1,734,873 research outputs found
Polystomophora arakensis Moghaddam
No full textPolystomophora arakensis Moghaddam (Fig. 38) Polystomophora arakensis Moghaddam: Moghaddam & Alikhani, 2010: 12–14. Holotype adult Ƥ, Iran, Arak. DIAGNOSIS. Mounted female broadly oval. Antennae 9 segmented. Legs moderately developed; claw with a denticle. Translucent pores present on hind femur and tibia. Cerarii absent except anal lobe cerarii; each with 2 long slender setae plus a few trilocular pores. Anal ring rounded, located at some distance from apex. Circulus present, notched on each side. Both pairs of ostioles well developed. Setae on dorsal and ventral surface all flagellate. Dorsal multilocular disc pores scattered across head, thorax, abdominal segments I–IV, and on margin and midline of abdominal segments V–VII; also present on venter, across midline of thorax and all abdominal segments, and posterior to vulva. Ventral quinquelocular disc pores present on midline of thoracic segments and anterior abdominal segments. Trilocular disc pores evenly distributed on dorsum and venter. Oral collar ducts sparse on both dorsum and venter. DISTRIBUTION. Palaearctic: Iran. In Iran, P. arakensis occurs in Markazi. MATERIAL EXAMINED: Markazi: Arak, 15 adult Ƥ, on Atraphaxis sp. (Polygonaceae), 1726 m., 10.x. 2009 (M. Alikhani).Published as part of Moghaddam, Masumeh, 2013, A review of the mealybugs (Hemiptera: Coccoidea: Pseudococcidae, Putoidae and Rhizoecidae) of Iran, with descriptions of four new species and three new records for the Iranian fauna, pp. 1-107 in Zootaxa 3632 (1) on page 68, DOI: 10.11646/zootaxa.3632.1.1, http://zenodo.org/record/21761
Chorizococcus pazoukii Moghaddam
No full textChorizococcus pazoukii Moghaddam (Fig. 8, distribution map Fig. 66 G) Chorizococcus pazoukii Moghaddam in Moghaddam & Alikhani, 2014. Field characteristics: Not recorded. Microscopic diagnosis: Slide-mounted adult female oval to broadly oval. Anal lobes poorly developed. Antennae each with 8 segments. Legs well developed, small for size of body. Hind leg with large translucent pores on dorsal surface of tibia; tarsal digitules pointed at tip and shorter than claw. Cerarii numbering a single pair situated on anal lobes, each cerarius with 2 large conical setae, 3 or 4 auxiliary setae and a group of trilocular pores, all situated on a slightly sclerotized area. Circulus divided by an intersegmental line. Ostioles well developed. Anal ring with 2 rows of pores and bearing 6 setae. Dorsum with setae short and stiff. Multilocular disc pores absent. Trilocular and discoidal pores evenly distributed. Oral rim tubular ducts of 2 sizes: larger type numerous in bands over entire dorsum, except for abdominal segment VIII; and smaller type present across posterior abdominal segments. Oral collar tubular ducts few, present on margins of abdominal segments IV ‒ VIII. Venter with normal flagellate setae. Multilocular disc pores each with 12 loculi, present medially in more or less double rows at posterior edges of abdominal segments VI and VII, and posterior to vulva. Trilocular pores and discoidal pores present. Oral rim tubular ducts of 2 sizes: larger type, similar to those on dorsum, present over body except for posterior to vulva; smaller type present in marginal and submarginal zones. Oral collar tubular ducts, same as those on dorsum, present mainly around lateral margins of posterior abdominal segments and at posterior edges of abdominal segments III ‒ VIII. Distribution: This Iranian species has been recorded from Markazi province (Moghaddam & Alikhani 2014). Host-plants: Chorizococcus pazoukii was found on the roots of an unknown plant (Moghaddam & Alikhani 2014). Economic importance: None. Natural enemies: None recorded. Comments: Chorizococcus pazoukii is similar to C. neomexicanus (Tinsley) in having numerous oral rim tubular ducts on the dorsum, but differs by lacking oral rim tubular ducts in medial areas of the venter. The accompanying illustration of C. pazoukii is reproduced from Moghaddam & Alikhani (2014), page 57, Fig. 1, with kind permission from the Editor of Journal of the Entomological Society of Iran.Published as part of MOGHADDAM, MASUMEH & WATSON, GILLIAN W., 2022, The Scale Insects Of Iran (Hemiptera: Coccomorpha) Part 2 The Mealybugs (Pseudococcidae And Rhizoecidae) And Putoidae, pp. 1-169 in Zootaxa 5126 (1) on pages 34-35, DOI: 10.11646/zootaxa.5126.1.1, http://zenodo.org/record/646041
Coccidohystrix burumandi Moghaddam
