species was referred to as “ <i>Argyrochlamys</i> sp. 1” in Brooks (2005).</p> <p> <b> <i>Phoomyia thailandensis</i> Naglis & Grootaert sp. nov.</b> (Figs 2, 3)</p> <p> <b>Type material. HOLOTYPE</b> 3: <b>THAILAND</b>: Chonburi province, Hat Nang Ram, 13.IX.2002, in crab burrows on sandy beach (reg. 22019, leg. P. Grootaert) (RBINS). <b>PARATYPES</b>: <b>THAILAND</b>: 4 3, 1Ƥ: same data as holotype; 1Ƥ: same data as holotype, but 19.IX.2002, sandy beach (22025, leg. P. Grootaert); 1 3, 1 Ƥ: Chonburi province, Sattahip, 11.IX.2002, sandy beach (reg. 22016, leg. P. Grootaert); 1 Ƥ: Rayong province, Laem Mae Phim, 24.IX.2002, sandy beach (reg. 22039, leg. P. Grootaert) (all RBINS).</p> <p> <b>Description. Male.</b> Length (holotype): body 2.8 mm, wing 2.7 mm. All setae brown yellow except as noted. <b>Head:</b> Face broad, at narrowest part 2.0 times as wide as distance between ocellar setae, with dense silvery-white pruinosity; clypeus rounded apically, not projecting beyond lower eye margin; palp short, yellow; antennal segments yellow, with yellow setae, arista black; scape with dorsal hairs; about as long as pedicel; pedicel short, with apical ring of short setulae; first flagellomere ovate, rounded apically, as long as high; arista dorsal to dorsoapical, bare, 2.0 times as long as first flagellomere, first segment short, second segment 3 times longer than first segment; frons broad, with dense grey pruinosity; pair of strong vertical setae on level of ocellar setae; pair of strong ocellar setae, as long as vertical setae; pair of strong postvertical setae; postocular setae uniseriate. <b>Thorax:</b> Mesonotum and scutellum covered with dense grey pruinosity; 6 pairs of strong dorsocentral setae, fifth pair offset medially; 5 pairs of acrostichal setae; scutellum with pair of strong marginal setae, and with small, hair-like lateral seta; pleura with dense grey pruinosity; propleura with small hairs and 1 strong seta in front of anterior spiracle, and with 1 strong seta ventrally. <b>Legs:</b> Entirely yellow (including coxae). Fore leg: coxa with some strong anterior setae; femur without strong setae; tibia with strong ad setae at 1/4 and 1/2, two small pd setae at same level, and 2– 3 apical setae; tarsomeres simple; relative podomere ratios: 50, 44, 24/ 12/9/7 /5. Mid leg: coxa bare; femur with 1 strong ad seta at 4/5; tibia with small ad seta at 1/5, strong ad setae at 1/4 and 2/3, small pd seta at 1/5, strong pd setae at 1/3 and 2/3, with strong av seta at 1/2, and with ring of strong apical setae; tarsomeres simple; relative podomere ratios: 60, 65, 43/30/ 20/15/5. Hind leg: coxa with 1 strong lateral seta at 1/2; femur with short dorsal and ventral setae on basal 2/3, and with strong ad seta at 2/3; tibia with a row of 3 ad/pd setal pairs on basal half, 1 strong ad seta subapically, 1 long and strong, sinuous dorsoapical seta, and with 1 pale and 1 black small apically curved posteroapical projections (MSSC) (Fig. 3C); basitarsus (Fig. 3C) with 1 strong dorsal seta; relative podomere ratios: 64, 65, 30/40/ 20/17/7. All claws and pulvilli simple. <b>Wing</b> (Fig. 3B): Hyaline, veins yellow; R3+4 slightly curved posteriad in distal half; M with strong anterior bend at 1/2 beyond dm-cu, converging with R4+5, joining costa anteriad of wing apex; crossvein dm-cu at 1/2 from base of CuA; CuAx ratio: 0.35; RMx ratio: 4.0; lower calypter and setae pale yellow; halter yellow. <b>Abdomen:</b> Mostly dark brown, with grey pruinosity, segments 1–5 with narrow yellow apical rim; epandrium entirely dark brown, cercus and surstylus yellow. Hypopygium (Fig. 2): epandrium about 2 times longer than wide; basiventral epandrial lobe not developed, basiventral epandrial seta positioned subapically; apicoventral epandrial lobe very reduced, bearing 1 curved and 1 straight ventral setae; ventral lobe of surstylus short with 1 subapical and 2 apical setae; dorsal lobe of surstylus digitiform with 1 subapical and 2 small apical setae; cercus with acute fin-like apicolateral projection (Fig. 2 A); hypandrium elongate, narrow in ventral view; phallus slender, finely wrinkled. <b>Female:</b> Similar to male, but face slightly broader; hind tibia lacking posteroapical projections; terminalia similar to that of <i>P. srilankensis</i>.</p> <p> <b>Etymology.</b> The name refers to the country where the species was found.</p> <p> <b>Remarks.</b> <i>Phoomyia thailandensis</i> was not only observed in crab burrows on bare sand in the more humid lower supralittoral zone, but also in crab burrows on the dryer, vegetated supralittoral embryonal dunes covered with Beach Morning Glory or Goat's Foot (<i>Ipomoea pes-caprae</i> (L.) R.Br.).</p> <p> <b>FIGURE 3.</b> <i>Phoomyia thailandensis</i> Naglis & Grootaert <b>sp. nov.</b>, male (A) habitus; (B) wing; (C) hind leg: apex of tibia and basal portion of tarsus. Abbreviation: t1, tarsomere 1 (basitarsus).</p>Published as part of <i>Naglis, Stefan, Grootaert, Patrick & Brooks, Scott E., 2013, Phoomyia, a new genus of Dolichopodinae from the Oriental Region (Diptera: Dolichopodidae), pp. 93-99 in Zootaxa 3666 (1)</i> on pages 94-97, DOI: 10.11646/zootaxa.3666.1.9, <a href="http://zenodo.org/record/283866">http://zenodo.org/record/283866</a&gt

    Dactylomyiini Naglis 2001

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    Tribe Dactylomyiini Naglis, 2001b <p> <b>Diagnosis</b> [modified from Naglis (2001b)]. Dorsocentral setae consisting of 2–3 strong setae and short reduced setae anteriad (except <i>Argentinia</i>, with 6–7 conspicuous setae); wing vein M 1 S-shaped, joining costa anteriad of apex close to vein R 4+5, with distance between M 1 and R 4+5 at wing margin at most half-length of crossvein m-m; legs devoid of major setae; male fore tarsomeres often modified with special chaetotaxy, expansions, or appendages; male claws of leg I sometimes modified or reduced; male hypopygium entirely or mainly yellow; female oviscapt with tergites 9+10 blunt and rounded, with sternite 10 (“cercus” in Naglis’ terminology) free from tergite 9+10.</p>Published as part of <i>Quevedo, Lucas, Capellari, Renato S. & Lamas, Carlos José E., 2024, A new monotypic genus of Dactylomyiini (Diptera: Dolichopodidae: Neurigoninae) from the Brazilian Chaco, with a discussion on the evolution of the lateral epandrial lobe, pp. 346-356 in Zootaxa 5403 (3)</i> on page 347, DOI: 10.11646/zootaxa.5403.3.4, <a href="http://zenodo.org/record/10561987">http://zenodo.org/record/10561987</a&gt
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