196,258 research outputs found
Laemostenus (Iranosphodrus) troglophilus Tahami & Muilwijk & Lohaj & Sadeghi 2017, sp. nov.
No full textLaemostenus (Iranosphodrus) troglophilus Muilwijk, Tahami & Lohaj, sp. nov. (Figs. 23, 25, 27, 29, 31) Type material. Holotype male, labelled: “Iran, Yazd, Tange Chenar, Nafas Cave; 28.viii.2014; leg. M.S. Tahami, H. Entezari, M. Sheykh Rasti, Meshki”/ Laemostenus troglophilus sp. nov. Muilwijk, Tahami & Lohaj; 2016 (red label Holotype). Paratypes: one male and one female, the same data as holotype / Laemostenus troglophilus sp. nov. Muilwijk, Tahami & Lohaj 2016 (red label Paratype). Holotype and one paratype deposited in ZM–CBSU, one paratype in cMU. Diagnosis. A medium sized, brachypterous Laemostenus sg. Iranosphodrus species. Upper side including legs and antennae dull black, female paratype with bluish shine on elytra. Pronotum and head shiny, with fine microsculpture, elytra matt, with distinct microsculpture (Fig. 23). Description. BL 15.8–16.0 (holotype) mm. Head narrow, parallel-sided, distinctly longer than wide (HL/HW: 1.1), with only posterior pair of supraorbital setae; frontal furrows short, but distinct; eyes flat, reduced (0.5 mm), shorter than temples (1.0 mm). Clypeus with two long setae; labrum with six setae. Antennae in holotype missing, antennae in paratypes rather short, reaching anterior eighth of elytral length, antennomere 3 without accessory pubescence except a few apical setae, antennomeres 4–11 pubescent. Mandibles short. Pronotum cordiform, distinctly wider than head, as long as wide (PL/ PW: 1.0). Basal impressions short, sparsely punctuated. Lateral bead with superficial punctuation, two antero-lateral setae present, basolateral setae missing. Anterior angles prominent, at apex rounded, posterior angles almost rectangular. Elytra ovate elongate, relatively broad (EL/EW: 1.6–1.7), with maximum width at apical third; disc with small antero-medial depression; base wider than pronotal base. Humeri moderately rounded, with a small tooth. Striae fine, with punctures, intervals flat. Scutellar striae present; scutellar setiferous punctures situated in striae 1. Umbilicate series consists of 18 setiferous punctures, more interrupted in middle; two setae at apex of stria 7. Mesosternum with small teeth before mesocoxae, in male paratype reduced. Metepisternum longer than wide (Fig. 25). Abdominal sternites black, with superficial wrinkles and dense microsculpture. Legs short. Meso- and metatibiae almost straight, not curved, with short ventral brush of setae near apex. Profemori without teeth at ventral sides. Protibiae with sparse, short hairs between cleansing apparatus and apical setae. Tarsi with decumbent pubescence at dorsal sides; claws with small teeth, male protarsi slightly dilated. Aedeagus: (Figs. 27, 29) median lobe of aedeagus short (1.93 mm), slightly arcuate, regularly narrowing to the apex in lateral view; dorsal view, apex moderately rounded in dorsal view. Ovipositor: as on Fig. 31. Differential diagnosis. L. (I.) troglophilus sp. nov. differs from L. yazdensis sp. nov. by the presence of small mesosternal tooth, absence of tooth on profemori, by the shape of pronotum and metepisternum as well as median lobe of aedeagus. Both new species differs from L. (I.) rudichae by straight mesotibiae, more oval elytra, and more short and stout appendages. Distribution and habitat. So far known only from the twilight zone of the type locality, Nafas cave. Etymology. The name is taken from the Greek origin, “ trōglo ” means “cave”, and “ philus ” means “liking” or “attracted to”, generally means cave-lover.Published as part of Tahami, Mohadeseh Sadat, Muilwijk, Jan, Lohaj, Roman & Sadeghi, Saber, 2017, Study of Laemostenus species across Zagros and Central zone of Iran, with the description of seven new cavernicolous species and notes on subgenus Iranosphodrus (Coleoptera: Carabidae: Sphodrini), pp. 115-136 in Zootaxa 4344 (1) on pages 130-131, DOI: 10.11646/zootaxa.4344.1.4, http://zenodo.org/record/104231
Laemostenus (Iranosphodrus) yazdensis Tahami & Muilwijk & Lohaj & Sadeghi 2017, sp. nov.
