2,135 research outputs found
Ametrodiplosis adetos Elsayed, Yukawa & Tokuda 2021, n. sp.
No full text<i>Ametrodiplosis adetos</i> Elsayed, Yukawa & Tokuda, n. sp. <p>[Figures 4–5 & 10–26]</p> <p> <i>Adult</i>. <i>Head</i>. Eyes connate; facets round, 7–8 facets long at vertex. Occiput with short dorsal protuberance (Fig. 10). Antenna: flagellomere XII with microtrichose, narrow apical prolongation (Figs 12–13); female flagellomeres cylindrical, with bare necks, node about twice as long as neck (Figs 12, 14); male flagellomeres with bare internode, evanescing after flagellomere VII (Figs 13, 15). Frons with 8–12 setae (n = 4). Mouthparts (Figs 10, 11): labrum pointed, without microtrichia, with 6–8 short setae (n = 4); hypopharynx elongate, pointed, with long microtrichia on margins; labellum elongate, narrow in frontal view, with 5–7 strong setae (n = 4); palpus 4-segmented, with noticeable palpiger, segments usually consecutively longer, but in one specimen palpal segment II as long as III.</p> <p> <i>Thorax</i>. Wing (Figs 4, 5) 1.5 mm long in females (n = 2), 1.3–1.4 mm long in males (n = 3); R 1 joining C before wing mid-length; R 5 strongly curved distally, joining C posteriad of wing apex; C broken after conjunction with R 5. Acromere (Fig. 16): claws untoothed, bent and slightly widened after midlength; empodia slightly shorter than claws; pulvilli ca. 0.3 times shorter than claws. Scutum with 4 longitudinal rows of setae; scutellum laterally with 6–7 setae (n = 4). Anepimeron with 4–5 setae (n = 4); anepisternum with 1–3 scales (n = 4); katepisternum bare.</p> <p> <i>Female abdomen</i> (Fig. 17). Tergites I–VII with anterior pair of trichoid sensilla; tergites I–VI rectangular, with few lateral setae, sparse scattered scales and 1 row of posterior setae; tergite VII with few lateral setae and 2 rows of posterior setae; tergite VIII membranous, differentiated from remainder of tergum only by anterior pair of trichoid sensilla and a few setae posteriorly. Sternites II–VI with scattered setae and setiform scales near midlength, and 1 row of posterior setae; sternites III–VII with anteromedial pair of trichoid sensilla located intersegmentally; sternite VII with scattered setae and few scales near midlength and 2 rows of posterior setae; sternite VIII differentiated from remainder of sternum only by anterior pair of lateral trichoid sensilla. Ovipositor (Fig. 18): protrusible portion bare dorsally, mostly covered with short and few long setae ventrally; cerci ovoid, ca. 2.3 times longer than wide, with short setae and 2 apicoventral setae slightly thicker and longer than surrounding setae; hypoproct with 2 apical setae.</p> <p> <i>Male abdomen</i> (Fig. 19). Tergites I–VI as in female but with fewer scales; tergite VII with anterior pair of trichoid sensilla, 2 setae each placed posterolaterally, without scales; tergite 8 differentiated from remainder of tergum only by anterior pair of trichoid sensilla. Sternites II–VII as in female; sternite VIII short, about 0.5 as wide as sternite VII, with anterior pair of trichoid sensilla placed intersegmentally, and posterior and midlength groups of setae coalesced. Terminalia (Figs 20, 21): gonocoxite narrowly cylindrical except for prominent mediobasal lobe slightly before midlength; gonostylus microtrichose basally, carinate and setose distally, with cluster of setae on base ventrally; each lobe of cerci trapezoid, microtrichose, with few setae along posterior margin; hypoproct longer than cerci, with microtrichia on distal half, bilobed, each lobe with 1 dorsal and 1 ventral subapical seta; aedeagus longer than hypoproct, slightly tapered to narrow and rounded apex, with lateral sensoria on posterior half; basal portion of aedeagus not extending anteriorly beyond base line of gonocoxite.</p> <p> <i>Pupal exuviae</i>. Antennal base with tiny, pointed anteroventral umbo-like sclerotized prolongation, antennal papillae invisible. Vertex with 2 cephalic papillae on each side, outermost papillae with long seta. Face with 2 setose and 2 asetose median papillae, and triplet lateral papillae on each side, 2 asetose and 1 setose (Fig. 22). Prothoracic spiracles (Fig. 23) pigmented, elongate, 24–27 μm long (n = 4), curved, with trachea extending to tip, ca. 4.5 times longer than cephalic seta. Segments II–VI with rudimentary abdominal spiracles. Terga I–VIII with pair of trichoid sensilla anteriorly and short spicules on anterior third; terga II–VIII with 2–3 horizontal rows of spine-like spicules on anteromedian third; terga I–VII with 2 asetose and 4 setose dorsal papillae; tergum VIII with 2 setose dorsal papillae.</p> <p> <i>Larva. Third instar</i>. In life creamy white, body cylindrical. Spatula with rounded lobes (Fig. 24). Ventral and dorsal papillar pattern basic for Cecidomyiidi (Möhn 1955). Terminal segment: ventrally (Fig. 25) with smooth median perineal pads each bearing 1 asetose anal papilla, 2 posterolateral smooth plaques each bearing 2 asetose anal papillae, surface anterior and lateral to anus covered with pointed and raised cuticular warts; anal opening surrounded by microtrichia; dorsally (Fig. 26) covered with tiny pointed verrucae and with 2 long setose papillae and 6 large corniform papillae, most anterior 2 corniform papillae thinner than others, innermost 2 shorter than others.</p> <p> <b>Etymology</b>. The species name is derived from the Greek word “adetos”, meaning free, for the fact that the larvae live freely in the flowers of <i>Tylophora aristolochioides</i> without gall-induction.</p> <p> <b>Holotype.</b> 1♂ (ELKU): Reared by A. K. Elsayed and emerged on 5.viii.2019 from a flower of <i>Tylophora aristolochioides</i> collected by A. K. Elsayed & K. Mochizuki in Shiramine Village, Hakusan City, Ishikawa Prefecture, Japan on 16.vii.2019.</p> <p> <b>Paratypes.</b> All collected from flowers of <i>Tylophora aristolochioides</i> in Japan; 5 larvae: collected on 8.ix.2019 by K. Mochizuki & S. Nemoto in Shirakawa City, Fukushima, Japan. The remaining paratypes were collected at the type locality by A. K. Elsayed & K. Mochizuki and reared by A. K. Elsayed: 3 pupal exuviae & 2 ♀♀: emerged on 1.viii.2019; 1 pupal exuviae & 1♂: emerged on 5.viii.2019; 2 pupal exuviae: emerged on 2.viii.2019; 1 pupal exuviae & 1♀: emerged on 30.vii.2019; 2 ♀♀: emerged on 14.vii.2019; 3 ♀♀: emerged on 2.viii.2019; 4 larvae: collected on 16.vii.2019.</p> <p> <b>Distribution.</b> Japan, Honshu: Ishikawa and Fukushima Prefectures.</p> <p> <b>Life history and biological notes</b>. Larvae of <i>A. adetos</i> are ectophagous on the flowers of <i>Tylophora aristolochioides</i> (Apocynaceae) and do not induce galls (Fig. 1). Pupation takes place in the soil. In the laboratory, the adults emerged 2–3 weeks after larvae were transferred to rearing cups. <i>Ametrodiplosis adetos</i> has several overlapping generations in summer due to the long flowering season of <i>T. aristolochioides</i> that extends from July to September.</p> <p> <b>Remarks</b>. <i>Ametrodiplosis adetos</i> is most similar to <i>A</i>. <i>mamajevi</i> Kovalev, 1972, sharing with it the curved R 5 distally, narrow gonocoxites, mediobasal lobes placed on the basal half of gonocoxites and trapezoid lobes of cerci. They can be distinguished as follows: aedeagal base ends before gonocoxal bases in <i>A</i>. <i>adetos</i> but exceeds the gonocoxal bases in <i>A</i>. <i>mamajevi</i>; mediobasal lobes are more pronounced in <i>A</i>. <i>adetos</i>; and the male flagellomere XII ends with a narrow apical prolongation in <i>A</i>. <i>adetos</i> but lacks the apical prolongation in <i>A</i>. <i>mamajevi</i> (Kovalev 1972).</p>Published as part of <i>Elsayed, Ayman Khamis, Yukawa, Junichi, Mochizuki, Ko, Tokuda, Makoto & Kawakita, Atsushi, 2021, Three new species of Ametrodiplosis (Diptera: Cecidomyiidae) from Japan, with a key to the Japanese species and a molecular phylogenetic analysis, pp. 151-172 in Zootaxa 4942 (2)</i> on pages 155-159, DOI: 10.11646/zootaxa.4942.2.1, <a href="http://zenodo.org/record/4600392">http://zenodo.org/record/4600392</a>
Stefaniella skuhravae Elsayed, new species
No full textStefaniella skuhravae Elsayed, new species Adult description. Color of freshly emerged individuals: head black, antennae light brown, thorax brown, wings transparent, legs light brown, upper and lower parts of abdomen light brown, lateral parts orange. Body length. 