No full textLaemostenus (Iranosphodrus) yazdensis Muilwijk, Tahami & Lohaj, sp. nov. (Figs. 22, 24, 26, 28, 30) Type material. Holotype male, labelled: “ Iran, Yazd, Taft, Badamestan Cave; 13.iii.2016; leg. M.S. Tahami, M. Tahami, V. Rahmatyan, M. Dehghanpour ”/ Laemostenus yazdensis sp. nov. Muilwijk, Tahami & Lohaj, 2016 (red label Holotype). Paratypes: one female, the same data as holotype, one male (immature) and one female labelled: “ Iran Yazd, Taft Badamestan Cave, 14.x.2014 leg. M.S. Tahami, H. Darvishnia”/ Laemostenus yazdensis sp. nov. Muilwijk, Tahami & Lohaj, 2016 (red label Paratype). Holotype and two paratypes deposited in ZM–CBSU, one paratype in cMU. Diagnosis. A small to medium sized, brachypterous Laemostenus sg. Iranosphodrus species; colour dark brown (holotype) to dull black (paratypes), antennae and legs reddish-brown. Pronotum and head with fine microsculpture, elytra dull, with distinct microsculpture (Fig. 22). Description. BL 12.4–15.5 mm (holotype 12.8 mm). Head narrow, parallel-sided, as long as wide (HL/HW: 1.0), with only posterior pair of supraorbital setae; frontal furrows fine, superficial, eyes flat, reduced (0.5 mm), shorter than temples (0.6 mm). Clypeus with two long setae; labrum with six setae. Antennae short, reaching anterior sixth of elytral length, antennomere 3 without accessory setae except a few apical ones; antennomeres 4– 11 pubescent. Mandibles short. Pronotum elongate, cordiform, distinctly wider than head, slightly wider than long (PL/ PW: 0.9). Basal impressions short, finely punctate. Lateral margins finely punctate, with two antero-lateral setae, basolateral setae missing. Anterior angles widely rounded, posterior angles almost rectangular. Elytra ovate elongate, rather wide (EL/EW: 1.6), with maximum width in posterior third; disc depressed; base wider than pronotal base. Humeri distinct, with small tooth. Striae fine, finely punctate, intervals slightly convex. Scutellar striae present; scutellar setiferous punctures situated in striae 1. Umbilicate series consists of 18 irregularly situated setiferous punctures; two setae at apex of stria 7. Mesosternum unarmed, without teeth before mesocoxae. Metepisternum as long as wide (Fig. 24). Abdominal sternites black, with superficial wrinkles and dense microsculpture. Legs short, meso- and metatibiae not curved, straight, ventral side of mesotibiae with a short apical brush, metatibiae with more extended brush. Profemori with tooth, which is reduced in immature male paratype. Tarsi dorsally with decumbent pubescence, claws with reduced teeth; male protarsi dilated. Aedeagus: (Figs. 26, 28) median lobe short (1.77 mm) and narrow, strongly arcuate and narrowing to apex in lateral view; apex widely rounded in dorsal view. Ovipositor: as on Fig. 30. Differential diagnosis. L. (I.) yazdensis sp. nov. differs from L. rudichae by straight mesotibiae in males, unarmed mesosternum, more oval elytra and more short and stout appendages. Distribution and habitat. So far known only from the twilight zone of the type locality, Badamestan cave, Yazd province. Etymology. Topotypic, named after the province in which the cave is situated.Published as part of Tahami, Mohadeseh Sadat, Muilwijk, Jan, Lohaj, Roman & Sadeghi, Saber, 2017, Study of Laemostenus species across Zagros and Central zone of Iran, with the description of seven new cavernicolous species and notes on subgenus Iranosphodrus (Coleoptera: Carabidae: Sphodrini), pp. 115-136 in Zootaxa 4344 (1) on pages 128-129, DOI: 10.11646/zootaxa.4344.1.4, http://zenodo.org/record/104231
Philorhizus flavocorpus Muilwijk & Seiedy & Wrase 2021, sp. n.
No full textPhilorhizus flavocorpus sp. n. urn:lsid:zoobank.org:act: 3F40F6E0-963D-47B3-9311-33F0A1DBE036 (Figs. 4.1, 4.2) Type material. Holotype ♂ labeled: “ Iran, Kerman / Kuh-e Bahr Aseman / 22.iv.2007 / Muilwijk leg.“ // “ Holotype / Philorhizus flavocorpus sp. n. / Muilwijk, Seiedy & Wrase des. 2021“ (black print on red label). Paratypes with same data but “ Paratype “. 1 ♀ same data as holotype and red label // “ Paratype “. 1 ♀: “ Iran, Kerman / Kuh-e Lalezar / 3400–3600 m / 04.vi.2014, 29°27´52´´ N 56°45´65´´ E / A. Weigel leg. IR 10a HF” // Philorhizus flavocorpus sp. n. // “ Paratype “ red label. Holotype deposited in HMIM, 1 paratype in CJM and 1 paratype in CAW. Description. A small sized Philorhizus (3.1–3.2 mm.) with elytra strongly widened to apex; unicolorous, slightly shining pale yellowish including appendages (Fig. 4.1); microsculpture superficially in male, more evident in female; brachypterous. Head longer than pronotum (HL/PL: 1.16); eyes slightly convex; temples oblique to neck. Labium convex, with 5 setae; antennomeres 1–3 without hairs except normal setae, 4–11 with fine, long, suberect hairs. Microsculpture consisting of large isodiametric superficial meshes. Pronotum slightly transverse (PW/PL: 1.3), widest at insertion of lateral seta. Anterior angles not protruding. Lateral sides regularly sinuate to rounded hind angles. Lateral furrow narrow, with 2 setae, one at anterior fourth, the other at the hind angle. Base medially more or less rectilinear, laterally toward hind angles oblique. Middle line well-developed. Basal impression prominent. Microsculpture consisting of superficially transverse, somewhat irregular meshes. Elytra long-oval (EL/EW: 1.35); humeri strongly rounded; strongly widened to apex, with maximum width at about 1/6 of the apex. Striae 1–3 superficially, other striae evanescent. Scutellar striole present. Marginal umbilicate series consist of 14 setigerous pores. Microsculpture consisting of superficial isodiametric meshes. Legs metatibia in females slightly bent, in males straight; meso- and metatarsi with long claws with 2–3 small dents; meso- and metatarsomere 1 about the same length as tarsomere 5. Ventral side with wide ligula, slightly rounded at apex; mentum without tooth. Anal segment with 2 setae on each side, segment 4–5 with 1 seta in middle on each side. Microsculpture consists of superficially transversal meshes; more evident on anal segment. Median lobe (Fig. 4.2) relatively stout, 0.84 mm long, apex pointed in lateral view. Endophallus with two twisted bands of scales and spines, the middle band more extensive than the apical one. Comparative notes. This wingless species resembles the mono-coloured species such as Philorhizus tinauti Anichtchenko, 2005 or P. alpinus Meschnigg, 1934 from the West Palearctic due to its yellow colour. However, the combination of features and especially the structure of the endophallus are different from all other species of this genus. Distribution. Iran (Kerman / Kuh-e Bahr Aseman). Habitat. The type specimens were collected in sub-alpine grasslands interspersed at lower elevations with shrubberies. This species is uncapable of flight, hence its distribution area is certainly restricted. 3.1 habitus holotype, scale: 5 mm. 3.2 median lobe holotype, lateral view, scale: 1 mm. 3.3 Atranus ruficollis habitus, scale: 5 mm. Alborz. 3.4 Atranus ruficollis median lobe, lateral view, scale: 1 mm. Alborz. 4.1 habitus holotype, scale: 2 mm. 4.2 median lobe holotype, rl = right lateral, v = ventral, ll = left lateral, scale 0.2 mm. Etymology. The name refers to its body colour. Identification references: Sciaky (1991); Wrase (2005); Wrase & Assmann (2008); Arndt et al. (2011); Allegro et al. (2015).Published as part of Muilwijk, Jan, Seiedy, Marjan & Wrase, David W., 2021, Contribution to the knowledge of Carabidae in Iran with the descriptions of five new taxa, proposing of four synonyms and providing of nine new records (Insecta: Coleoptera), pp. 55-81 in Zootaxa 5067 (1) on pages 62-64, DOI: 10.11646/zootaxa.5067.1.3, http://zenodo.org/record/565650
Atranus dariushi Muilwijk & Seiedy & Wrase 2021, sp. n.