0.7 mm (n= 7) in males and 1.1 mm (n= 5) in females. Head (Fig. 24). Compound eyes with circular facets. Gap between eyes on vertex about 0.5-1.5 times as wide as facet. Palpi 2 –segmented, second segment elongated, nearly longer than the first. Antenna 2 + 9–10 flagellomeres; scape conical, pedicel rounded, flagellomeres barrel-shaped in both sexes; each with two connected rings of circumfila (Fig. 25); the apical two flagellomeres sometimes fused. Thorax.Wing (Fig. 26) length 0.6 mm (n= 7) in males and 1.0 mm (n= 5) in females. Vein R 5 joining C before its mid-length; C broken beyond the attachment point with R 5; M present; CuA simple. Tarsal claws (Fig. 27) toothed. Empodia as long as, or shorter than, claws. Abdomen, male. Tergites 1–7 rectangular, with a posterior row of strong, hyaline setae; tergites 2–7 with one pair of anterior, small, trichoid sensilla; tergite 8 very narrow, about 0.33 width of tergite 7, with median pair of trichoid sensilla. Sternites 1–7 with scattered setae, in addition to posterior row of strong hyaline setae; sternum 8 undifferentiated from the surrounding membranous tissue. Genitalia (Fig. 28): Gonocoxite slightly elongate, with setulose mediobasal lobe. Gonostylus 0.5 times as long as gonocoxite, arched, ending with a strong tooth. Cerci fused at base, forming one apically notched sclerite, setulose. Hypoproct entire, setulose, shorter than tips of cerci, with rounded tip. Parameres dorsally covered with dense tiny setae, and surrounding aedeagus. Aedeagus cylindrical, straight, slightly longer than parameres, with truncate tip. Female: (Fig. 29): Tergites 1–7 with 1–2 posterior rows of strong, hyaline, setae, and anterior trichoid sensilla; tergite 8 weakly sclerotized, about half width of tergite 7, divided into two sclerites. Sternites 2–7 with 1–2 posterior rows of hyaline setae. Ovipositor: segment 8 with lateral group of strong, curved, internally directed setae; membranous part rugose with papillae surrounded with tiny spines. Segment 9 (ovipositor trunk) with two sclerotized rods that widen posteriorly, forming weakly sclerotized triangular plate covered with tiny spines. Lateral plate bearing about 25 thick and strong setae. Aculeus straight, thick, tapered at apex, with two rows of tiny setae, every row consists of ~ 22 setae. Apical lamella rectangular and setose. Holotype. Female, Egypt, El-Amria district (30 ° 59 '54.00"N, 29 ° 49 '7.00"E), 4. VI. 2013, A. K. Elsayed reared from galls on male floral inflorescences of Atriplex halimus. Paratypes. All material from Egypt, El-Amria district, Alexandria, reared by A. K. Elsayed from galls on male floral inflorescences of A. halimus. 1 males, 12.VI. 2013; 1 male, 13.VI. 2013; 2 females, 14.VI. 2013; 2 females, 3 males, 16.VI. 2013; 4 males, 17.VI. 2013; 5 males, 2 females, 19.VI. 2013; 8 males, 20.VI. 2013; 1 female, 22.VI. 2013; 2 females, 24.VI. 2013. Distribution. Egypt (El-Amria district). Etymology. This species is named in honor of Mrs. Marcela Skuhravá, the Czech entomologist and expert on the family Cecidomyiidae (Diptera). Biology. Larvae of S. skuhravae induce small, slight swellings (Fig. 9 and 10) on male floral inflorescences of the salt marsh plant Atriplex halimus. The gall consists of a single chamber, and pupation takes place inside it. The pupal exuviae protrude from the emergence hole, and can be distinguished by their hyaline color. The galls were collected and the adults emerged from the end of May to October 2013. Remarks. The genus Stefaniella contains 9 species (Gagné & Jaschhof 2014). Dorchin & Freidberg (2008) revised all the species and found no significant differences between them in morphological characters. They concluded that study of the immature stages is needed, and molecular study will be useful to determine relationships between the species. They added that the currently the best characters for distinguishing species of Stefaniella are those of their galls. There are two known species of Stefaniella that induce galls on A. halimus: S. atriplicis Kieffer, 1898 and S. trinacriae De Stefani, 1900 (Dorchin & Freidberg 2008, Gagné & Jaschhof 2014). Stefaniella atriplicis induces small stem galls, each gall about 4–5 mm in diameter and multiple chambers (Skuhravá et al. 2007). Stefaniella trinacriae induces large galls on the stems, each gall about the size of a hazelnut and having multiple chambers. In contrast to the preceding species, S. skuhravae induces small galls on the male floral inflorescence, and each gall consists of only a single chamber. Therefore, we consider it to be a new species.Published as part of Elsayed, Ayman Khamis, Skuhravá, Marcela, Karam, Hedaya Hamza, Elminshawy, Abdelaziz & Al-Eryan, Mohamed Awad, 2015, New records and new species of gall midges (Diptera: Cecidomyiidae) developing on Chenopodiaceae in Egypt, pp. 105-115 in Zootaxa 3904 (1) on pages 112-114, DOI: 10.11646/zootaxa.3904.1.6, http://zenodo.org/record/23434
Ametrodiplosis aeroradicis Elsayed, Yukawa & Tokuda 2021, n. sp.
No full textAmetrodiplosis aeroradicis Elsayed, Yukawa & Tokuda, n. sp. [Figures 6, 7 & 27–45] This species is morphologically similar to A. adetos except for the following characters: Adult. Head (Figs 27–32). Frons with 5–9 setae (n = 7). Male flagellomeres with bare internode evanescing after flagellomere X but slight constriction remaining in flagellomeres XI–XII (Fig. 30). Mouthparts: labrum with 6–8 short setae (n = 7); labellum with 8–10 strong setae (n = 9). Thorax. Wing (Figs 6, 7) 2.2–2.6 mm long in females (n = 5), 1.8–1.9 mm long in males (n = 3); R 5 slightly curved distally. Scutellum laterally with 5–11 setae (n = 7). Anepimeron with 4–7 setae (n = 8); anepisternum with 2–5 scales (n = 5). Female abdomen (Figs 34, 35). Tergite VII with scales only on posterior third. Sternites VI with 2 rows of posterior setae; sternite VII with 3 rows of posterior setae. Ovipositor: protrusible portion with 2 lateral setae postero-dorsally; cerci ca. 2.6 longer than wide. Male abdomen (Figs 36–38). Tergite VI with few scales, no lateral setae and 1 row of posterior setae. Terminalia: gonostylus with microtrichia from base to before midlength dorsally and from base to midlength ventrally; hypoproct with deep triangular notch. Pupal exuviae (Figs 39, 40). Antennal base with elongate, pigmented, serrate protuberance, nearly as long as cephalic seta. Prothoracic spiracles 26–27 μm long (n = 2), ca. 3.3 times longer than cephalic seta. Terga II–VIII with 3–4 horizontal rows of spine-like spicules on anteromedian third of segment. Larva. Third instar (Figs 41–43). Orange in life. Spatula with pointed lobes. Anus surrounded by median perineal pads covered with pointed and raised cuticular warts. Most anterior pair of corniform terminal papillae thinner than others; median pair wider than others. Larva. Second instar (Figs 44, 45). Light orange in life, body cylindrical. Spatula absent. Integument ventrally with anteromedian field of tiny pointed cuticular warts on anterior half of thoracic segments and abdominal segments I–VIII, dorsally and laterally covered pointed verrucae on anterior two thirds and rounded verrucae on distal third. One group of two setose and one asetose lateral papillae present on each thoracic segment. Ventral papillae invisible. Two sternal papillae on each thoracic segment and four sternal papillae on abdominal segments I– VIII, all without setae. Two pairs of setose pleural papillae present on thoracic and abdominal segments I–VIII. Terminal segment: ventrally with median perineal pads each bearing 1 asetose anal papilla and few rows of pointed cuticular warts surrounding anal opening, 2 posterolateral smooth plaques each bearing 2 asetose anal papillae, surface anterior and lateral to anus covered with tiny, pointed cuticular warts; dorsally covered with pointed verrucae and with 2 long setose papillae and 6 large corniform papillae, most anterior 2 corniform papillae thinner than others, median 2 wider than others, innermost 2 shorter than others. Etymology. The species name, aeroradicis, is based on the two Latin words “aer” for air and “radices” for root to indicate that galls of this species are formed on the aerial roots. Holotype. 1♂ (ELKU): emerged on 23–24.vi.2011 from gall on aerial root on Trachelospermum asiaticum collected by S. Sato on 19.vi. 2011 in Kanou, Kiyotake, Miyazaki City, Miyazaki Prefecture, Japan. Paratypes. All from aerial root galls on Trachelospermum spp. in Japan; 4 second instars: collected by K. Ogata on 18.ix.2019 from galls on T. gracilipes var. liukiuense in Tanegashima, Nishinoomote City, Kagoshima Prefecture and dissected on 2.x.2019 by A. K. Elsayed; 1♂: same as the holotype; 2♀♀ & 2 pupal exuviae: collected by S. Sato on 19.vi.2011 from galls on T. asiaticum at the type locality, emerged on 22.vi.2011; The following specimens were reared or obtained from the aerial root galls on T. asiaticum that were collected by H. Ikenaga from Shiroyama, Kagoshima City, Kagoshima Prefecture: 3♀♀ emerged on 22.vi.1977; 1♂, 1♀ on 15.iv.1978; 1♀ on 26.vi.1977; 1♀ on 28.vi.1977; 1♀ & 1 pupal exuviae on 22.vi.1977; 1♂ & 1 pupal exuviae on 29.vi.1977; 1♀ on 26.vi.1977; 1♀ on 29.vi.1977; 2 third instars obtained on 4.iii.1978. In addition, J. Yukawa reared or obtained the following specimens from the root galls on T. asiaticum in Shiroyama: 1♂, reared on 15.iv.1978; 1♀, on 23.iv.1978; 2 third instar larvae obtained on 27.v.1972. Distribution. Widespread in Japan: Honshu (Fukushima, Chiba, Tokyo, The Izu Islands, Shizuoka, and Wakayama Prefectures), Kyushu (Fukuoka, Oita, Miyazaki, and Kagoshima Prefectures), and Okinawa Prefecture. Life history and biological notes. Ametrodiplosis aeroradicis induces irregular, brown, spherical galls of 2.3– 7.0 mm in diameter on the aerial roots of Trachelospermum asiaticum and T. gracilipes var. liukiuense (Apocynaceae) (Fig. 2). The galls are sometimes fused and each harbors one to four larvae in separate chambers. Larvae can be found in the galls in all seasons of the year and pupation takes place in the gall. Remarks. Two species of Ametrodiplosis are known to induce irregular spherical galls on the roots of their host plants: A. recondita Gagné on Pycnanthemum tenuifolium Schard. (Lamiaceae) in North America (Gagné 2018a) and A. auripes (Löw) on Galium mollugo (Rubiaceae) in Europe (Löw 1888). Although the life history and the gall shape of these two species resemble those of A. aeroradicis , they are associated with distant hosts and are very