No full textAtranus dariushi sp. n. urn:lsid:zoobank.org:act: D65213E8-E48D-4639-A60E-1BA6A5E3FC7A (Figs. 3.1, 3.2) Type material. Holotype, ♂, labelled: “ Iran, Kohgiluyeh and Boyer-Ahmad / Suq cave, 812 m / 30°49ˊ25.9´´ N, 50° 25ˊ37.9´´ E / 16.vi.2018 / S. Rahideh & J. Muilwijk leg.” // Atranus dariushi sp. nov. / Muilwijk, Seiedy & Wrase des. 2021“ (black print on red label). Holotype deposited in HMIM. Comparing material: Atranus ruficollis (Gautier des Cottes, 1858): 1 ♂, Iran, Alborz, Mazandaran, near Siah Bisheh, ca 2000 m, 14.vi.2017; Muilwijk leg.; CJM. Description. Elytra and appendices yellow, head and pronotum slightly shiny yellowish red; dorsum pubescent; eyes flat; pronotum slender; elytra elongate elliptical; micropterous. (Fig. 3.1). Body length 6.4 mm. Head temples long; eyes flat; frons lateral flat impressed, wrinkled in the middle; finely pubescent on basal part of vertex; with anterior margin weakly emarginate, clypeus with two setae; mandibles long and pointed; palpi long and slender; 2 pairs of supraorbital setae, the posterior setae far behind the eye; antennae from antennomere 2 onwards pubescent; reaching to one third of the elytra length. Pronotum slender, cordate (PW/PL 1.02), lateral sides slightly sinuated and constricted to the base; greatest width at about the anterior fourth; anterior angles protruding, posterior angles subrounded, apically slightly upwards pointed; basal fovea moderately deep and round extending along the lateral side; lateral margin with 2 pairs of setae, one pair basolateral, the other pair at the widest point; middle line from apex to base. Pubescence less dense and finer on the disc. Elytra elongate-elliptical (EL/EW: 1.55); shoulders strongly rounded, without teeth; striae at the disc coarsely punctated, finer towards the apex, striae 1–4 at the base impressed, superficially near the apex; intervals slightly convex, with 2–3 rows of hairs; scutellar striole at base with a scutellar setiferous pore; three setiferous punctures in intervals 3; series umbilicata with 14 setiferous punctures. Microsculpture head with isodiametric meshes, pronotum and elytra with dense transverse meshes. Legs rather short and stout; tarsi with dorsal dense pubescence, ventral with hairbrush; protarsomeres 1–3 dilated (male), protarsomere 4 slightly bilobed. Ventral side colour underside yellow except the yellowish-red head; mentum with median tooth, slightly emarginate in the middle; sterna with pubescence; ratio length/width at anterior margin metepisterna: 1.67. Median lobe (Fig. 3.2) in lateral view comma-shaped, 1.44 mm long; the large erected hook at apex spatulashaped, directed at the apical lamella at an acute angle (seen laterally, in A. ruficollis the hook is located at an approximate right angle on the apical lamella, Fig.3.4). Comparative notes. Atranus dariushi sp. n. is compared with numerous specimens of A. ruficollis (Gautier des Cottes, 1858) from almost the entire range (Fig. 3.3). The new species differs from A. ruficollis by colour, the slenderer habitus, the pronotum less sinuate before hind angles, slightly weaker shoulders, reduced hind wings, metepisterna: ratio length/width at anterior margin (1.67) shorter as in A. ruficollis (2.11), and by the apex of the median lobe which is curved (as in P. ruficollis, Fig. 3.4) but the apex is spatula-shaped and the hook is located at an acute angle on the apical lamella (seen laterally). A. ruficollis is lighter or darker red-brown, often pronotum somewhat lighter, the pronotum and elytra a little shorter (PW/PL about 1.03-1.10, EL/EW 1.47-1.50, but measuring of more specimens of A. dariushi sp. n. in future should show the variability of these characters), pronotum as a rule somewhat more strongly sinuate before hind angles, shoulders less rounded, hind wings fully developed (distinctly longer than elytra, at the end folded twice, hence the species is capable of flight (some specimens from Greece were collected by car-catcher). Distribution. Iran (Kohgiluyeh and Boyer-Ahmad). Only known from the same small cave where Trechus kouroshi sp. n. was found. Habitat. This species was collected about 50 m from the entrance. Remarks: Atranus dariushi sp. n. is likely a cave-adapted beetle, lacking of functional hind wings and without pigmentation, and differs considerably in its ecology from the other species of this genus. Both A. ruficollis and the North American species A. pubescens Dejean, 1828 are epigean species from river banks, vegetal debris and floodplain forests (Larochelle & Larivière 2003, own observations) and are capable of flight. Etymology. The species epithet refers to Dariush, king of Persia, of the house of the Achaemenids. Identification reference. Arndt et al. (2011).Published as part of Muilwijk, Jan, Seiedy, Marjan & Wrase, David W., 2021, Contribution to the knowledge of Carabidae in Iran with the descriptions of five new taxa, proposing of four synonyms and providing of nine new records (Insecta: Coleoptera), pp. 55-81 in Zootaxa 5067 (1) on pages 61-62, DOI: 10.11646/zootaxa.5067.1.3, http://zenodo.org/record/565650
A remarkable new lebiine genus with two new species from Pakistan, Afghanistan and Iran (Coleoptera: Carabidae: Lebiini)
No full textKataev, Boris M., Muilwijk, Jan (2020): A remarkable new lebiine genus with two new species from Pakistan, Afghanistan and Iran (Coleoptera: Carabidae: Lebiini). Zootaxa 4763 (1): 73-84, DOI: https://doi.org/10.11646/zootaxa.4763.1.
Trechus kouroshi Muilwijk & Seiedy & Wrase 2021, sp. n.
No full textTrechus kouroshi sp. n. urn:lsid:zoobank.org:act: 83977913-FB4E-48E7-AC91-5C77E066A015 (Figs. 2.1, 2.2) Type material. Holotype, ♂ labeled: “ Iran, Kohgiluyeh and Boyer-Ahmad / Suq cave, 812 m / 30°49ˊ25.9´´ N, 50° 25ˊ37.9´´ E, 5.vi.2019 / S. Rahideh, Y. Fatemii & J. Muilwijk leg.” // “ Holotype / Trechus kouroshi sp. n. / Muilwijk, Seiedy & Wrase des. 2021” (black print on red label), deposited in HMIM. Paratype, 1 ♀ with same data but “ Paratype ”, in CJM. Comparing material: Trechus saulcyanus Csiki, 1928: 1 ♂, Israel, Upper Galilee Ya'ar Bar'am, 683 m, 23.iii.2011, 33°02´21´´ N, 35°2´26´´ E; Meybohm leg.; W. Ziegler det. 2011. CJM. Description. A small to middle sized and bicoloured Trechus species (3.5 mm) with piceous brown head and pronotum and yellowish elytra; appendages yellow; pronotum slightly shiny; micropterous (Fig. 2.1). Head eyes slightly convex, much longer than tempora (tempora/eye: 0.33–0.36). Frontal furrows moderately deep, evenly impressed, semi-circular. Clypeus with two pairs of setae, located in a slight deepening of the clypeus. Two pairs of supraorbital setae, the anterior setae at level of the middle of the eye, the posterior setae behind the eye. Antennae short (1.95 mm), reaching to one quarter of the elytra. Pronotum cordate (PW/PL: 1.28–1.37); slightly convex. Lateral sides of pronotum sinuate towards posterior angles, posterior angles distinct, nearly rectangular, apically upwards pointed. Anterior angles rounded. Posterior margin slightly emarginate laterally. Basal fovea deep. Middle line deep on disc, not reaching the anterior edge; lateral margin with 2 pairs of setae, one pair in the posterior angles, the other pair at the widest point. Elytra oblong-ovate (EL/EW: 1.3–1.4); humeri prominent, though rounded; maximum width around the middle; the marginal border reaching anteriorly interval 5. Three setiferous punctures in intervals, the anterior 2 pores situated along striae 3, the preapical pore connects stria 2 with 3; all striae punctated; striae 1–2 deep, ending at apical setiferous punctures, striae 3–4 superficial, towards apex shortened, striae 5–6 consists of a row of punctures, strongly shortened, stria 7 indistinct, consisting of a few punctures in the middle. Parascutellar striole short, superficially; parascutellar pore small surrounded through a keel formed by the thickened suture. The deep recurrent striole not reaching stria 5; series umbilicata with 8 setiferous punctures, 4 punctures at the base, 2 punctures just behind the middle, and 2 punctures apically. Microsculpture of head with isodiametric meshes, pronotum with more transverse meshes and elytra with superficial, transverse meshes. Legs slender; protarsomeres 1–2 dilated in male, each with a massive inner and obtuse outer tooth. Front tibiae grooved on exterior surface. Ventral side underside colour yellow except the dark meso- and metasternum and mes- and metepisternum. Median lobe (Fig. 2.2) small, 0.5 mm long, regularly bent, apex in vertical view asymmetric. Sclerites of endophallus in lateral view (l) triangular plate shaped, in dorsal view (d) and ventral view (v) arrow shaped, pointed to the apex. Left paramere with three setae, right paramere with four setae. 2.1 habitus holotype, scale: 2 mm. 2.2 median lobe holotype, l = lateral, v =ventral, d= dorsal view, scale: 0.5 mm. 2.3 Trechus saulcyanus, median lobe, l = lateral, v =ventral view, scale: 0.5 mm; Israel. Comparative notes. So far only three Trechus species are known from Zagros: T. liopleurus Chaudoir, 1850, T. melanocephalus Kolenati, 1845 and T. quadristriatus (Schrank, 1781) (Pawłowski, 1979). T. kouroshi sp. n. differs from these species by the shape of the pronotum, shape of median lobe and sclerites in the endophallus. T. quadristriatus and T. liopleurus have slightly convex lateral sides of the pronotum. T. melanocephalus has a dark brown dorsal and ventral side and elongated elytra. Probably Trechus kouroshi sp. n. is more related to T. saulcyanus Csiki, 1928 by the combination of punctated striae, small body size, cordiform pronotum and lamelliform sclerite in the median lobe (Fig. 2.3). However, in T. saulcyanus the sclerite has in horizontal view an arch shape and in lateral view a plate shape. The median lobe is more bent than in Trechus kouroshi sp. n. Distribution. Iran (Kohgiluyeh and Boyer-Ahmad). Only known from the collecting site, a small cave. Habitat. The cave is located in a gorge where a small brook flow through. The cave is approximately 100 m long, almost horizontal, with a streamlet between stones and small rocks. At the far end of the cave streams a larger brook, too narrow and too low to follow without