different morphologically (Gagné 2018a; Möhn 1955). Ametrodiplosis aeroradicis is most similar to A. adetos with which it shares mediobasal lobes that are closer to base than to distal part of gonocoxites, narrower gonostylus from base to apex, and aedeagal base not exceeding the gonocoxal base. They can be distinguished easily as follows: male flagellomeres VIII–XII are slightly constricted in A. aeroradicis, but cylindrical in A. adetos; R 5 is less curved distally in A. aeroradicis; mediobasal lobes are less protruding from gonocoxites in A. aeroradicis; pupal antennal bases of A. aeroradicis possess long serrate apical protuberances but tiny, acute apical protuberances in A. adetos; and anterior lobes of larval spatula are pointed and narrower in A. aeroradicis but rounded and wider in A. adetos.Published as part of Elsayed, Ayman Khamis, Yukawa, Junichi, Mochizuki, Ko, Tokuda, Makoto & Kawakita, Atsushi, 2021, Three new species of Ametrodiplosis (Diptera: Cecidomyiidae) from Japan, with a key to the Japanese species and a molecular phylogenetic analysis, pp. 151-172 in Zootaxa 4942 (2) on pages 161-165, DOI: 10.11646/zootaxa.4942.2.1, http://zenodo.org/record/460039
A taxonomic revision and molecular phylogeny of the eastern Palearctic species of the genera Schizomyia Kieffer and Asteralobia Kovalev (Diptera, Cecidomyiidae, Asphondyliini), with descriptions of five new species of Schizomyia from Japan
No full textThe genus Asteralobia (Diptera, Cecidomyiidae, Asphondyliini, Schizomyiina) was erected by Kovalev (1964) based on the presence of constrictions on the cylindrical male flagellomeres. In the present study, we examine the morphological features of Asteralobia and Schizomyia and found that the male flagellomeres are constricted also in Schizomyia galiorum, the type species of Schizomyia. Because no further characters clearly separating Asteralobia from Schizomyia were observed, we synonymize Asteralobia under Schizomyia. Molecular phylogenetic analysis strongly supports our taxonomic treatment. We describe five new species of Schizomyia from Japan, S. achyranthesae Elsayed &amp; Tokuda, sp. n., S. diplocyclosae Elsayed &amp; Tokuda, sp. n., S. castanopsisae Elsayed &amp; Tokuda, sp. n., S. usubai Elsayed &amp; Tokuda, sp. n., and S. paederiae Elsayed &amp; Tokuda, sp. n., and redescribe three species, S. galiorum Kieffer, S. patriniae Shinji, and S. asteris Kovalev. A taxonomic key to the Japanese Schizomyia species is provided
Ametrodiplosis stellariae Elsayed, Yukawa & Tokuda 2021, n. sp.
No full textAmetrodiplosis stellariae Elsayed, Yukawa & Tokuda, n. sp. [Figures 8, 9 & 46–62] This species is morphologically similar to A. adetos except for the following characters: Adult. Head (Figs 46–51): Frons with 5–7 setae (n = 5). Internode of male flagellomeres bare, evanescing after flagellomere IX. Mouthparts: labrum with 5–6 short setae (n = 6); labellum with 6–9 strong setae (n = 6). Thorax. Wing (Figs 8, 9) 2.2–2.6 mm long in females (n = 4), 2.0– 2.1 mm long in males (n = 3). Acromere (Fig. 52): empodia nearly as long as claws. Scutellum laterally with 5–9 setae (n = 6). Anepimeron with 5–7 setae (n = 6); anepisternum with 2–4 scales (n = 6). Female abdomen (Figs 53, 54). Tergites I–VI with few lateral setae, few scattered scales on midlength and 1 row of posterior setae; tergite VII with few lateral setae and few scattered on 2 rows of posterior setae. Ovipositor: cerci, about twice longer than wide, with short setae on distal half. Male abdomen (Figs 55–57). Tergites without scales; tergite 7 with 1–2 setae placed posterolaterally. Terminalia: Basal portion of aedeagus extending anteriorly beyond baseline of gonocoxites. Pupa (Figs 58, 59). Antennal bases with pigmented, short, pointed protuberances. Prothoracic spiracle 26 μm long (n = 1), ca. 4.3 times longer than cephalic seta. Terga II–VIII with 3–4 horizontal rows of spine-like spicules on anteromedian third of segment. Larva. Third instar (Figs 60–62). Orange in life. Spatula with pointed lobes. Anus surrounded by median perineal pads covered with pointed and raised cuticular warts. Corniform terminal papillae similar in length; most anterior 2 corniform papillae thinner than others. Etymology. The species name, stellariae, is based on the host genus name. Holotype. 1♂ (ELKU): Reared by R. Sonobe and emerged on 5.viii.2019 from a leaf bud gall on Stellaria uliginosa collected by R. Sonobe in Ryuzu, Nikko, Tochigi Prefecture, Japan on 16.vii.2019. Paratypes. All collected and reared from leaf bud galls on Stellaria uliginosa in Japan; 2 third instar larvae: collected on 16.xi. 2014 in Yakumo village, Matsue City, Shimane Prefecture by T. Nodue. 2♂♂ 5♀♀: same as the holotype. 1 pupa: collected on 14.x.2016 by R. Sonobe at the type locality. 1 pupa & 4♂♂ 1♀: same as the holotype, but collected on 9–15.xi.2016. Distribution. Japan, Honshu: Tochigi, Shiga, Nara, Hiroshima, and Shimane Prefectures. Life history and biological notes. Ametrodiplosis stellariae induces leaf bud galls on Stellaria uliginosa (Fig. 3). Usually one, sometimes two or more larvae develop in each gall. Larva usually pupates in the gall but occasionally in the soil. The number of annual generations has not been clarified. A similar sort of leaf bud gall was found on two congeneric plant species, Stellaria sessiliflora Y. Yabe, in Nara and Tokushima Prefectures, and Stellaria uchiyamana Makino var. apetala (Kitam.) Ohwi in Nara Prefecture. In addition, Cerastium fontanum Baumg. (Caryophyllaceae) also bears similar leaf bud galls in Hokkaido. The host range of A. stellariae requires further study. Remarks. One European species of Ametrodiplosis is presently known from Stellaria L., namely A. duclosii (Tavares) that induces leaf bud galls on S. graminea L. in France and Germany (Tavares 1930; Stelter 1961; Gagné & Jaschhof 2017). Ametrodiplosis stellariae is distinct from A. duclosii as follows: the gonocoxite of A. stellariae are much narrower, its mediobasal lobe are closer to the gonocoxal bases but closer to the distal parts of gonocoxites in A. duclosii (Stelter 1961), and the anterior lobes of the larval spatula are more pointed in A. stellariae (Tavares 1930). Ametrodiplosis stellariae is most similar to A. mamajevi and A. adetos with which they share the curved R 5 distally, narrow gonocoxites and mediobasal lobes placed on the basal half of gonocoxites (Kovalev 1972). Ametrodiplosis stellariae can be distinguished from A. adetos by the less pronounced mediobasal lobe, the basal portion of aedeagus that exceeds anteriorly beyond the base line of gonocoxite, the short-pointed protuberance on pupal antennal bases, the narrow anterior lobes of larval spatula and the more elongate corniform terminal papillae in third instars. Ametrodiplosis stellariae differs from A. mamajevi as follows: the male hypoproct is deeply notched in A. stellariae , but shallowly notched in A. mamajevi; male cerci are truncate in A. stellariae, but lobes of cerci trapezoid in A. mamajevi; male flagellomere XII ends with narrow apical prolongation in A. stellariae but without such prolongation in A. mamajevi.Published as part of Elsayed, Ayman Khamis, Yukawa, Junichi, Mochizuki, Ko, Tokuda, Makoto & Kawakita, Atsushi, 2021, Three new species of Ametrodiplosis (Diptera: Cecidomyiidae) from Japan, with a key to the Japanese species and a molecular phylogenetic analysis, pp. 151-172 in Zootaxa 4942 (2) on pages 165-166, DOI: 10.11646/zootaxa.4942.2.1, http://zenodo.org/record/460039
Primofavilla aegyptiaca Elsayed, new species
No full textPrimofavilla aegyptiaca Elsayed, new species Adult description. Color of freshly emerged females: head black, antennae brown, thorax dark brown, wings transparent, legs light brown, upper and lower parts of abdomen black, lateral parts red and covered with scales. Body length. 1.8 mm (n= 2) in female. Head (Fig. 19). Compound eyes with circular facets. Gap between eyes on vertex about 0-2 times as wide as facet. Palpi 2 -segmented, the second segment slender, elongated, slightly longer than the first. Antenna 2 + 10 - segmented; scape conical; pedicel rounded; flagellomeres barrel-shaped, each with two connected rings of circumfila, except the terminal flagellomere, with two sets of two connected rings of circumfila (Fig. 20). Thorax. Wing (Fig. 21) length about 1.5 mm (n= 2) in female; vein R 5 joining C at its mid-length; C broken beyond the junction point with R 5; M present; CuA simple. Tarsal claws (Fig. 22) curved and toothed. Empodia much shorter than claws, about as long as the small basal tooth. Female Abdomen. (Fig. 23): Tergites 1–7 rectangular, setulose, and with 1–2 posterior rows of strong setae; tergite 8 about 0.3 width of the tergite 7. Sternites 2–7 rectangular, setulose, with posterior row of hyaline setae. Ovipositor: segment 8 with lateral group of strong curved setae, the membranous part rugose with papillae surrounded with tiny spines. Lateral plate with ~ 26 strong, straight and thick setae. Aculeus curved and bare. Apical lamella oval, setose; the basal third of the dorsal margin covered with filiform short setae and the apical two thirds covered with short lanceolate setae. Holotype. Female, Egypt, Alexandria, El-Amria district (30 ° 59 '54.00"N, 29 ° 49 '7.00"E), 26.V. 2013, A. K. Elsayed, reared from leaf galls on leaves of Atriplex halimus. Paratypes. 1 female, 26.V. 2013, El-Amria district, Alexandria, reared by A. K. Elsayed from galls on leaves of Atriplex halimus. Distribution. Egypt, Alexandria, El-Amria district. Etymology. The name of that species is derived from Egypt. Biology. Larvae of P. aegyptiaca induce globular galls, 2-3 mm in diameter, on both surfaces of leaves (Fig. 8) of the salt marsh plant A. halimus. Each infested leaf has 2-6 galls, usually beyond the mid-rib. The galls were found in May and June 2013. The galls were collected and preserved in test tubes to rear the adult stage, but that method was not very successful, as only 2 females emerged and the pupation site was not determined. Another method was tested to rear the adults, the galls were collected and put in a plastic jar with soil at the bottom, but no adults emerged. Remarks. The three known species of Primofavilla, P. initialis Mamaev 1972, P. kaplini Fedotova 1991, and P. cystiphorae Fedotova 1991, are associated with Atriplex salina Siev., A. nana Parr-Sm., and A. tatarica L., respectively (Fedotova 1991 b, Gagné & Jaschhof 2014). The identification of Primofavilla species could be determined by the diagnostic morphological characters of the female ovipositor (Mamaev 1972, Fedotova 1991 b). The dorsal margin of the ovoid apical lobe of P. kaplini has only filiform setae, in contrast to P. aegyptiaca, P. cystiphorae and P. initialis, which have the filiform setae only on the basal third of the dorsal margin with the remaining setae either lanceolate (P. aegyptiaca and P. cystiphorae) or squamiform (P. initialis). Primofavilla aegyptiaca is closest to P. cystiphorae, but the former