diving equipment. There is hardly any debris or mud between the stones. The specimens were found not far from the cave entrance. The cave is surrounded by an extensively agricultural area with very high temperatures (more than 35 ° C) and very low humidity in summer. Etymology. The species’ epithet refers to Koerosh (Cyrus the Great), the founder of the Achaemenid Empire. Identification references: Morvan & Pawłowski (1977); Pawłowski (1979).Published as part of Muilwijk, Jan, Seiedy, Marjan & Wrase, David W., 2021, Contribution to the knowledge of Carabidae in Iran with the descriptions of five new taxa, proposing of four synonyms and providing of nine new records (Insecta: Coleoptera), pp. 55-81 in Zootaxa 5067 (1) on pages 59-61, DOI: 10.11646/zootaxa.5067.1.3, http://zenodo.org/record/565650
Leistus (Pogonophorus) spinibarbis subsp. zagrosensis Muilwijk & Seiedy & Wrase 2021, ssp. n.
No full textLeistus (Pogonophorus) spinibarbis zagrosensis ssp. n. urn:lsid:zoobank.org:act: AEF2F2E3-B739-4A34-8D34-36C96041C008 (Fig. 1.1, 1.2) Type material. Holotype ♂, labelled: “ Iran, Chahar Mahaal and Bakhtiari / Chery pass, 31.v.2019, 3260 m / Muilwijk leg. / 32°10ˊ51´´ N, 50°09ˊ59´´ E ” // “ Holotype / Leistus (Pogonophorus) spinibarbis / zagrosensis ssp. n. / Muilwijk, Seiedy & Wrase des. 2021” (black print on red label). Deposited in HMIM. Description. Upper surface of head and pronotum dark with bright blue-violet lustre, elytra more brownish with blue lustre; lateral margins of pronotum translucent reddish; propleura yellow, epipleura brown, appendages entirely yellow, underside dark except brownish apical sternites (Fig. 1.1). BL: 8.4 mm, EW: 3.3 mm. 1.1 habitus holotype, scale: 5 mm. 1.2 median lobe holotype, lateral view, scale: 1 mm. Head punctate, longitudinally superficial rugulose; eyes hemispherical; antennae long and slender. Pronotum transverse (PW/PL:1.37); punctuation on the base, lateral margins and near the apex strong, punctation on the disc rather fine; lateral furrow broad, sides narrowed to the rectangular basal angles. Middle line fine and only visible on disc. Elytra subparallel (EL/EW:1.7); weakly rounded laterally, their maximum width apical of middle; basal margin concave, humeral denticle small and sharp; striae rather deep, superficial apically, evidently punctate, intervals moderately convex, flattened near the apex with fine punctuation. Ventral side with dense and rather strong punctuation on prosternum, pro-, mes- and metepisterna, mesosternum, lateral sides of metasternum, lateral sides of abdominal sternites 1 and 2, lateral sides of sternite 3 with finer punctation, middle of sternite 3 and sternites 4–7 with fine scarce punctation; prosternal lobe pointed and not bordered at the tip with brown colour. Microsculpture superficially, isodiametric on head, and pronotal disk; elytra with less superficially, isodiametric meshes. Median lobe (Fig. 1.2) with a regularly concave apex. Comparative notes. L. (P.) spinibarbis zagrosensis ssp. n. differs from the other subspecies, Leistus (P.) spinibarbis rufipes Chaudoir 1843, L. (P.) spinibarbis ponticus Kryzhanovskij & Shilenkov, 1999 and L. (P.) spinibarbis abdominalis Reiche & Saulcy, 1855, in the Middle East by the concave apex of the median lobe, which is clearly asymmetrical in the other subspecies. In addition, L. (P.) spinibarbis rufipes differs from L. (P.) spinibarbis zagrosensis ssp. n. by shorter antennae; antennomere 3 shorter and thicker; more transverse pronotum; elytral sides more rounded; prosternal lobe apically rounded and completely bordered and black coloured. L. (P.) spinibarbis ponticus differs from L. (P.) spinibarbis zagrosensis ssp. n. by more coarsely and densely punctation of head, head and pronotum brownish black, elytra less parallel, widest point in the middle. L. (P.) spinibarbis abdominalis differs from L. (P.) spinibarbis zagrosensis ssp. n. by blue-green elytra, elytral striae more finely and densely punctured, the intervals completely flat and ventral side completely brown-red. Distribution. Iran (Chaharmahal and Bakhtiari). So far only known from collecting place. Habitat. The specimen was collected under a stone at high altitude, near a small snow field. Etymology. The name of the subspecies refers to the Zagros Mountains, where the specimen was collected.Published as part of Muilwijk, Jan, Seiedy, Marjan & Wrase, David W., 2021, Contribution to the knowledge of Carabidae in Iran with the descriptions of five new taxa, proposing of four synonyms and providing of nine new records (Insecta: Coleoptera), pp. 55-81 in Zootaxa 5067 (1) on pages 57-59, DOI: 10.11646/zootaxa.5067.1.3, http://zenodo.org/record/565650
Disciferella psammophila Kataev & Muilwijk 2020, sp. n.
No full textDisciferella psammophila sp. n. (Figs 15–17, 19, 21, 23, 26–29, 30 b) urn:lsid:zoobank.org:act: 7CC4567A-6590-4006-ACA2-E6646B512304 Type material. Holotype: ♂, “S. Iran, 400m / 6 km W Genu / 7.– 9.5.1977 // Loc. No. 323 Czechoslovak-Iranian entomological expedition” (MNP). Paratypes: 1 ♂, 7 ♀ “ Iran, Ziarat / 26 km NWN Bila’i / 14– 15.4.1977 // Loc. No. 330 / Czechoslovak-Iranian entomological expedition” (MNP, RMNH, HMIM, ZIN), 1 ♀, “ Iran Belutschistan / Iranshar 800m / 11.–21.IV.1954 / Richter u. Schäuffele” (SMNS); 1 ♂, 1♀, “SE Iran / Bahu-Kalat / 3– 4.4.1973 // Loc. No. 147 / Czechoslovak-Iranian entomological expedition” (MNP). Description. Habitus as in Figs 15 and 17. Body length 4.4–4.8 mm, width 2.3–2.6 mm. Characters as described for D. kabakovi sp. n. except as follows. Colour: Brownish yellow, mat, with elytral disc, particularly medio-basally, slightly darker than head and pronotum, with yellowish suture; apex and inner margin of mandibles blackish brown (one female completely black, with small yellow elytral border). FIGURES 18–23. Disciferella gen. n. 18, 20, 22, D. kabakovi sp. n.; 19, 21, 23, D. psammophila sp. n. 18, 19, head; 20, 21, pronotum, 22, 23, left metepisternum. Scale bar = 1.0 mm. Head (Fig. 19): Approximately as wide as pronotum (HWmax/PWmax 0.97–1.03 and HWmin/PWmax 0.60– 0.66). Eyes larger and more convex than those in D. kabakovi sp. n. Tempora convex, about half as long as eye. Labrum also with three pairs of setigerous pores, one setigerous pore at each apical angle and two pair near middle, with clear incision at middle of anterior margin. Antennae densely pubescent from apical half of antennomere 4; antennomeres 1–3 with scattered short setae. Pronotum (Fig. 21): Similar in shape to that of D. kabakovi sp. n. (PWmax/PL 1.17–1.25, PWmax/PWmin 1.44–1.62, and PWmax/PWbas 1.37–1.44). Sides sinuate at posterior fifth, bearing each, in addition to seta in basal angle, a strong seta approximately in widest point of pronotum (at the end of anterior fifth); apical angles each with a seta at apex; this seta shorter than anterior lateral seta and seta in basal angles but markedly longer than background pubescence. Lateral furrow reaching basal angle, narrow almost throughout, only slightly narrowed to apical angles. Dorsal microsculpture very distinct throughout, consisting of fine irregular meshes. Elytra: Similar in shape to that of D. kabakovi sp. n. (EL/EW 1.14–1.18, EL/PL 3.30–3.53, EW/PWmax 2.34– 2.54). Sutural angle nearly right or narrowly rounded (not related to sex). Discal setigerous pore absent. In addition to marginal umbilicate series consisting of 12–14 large setigerous pores, one pore located at apex of position of stria 3 and one pore at apex of stria 6. Microsculpture very distinct throughout, not finer than that on head or pronotum, consisting of granulate meshes. Ventral surface of thorax and abdomen: Prosternum throughout covered with partly slightly transversal and partly isodiametric microsculpture and more scarce setae than on proepisterna. Metepisternum (Fig. 23) longer than that of D. kabakovi sp. n., about twice as long as wide. Apex of last visible abdominal sternite in both sexes with one pair of marginal setae, concave in middle. Legs: Protibia with one apical spur (ca half of protarsomere 1). Two inner apical spurs of metatibia ca 0.3–0.4 as long as metatarsomere 1 (Fig. 16). Tarsi similar to those of D. kabakovi sp. n., but mesotarsomeres 3 and 4 slightly dilated in males. Female genitalia (Fig. 29): Laterotergite wider than long, with wide and deep emargination on mesal margin anteriorly; its apical membranous portion reduced; gonosubcoxite elongate, markedly longer than laterotergite, without setae and spines. Male genitalia (Figs 26–28): Median lobe of aedeagus in lateral view slightly less arcuate than in D. kabakovi sp. n., with apical portion less markedly bent ventrally. Internal sac with a small field of slightly longer spines on the right side medio-ventrally and with spiny patch medially. Etymology. The name of this new species refers to its assumed mode of life. Distribution (Fig. 30b). This species is known from south-eastern Iran (Hormozgan and Sistan va Baluchestan provinces). Bionomics. Most specimens of the type series of this new species were collected by the Czechoslovak-Iranian entomological expedition in 1973 and 1977. According to the description of the sampling localities (Hoberlandt, 1981, 1983), the species seems to be a psammophile. The habitats in “Ziarat” are sandy dunes in the vicinity of date-palms and trees; the habitats in “Bahu-Kalat” and in “ 6 km W of Genu” are respectively a gravel sandy river bed and a savanna vegetation formation. In all localities, material was collected both by hand and by light trap, but there is no information concerning the specimens of the new species. Comparative remarks. This new species is similar to D. kabakovi sp. n. in habitus and aedeagus, but can be easily distinguished from it, among other characters mentioned in the description, by using the following key: 1. Pronotum (Fig. 20) without anterior lateral seta at widest point. Each elytron in position of interval 3 with one very small discal setigerous pore located at the end of basal quarter. Head (Fig. 18) slightly narrower than pronotum, with smaller eyes. Metepisternum (Fig. 22) shorter, about 1.5 times as long as wide. Median lobe of aedeagus (Figs 13, 14, 24, 25) in lateral view slightly more arcuate, with apical portion more markedly bent ventrally; internal sac with a small, nearly transverse row of finer spines on the right side medially. Laterotergite (Fig. 10) about as long as wide, with shorter emargination on mesal margin