has dorsal and ventral sclerotized plates at the base of segment 9 of the female abdomen, while the latter does not. Primofavilla aegyptiaca also differs from P. cystiphorae in the distribution and number of strong setae on the lateral plate, which do not extend to the ventral side in P. aegyptiaca, unlike in P. cystiphorae. In addition to these differences, P. aegyptiaca has a bare saddle-like projection at the base of the aculeus. Therefore, P. aegyptiaca is considered to be a new species. Very similar galls were shown by De Stefani (1942) on the same host plant in Sicily, Italy, but the gall inducer was not identified, strongly suggesting the presence of P. aegyptiaca in Italy.Published as part of Elsayed, Ayman Khamis, Skuhravá, Marcela, Karam, Hedaya Hamza, Elminshawy, Abdelaziz & Al-Eryan, Mohamed Awad, 2015, New records and new species of gall midges (Diptera: Cecidomyiidae) developing on Chenopodiaceae in Egypt, pp. 105-115 in Zootaxa 3904 (1) on pages 111-112, DOI: 10.11646/zootaxa.3904.1.6, http://zenodo.org/record/23434
Baldratia karamae Elsayed and Skuhrava, new species
No full textBaldratia karamae Elsayed and Skuhravá, new species Adult description. Color (freshly emerged individuals): head black, antennae brown, thorax dark brown, wings smoky grey, legs light brown, upper and lower parts of abdomen dark brown, lateral parts red. Body length. 1.8 mm (n= 5) in female when the ovipositor not extended and 1.6 mm (n= 5) in male. Head (Fig. 11): Compound eyes with rounded facets, gap between eyes on vertex about 0-1 times as wide as facet. Palpi one-segmented; labella globular, setose, widely separated. Antenna 2 + 10 –segmented (n= 23), scape conical, pedicel rounded, flagellomeres 1–9 subequal, slightly longer than wide, each with two connected rings of circumfila in both sexes; male terminal flagellomere with circumfila arranged in a network pattern (Fig. 12); female terminal flagellomere (Fig. 13) consisting of the fusion of the three distal flagellomeres. Thorax: Wing (Fig. 14) length about 1.3 mm (n= 5) in male and 1.4 mm (n= 5) in female. Vein R 5 joining C approximately at mid-length; C broken behind the junction point with R 5; Sc and M present; CuA simple. Tarsal claws (Fig. 15) toothed and curved. Empodia shorter than claws. Hind legs of males much longer and thicker than fore- and midlegs of the female. Abdomen, Male: Tergites 1–7 rectangular with posterior row of setae; tergites 3–7 with anterior pair of trichoid sensilla. Tergite 8 about 0.3 times as wide as tergite 7. Sternites rectangular; sternites 1 and 3–5 with posterior row of setae; sternites 2, 6 and 7 with two posterior rows of setae. Genitalia (Fig. 16): Gonostylus about 0.6 times as long as gonocoxite, arched, setulose and setose, apically with blunt tooth. Gonocoxite wide, massive with scattered long setae. Mediobasal lobes small. Cerci fused, notched, setose and setulose, with rounded tips. Hypoproct entire, rounded apically. Aedeagus slender, and rounded at apex, surrounded with wide setulose parameres. Female (Fig. 17): Tergites 2–7 rectangular, with anterior pair of trichoid sensilla and posterior row of setae; tergite 8 about half tergite 7. Sternites rectangular; sternites 3–6 with posterior row of setae; sternites 6 and 7 with 1–2 posterior rows. Ovipositor (Fig. 18): segment 9 anteriorly with dorsal and ventral dark sclerotized patches, posteriorly with some hyaline setae; the two sclerotized rods widened basally. Lateral plate bears ~ 21 straight, hyaline, split setae. Aculeus concave ventrally, with three rows of strong, squamiform, apically hooked setae on the dorsal site. Sclerotized thin spine extends dorsally along the lateral plate. Apical lamella ovoid, densely covered with short setae. Holotype. Female, Egypt, El-Amria district (30 ° 59 '54.00"N, 29 ° 49 '7.00"E), 27.I. 2013, A. K. Elsayed, reared from pustule galls on leaves of Suaeda pruinosa. Paratypes. All from Egypt, Alexandria, and reared by A. K. Elsayed from leaf galls on Suaeda pruinosa. El- Amria district: 2 females, 1 male, 29.I. 2013; 2 females, 30.I. 2013; 1 female, 17.III. 2013; Abo-Talat district: 1 male, 7.III. 2013; 1 female, 27.IV. 2013; 1 female, 30.IV. 2013; Sidi Kreer district: 2 females, 1 male, 4.V. 2013; 1 female, 1 males, 5.V. 2013; 1 female, 1 male, 7.V. 2013; 4 females, 8.V. 2013; 1 female, 2 males, 15.V. 2013. Distribution. Egypt (Sidi Kreer, Abo-Talat, and El-Amria district). Etymology. This species is named in honor of Mrs. Hedaya H. Karam, professor of Economic Entomology at Alexandria University, Egypt. Biology. Larvae of B. karamae develop inside leaves of S. pruinosa (Chenopodiaceae). Attacked leaves do not show any external signs of infestation except for a dark reddish spot, but can be recognized once adults have emerged, leaving behind emergence holes and the protruding pupal exuviae. Each gall consists of a single chamber in which pupation takes place. The adults were collected from plants from the end of January to the beginning of March, and from the end of April to the middle of October, 2013. Baldratia karamae may have more than two generations per year. Remarks. According to Fedotova (1991 a) the genus Baldratia is divided into five groups on the basis of morphological characters of adults. By reviewing these characters, it was clear that the new species, B. karamae, belongs to the salicorniae Group, which is characterized by the apical lamella positioned at an obtuse angle relative to segment 9, and the lateral plate embraces