anteriorly. Brownish yellow, with elytra slightly paler than head and pronotum. Larger: body length 4.7–5.2 mm. Afghanistan, Pakistan.............................................................................. Disciferella kabakovi sp. n. - Pronotum (Fig. 21) on each side with a strong anterior lateral seta at widest point in addition to setae in basal and apical angles. Elytron without discal pore in position of interval 3. Head (Fig. 19) wider, approximately as wide as pronotum, with larger eyes. Metepisternum (Fig. 23) longer, about twice as long as wide. Median lobe of aedeagus (Fig. 26) in lateral view slightly less arcuate, with apical portion less markedly bent ventrally; internal sac with a small field of slightly longer spines on the right side medio-ventrally. Laterotergite wider than long, with wider emargination on mesal margin anteriorly (Fig. 29). Usually darker, with elytral disc slightly darker than head and pronotum. Smaller: body length 4.4–4.8 mm. South-eastern Iran...................................................................................... Disciferella psammophila sp. n.Published as part of Kataev, Boris M. & Muilwijk, Jan, 2020, A remarkable new lebiine genus with two new species from Pakistan, Afghanistan and Iran (Coleoptera: Carabidae: Lebiini), pp. 73-84 in Zootaxa 4763 (1) on pages 79-81, DOI: 10.11646/zootaxa.4763.1.6, http://zenodo.org/record/374407
FIGURE 30. Disciferella gen. n., distribution. a, D in A remarkable new lebiine genus with two new species from Pakistan, Afghanistan and Iran (Coleoptera: Carabidae: Lebiini)
No full textFIGURE 30. Disciferella gen. n., distribution. a, D. kabakovi sp. n.; b, D. psammophila sp. n.Published as part of Kataev, Boris M. & Muilwijk, Jan, 2020, A remarkable new lebiine genus with two new species from Pakistan, Afghanistan and Iran (Coleoptera: Carabidae: Lebiini), pp. 73-84 in Zootaxa 4763 (1) on page 83, DOI: 10.11646/zootaxa.4763.1.6, http://zenodo.org/record/374407
Disciferella kabakovi Kataev & Muilwijk 2020, sp. n.
No full textDisciferella kabakovi sp. n. (Figs 1–14, 18, 20, 22, 24, 25, 30 a) urn:lsid:zoobank.org:act: DF46AD36-5BF0-48B7-8F0C-4D407FCD0E88 Type material. Holotype: ♂, “ Afghanistan / Jalalabad 600 m / 10.5.1971 Kabakov ” (ZIN). Paratypes: 1 ♀, Pakistan, “bridge over the Indus River / 70 km E Peshavar / 30.08.2005 ” (in Cyrillic) // ex coll. / S.V. Ovchinnikov / Donation to ZIN ” (ZIN). Description. Habitus as in Figs 1 and 2. Body length 4.7–5.2 mm, width 2.6–2.7 mm. Colour: Brownish yellow, mat, with elytra slightly paler than head and pronotum; apex and inner margin of mandibles as well as tibiae basally blackish brown; elytra with rows of indistinct small brown spots corresponding positionally to elytral striae; general fine setation of body yellow. Head (Fig. 18): Medium-sized, with moderately deep neck constriction, slightly narrower than pronotum, narrowest just behind tempora (HWmax/PWmax 0.95–0.96 and HWmin/PWmax 0.59–0.60); neck slightly widened posteriorly. Eyes large, convex, touching ventrally buccal fissure. Tempora long and very convex, about 0.8 times as long as eye. Labrum with three pairs of setigerous pores, flat, transverse; its anterior margin almost straight or with shallow incision at middle. Clypeus flat, sloped to labrum, transverse, with almost straight anterior margin and rounded posterior margin, with one setigerous pore at each apical angle. Fronto-clypeal suture distinct, not deepened. Frons somewhat flat, with very fine elongate wrinkles along inner margins of eyes. No frontal foveae, frontal and supraorbital furrows. Two pairs of very small, indistinct supraorbital setigerous pores: anterior pore situated at middle of upper margin of eye, and posterior one located behind eye, close to neck constriction. Mandibles elongate, similar to each other, very slightly widened at base and with distinct basal scrobe; their apices acute. Maxillae (Fig. 5) with ultimate palpomere slightly longer than penultimate one, fusiform. Lacinia with moderately dense row of straight slender setae along occlusal margin; tooth acute, long. Mentum (Fig. 4) with acute apical angles and with acute median tooth, completely fused with submentum; epilobes very narrow; two mental setigerous pores at apical margin laterally of median tooth. Ligular sclerite wide and short, with almost right, slightly prominent laterally apical angles and with two ventroapical setae. Paraglossae membranous, glabrous, wide, rounded at apex, longer than ligular sclerite and touching it by their inner ventral margin. Labial basal palpomere comparatively wide; penultimate palpomere slightly widened anteriorly, with one seta at anterior margin; ultimate palpomere approximately as long as penultimate one, fusiform, narrowed to truncate apex. Dorsal microsculpture very distinct throughout, consisting of large granulate meshes. Setigerous micropunctures very fine, indistinct, present both dorsally and ventrally; setae inclined anteriorly. Antennae long and slender, surpassing elytral base approximately by three apical antennomeres and densely pubescent from antennomere 4; antennomere 3 covered with sparser setae and antennomeres 1–2 with a few scattered short setae; antennomere 1 short, slightly longer than antennomere 2, with one longer dorsoapical seta; antennomeres 3–11 similar to each other in length, each only slightly longer than antennomeres 1+2; antennomeres 1–4 almost circular in cross section, other ones flattened, each with a narrow glabrous strip on outer margin. Pronotum (Fig. 20): Relatively small, cordate, slightly transverse (PWmax/PL 1.22–1.26), widest approximately at the end of anterior fifth, strongly narrowed posteriorly (PWmax/PWmin 1.42–1.48, PWmax/PWbas 1.37–1.44). Sides rounded almost along entire length, with very short sinuation just before very small and sharp basal angles; the latter removed ahead from rounded basal margin and bearing each a strong seta; lateral margins of pronotum behind basal angles very fine, not reaching the rounded basal margin, making basal portion of proepipleuron visible in dorsal view; visible margins of proepipleura behind basal angles slightly divergent posteriorly; lateral groove narrow, gradually widened posteriorly and vanishing just before basal angles; anterior lateral seta absent. Apical margin nearly straight, not bordered; apical angles rounded, not projecting ahead, each with a short seta at apex; this seta only slightly longer than background pubescence. Basal margin not bordered. Disc moderately convex, more strongly sloped to apical angles and latero-basally; base of pronotum behind basal angles slightly depressed. Anterior transverse depression long, arcuate, slightly deepened. Median line fine, present only in central part of pronotum. Surface impunctate, covered with very fine transverse wrinkles. Setigerous micropunctures very fine, indistinct, their setae inclined posteriorly. Dorsal microsculpture very distinct throughout, consisting of granulate meshes, slightly finer than on head. Scutellum small, with acute apex. Elytra: Discoidal, very wide, widest at middle (EL/EW 1.13–1.18, EL/PL 3.39–3.54, EW/PWmax 2.38–2.47) and slightly obliquely, almost transversely truncate at apex. Humeri widely rounded. Sutural angle nearly right, with apex sharp in male and very narrowly rounded in female. Basal border incomplete, very thin, curving along humerus up to mesothoracic peduncle. Disc moderately convex, with wide explanation along sides and with basal depression at pronotal base. Striae, including parascutellar striole, absent; at places of striae rows of indistinct slightly darker spots. Parascutellar setigerous pores present, small, removed from elytral base. Each elytron in position of interval 3 with one very small discal setigerous pore located at the end of basal quarter and with one larger preapical pore. Marginal umbilicate series removed rather far from elytral sides, continuous, consisting of 12–14 large setigerous pores; one pore located separately at outer apical angle. Setigerous micropunctures very fine, indistinct, their setae inclined posteriorly. Microsculpture visible throughout, somewhat similar to that on pronotum, consisting of very distinct isodiametric meshes. Epipleura markedly widened prebasally, distinctly bordered along inner margin. Wings fully developed. Ventral surface of thorax and abdomen: Proepisterna (propleura) impunctate, covered with very distinct isodiametric microsculpture and very short setae. Prosternum throughout covered with finer microsculpture and slightly longer setae; prosternal process not bordered and without long setae at apex. Metepisternum (Figs 9, 22) impunctate, about 1.5 times as long as wide, narrowed posteriorly. Three penultimate abdominal sternites (ventrites), in addition to fine background setae, each with a pair of long setae at posterior margin. Apex of last visible abdominal sternite with one pair of marginal setae, in male concave in middle (Fig. 6) and in female subtruncate (Fig. 7) [since only two specimens are available, these differences should be checked on the additional material]. Legs: Slender, comparatively long. Protibia with one slender apical spur (ca 0.5–0.6 as long as protarsomere 1). Inner apical spur of meso- and metatibia (Fig. 3) short (ca 0.4–0.5 as long as meso- and metatarsomere 1, respectively) and not serrate. Metafemur with one longer seta at middle of posterior margin. Tarsomeres slender, parallel-sided, finely and densely setose; tarsomere 4 truncate at apex; tarsal claws elongate, not dentate, not pectinate; metatarsomere 1 long, only slightly shorter than 2+3 (Fig. 8); male protarsomeres 1–4 slightly dilated and covered ventrally with sparse spongy adhesive vestiture. Female genitalia (Fig. 10): Laterotergite about as long as wide, asymmetrical, with deep emargination on mesal margin anteriorly, widely membranous apically, asetose; gonosubcoxite elongate, as long as laterotergite, without setae and spines; gonocoxite short and wide, with a sensory pit just after middle of inner margin. Male genitalia: Median lobe of aedeagus (Figs 13, 14) slender, with relatively small basal bulb, in lateral view arcuate, in dorsal view almost straight, with terminal lamella short, narrowed apically; apical orifice shifted to left, not clearly delimited proximally. Internal sac medially with oval spiny patch and with a small, nearly transverse row of fine spines on the right side (Fig. 24, 25). Parameres, especially the right one, very narrow, both rounded at apices (Figs 11, 12). Etymology. Named after the geologist and coleopterologist Oleg N. Kabakov (1928–2009), an excellent specialist on Scarabaeidae, who first collected this species. Distribution (Fig. 30a). This species is known from Afghanistan and northern Pakistan. Bionomics. Unknown, but based on the body shape, the species seems to be a psammophile like D. psammophila sp. n., species of the genus Discoptera and some species of the genus Graphipterus having wide discoidal elytra (e. g. the species of the G. serrator group).Published as part of Kataev, Boris M. & Muilwijk, Jan, 2020, A remarkable new lebiine genus with two new species from Pakistan, Afghanistan and Iran (Coleoptera: Carabidae: Lebiini), pp. 73-84 in Zootaxa 4763 (1) on pages 76-79, DOI: 10.11646/zootaxa.4763.1.6, http://zenodo.org/record/374407
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