the entire base of the apical lobe. The salicorniae Group previously contained three species, viz. B. salicorniae, B. suaedifolia, and B. balchanensis (Fedotova 1991 a, 1992). The thin spine of the female ovipositor is longer and thinner in B. suaedifolia, and B. balchanensis than in B. salicorniae. This new species has a long thin spine that does not exceed the base of the aculeus, in contrast to B. balchanensis which has a longer thin spine. Baldratia suaedifolia has a thin spine covered with split setae, while it is bare in B. karamae. Currently only five gall midge species are known to be associated with the host plant Suaeda (Gagné & Jaschhof 2014). Two of them, B. aelleni Möhn, 1969, and B. suaedae Möhn, 1969, were described on the basis of larvae alone (Möhn 1969), and can therefore not be compared to other adults in the genus. Baldratia karamae larvae that develop in leaves of Suaeda pruinosa, differ from the three other species, viz. B. occulta Dorchin, 2001, associated with S. monoica Forssk; B. suaedifolia Fedotova, 1991, associated with Suaeda acuminata (Meyer); and B. terteriani Mamaev & Mirumian, 1990, associated with Suaeda altissima (L.), on the basis published descriptions of these species (Mamaev & Mirumian 1990, Fedotova 1991 a, Dorchin 2001). An unique feature of the B. karamae is the stable number of antennal flagellomeres (2 + 10) in both sexes, in contrast to other species of Baldratia which have a variable number of flagellomeres between the sexes, viz. B. suaedifolia (2 + 12 in female versus 2 + 10 in male), B. occulta (2 + 13-14 in female versus 2 + 12 in male), and B. terteriani (2 + 14 in female versus 2 + 12 in male). The lateral plate of the ovipositor of B. karamae is broad at its base, narrow in the middle and covered with split setae, but B. occulta has a lateral plate which is narrow at the base and bearing 10-15 straight setae, with split setae only on the basal part. The lateral plate of B. terteriani has a small lateral projection at the base that is not present in B. karamae.Published as part of Elsayed, Ayman Khamis, Skuhravá, Marcela, Karam, Hedaya Hamza, Elminshawy, Abdelaziz & Al-Eryan, Mohamed Awad, 2015, New records and new species of gall midges (Diptera: Cecidomyiidae) developing on Chenopodiaceae in Egypt, pp. 105-115 in Zootaxa 3904 (1) on pages 108-110, DOI: 10.11646/zootaxa.3904.1.6, http://zenodo.org/record/23434
Elsayed Elshahed, Das Problem des transzendenten sinnlichen Wahrnehmung in der spätmu-ʽtazilitischen Erkenntnistheorie nach der Darstellung des Taqīaddīn an-Nağrānī. Berlin, Klaus Schwarz Verlag, 1983
No full textPeters Jan. Elsayed Elshahed, Das Problem des transzendenten sinnlichen Wahrnehmung in der spätmu-ʽtazilitischen Erkenntnistheorie nach der Darstellung des Taqīaddīn an-Nağrānī. Berlin, Klaus Schwarz Verlag, 1983. In: Bulletin critique des annales islamologiques, n°3, 1986. pp. 51-52
Family violence and its relationship between Coping strategies and Psychological hardness among female students of king Abdul Aziz University and Omm Al-Qura University
No full textThis study aims to identifying the relationship between family violence and all of coping strategies and Psychological hardness to University students, identifying the differences between high of family violence university students and low of family violence university students in coping strategies and Psychological hardness, and identifying the differences between students of king Abdul Aziz University and Omm Al-Qura University in family violence, coping strategies and Psychological hardness, and identifying the differences between students in Scand level semester and eighth level semester in family violence, coping strategies, and Psychological hardness, participants sample of university students (females)(N=500)(250 of king Abdul Aziz University and 250 Omm Al-Qura University). The apparatus are family violence scale prepared by Ghanema, H.A (2011), coping strategy scale prepared by Elsayed, F. KH, (2011)and Psychological hardness prepared by Elsayed ,F.KH(2007) . Results revealed that there is a relationship between family violence and all of coping strategies and Psychological hardness to University students, there is a differences between high of family violence university students and low of family violence university students in coping strategies and Psychological hardness, but on differences between students of king Abdul Aziz University and Omm Al-Qura University in family violence, coping strategies and Psychological hardness, and no differences between students in Scand level semester and eighth level semester in family violence, coping strategies, and Psychological hardness. Key words:- Family violence - Coping strategies- Psychological hardness - University students
Pseudasphondylia tominagai, a new gall midge species (Diptera: Cecidomyiidae) inducing flower bud galls on Eleutherococcus spinosus (Araliaceae) in Japan
No full textThe genus Pseudasphondylia (Diptera: Cecidomyiidae: Asphondyliini: Asphondyliina) comprises ten Palearctic, Oriental and Australian species associated with various hosts belonging to at least ten plant families.
A new species, Pseudasphondylia tominagai Elsayed & Tokuda n. sp., that induces flower bud galls on Eleutherococcus spinosus (L.f.) S.Y.Hu (Araliaceae) is described. This species is considered to alternate between host plants seasonally. A key to males of known Pseudasphondylia species is provided
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