No full textCoccidohystrix burumandi Moghaddam (Fig. 8) Coccidohystrix burumandi Moghaddam: Moghaddam & Alikhani, 2009: 175–179. Holotype adult Ƥ, Iran, Arak (HMIM) DIAGNOSIS. Body of adult female broadly oval. Antennae 9 segmented. Legs well developed. Translucent pores few on hind tibia and tarsus; claw with a denticle. Cerarii numbering 17 pairs, each situated on a sclerotized plate. Anal lobe cerarii each comprising 4 or 5 blunt conical setae, 2 or 3 very small blunt setae, and 1 or 2 small discoidal pores. Anterior cerarii each with 6–8 conical setae, 2 or 3 small blunt setae and 1 or 2 small discoidal pores. Dorsal cerarii similar to lateral cerarii, but smaller, present on midline from head to abdominal segment VII; submedial series of dorsal cerarii situated from head to abdominal segment VI, and intermediate cerarii occurring between medial and submedial cerarii on mesothorax and abdominal segments I–IV. Circulus and ostioles absent. Dorsal surface with minute and sparse lanceolate setae; venter with slender flagellate setae. Multilocular disc pores absent from dorsum, but present ventrally posterior to vulva and across abdominal segments V–VII. With a compact group of more than 100 small trilocular pores and 2–5 quinquelocular pores adjacent to all spiracular openings. Trilocular pores evenly scattered. Discoidal pores sparse. Oral collar ducts numbering 4, present across abdominal segment VI. DISTRIBUTION. Iranian. In Iran, C. burumandi occurs in Markazi. MATERIAL EXAMINED. Markazi: Arak, 10 adult Ƥ, on Euphorbia sp. (Euphorbiaceae), 25.v. 2009 (M. Alikhani).Published as part of Moghaddam, Masumeh, 2013, A review of the mealybugs (Hemiptera: Coccoidea: Pseudococcidae, Putoidae and Rhizoecidae) of Iran, with descriptions of four new species and three new records for the Iranian fauna, pp. 1-107 in Zootaxa 3632 (1) on pages 18-20, DOI: 10.11646/zootaxa.3632.1.1, http://zenodo.org/record/21761
Spilococcus mirzayansi Moghaddam
No full textSpilococcus mirzayansi (Moghaddam) (Fig. 47) Chorizococcus mirzayansi Moghaddam, 2010 a: 64. Spilococcus mirzayansi (Moghaddam), Tsai & Wu, 2011: 29-30. Holotype Ƥ, Iran, Tehran: Tehran (HMIM). DIAGNOSIS. Adult female broadly oval. Antennae 8 segmented. Translucent pores large, present on hind coxa and tibia; claw with a small denticle. Cerarii numbering 8 pairs, all on abdominal segments; anal lobe cerarii each with 2 small conical setae, plus 3 or 4 auxiliary setae and a group of trilocular pores. Anterior cerarii, each with 2 conical setae but without auxiliary setae, plus a small concentration of trilocular pores. Circulus absent. Ostioles well developed. Dorsal surface with short, conical setae. Ventral surface with long flagellate setae, and slender conical setae near margins. Multilocular disc pores absent from dorsum; ventrally present posterior to vulva, and in 2 rows across medial posterior edges of abdominal segments VI and VII, and in a single row on posterior edges of abdominal segments III–V; also along anterior edges of abdominal segments V–VII, reaching to margins; a few multilocular disc pores also present in submedial area of abdominal segment II. Trilocular pores present on dorsum and venter. Discoidal pores minute, scattered on dorsum and venter. Dorsal oral rim ducts arranged in a single row across abdominal segments, except segment VIII, plus a few scattered over thorax and head. Oral rim ducts on venter, similar to those on dorsum, present in small numbers along margins of abdominal segments, and across head and thoracic segments. Dorsal oral collar ducts present in marginal groups on abdominal segments II–VII. Ventral oral collar ducts occurring mainly in groups in lateral areas of abdominal segments III–VIII and across segments III–VII; also present in a small group lateral to each first coxa. DISTRIBUTION. Palaearctic: Iran. In Iran, S. mirzayansi occurs in Tehran. MATERIAL EXAMINED. Tehran, Tehran, 3 adult Ƥ, on Opuntia ficus – indica (Cactaceae), 18.ii. 2002 (F. Parsi). COMMENTS. Tsai & Wu (2011) transferred Ch. mirzayansi to the genus Spilococcus based on the presence of more than 5 pairs of cerarii, a major diagnostic character of Spilococcus, distinguishing it from Chorizococcus, based on the definition of Williams’s (2004). The accompanying illustration is taken from Moghaddam (2010 a).Published as part of Moghaddam, Masumeh, 2013, A review of the mealybugs (Hemiptera: Coccoidea: Pseudococcidae, Putoidae and Rhizoecidae) of Iran, with descriptions of four new species and three new records for the Iranian fauna, pp. 1-107 in Zootaxa 3632 (1) on pages 83-86, DOI: 10.11646/zootaxa.3632.1.1, http://zenodo.org/record/21761
Dysmicoccus caspianensis Moghaddam
No full textDysmicoccus caspianensis Moghaddam (Fig. 15, distribution map Fig. 67 B) Dysmicoccus caspiansis Moghaddam, 2018: 181. Field characteristics: Not recorded. Microscopic diagnosis: Slide-mounted adult female elongate oval. Antennae each with 8 segments. Legs well developed; hind leg with translucent pores on ventral and dorsal surfaces of femur and tibia. Claw moderately stout, without a denticle; claw digitules knobbed; tarsal digitules each pointed at tip and longer than claw. Cerarii numbering 7 pairs, all situated on abdominal segments; anal lobe cerarii each containing 2 enlarged conical setae, 4 or 5 auxiliary setae, and a group of trilocular pores, all situated on a lightly sclerotized area. Circulus absent. Ostioles well developed. Anal ring situated at apex of abdomen, with 2 rows of pores and 6 setae. Dorsum with long, slender flagellate setae. Trilocular pores present in a regular arrangement. Discoidal pores minute, scattered. Oral collar tubular ducts of 2 sizes, forming groups of 3, groups distributed mainly across dorsal segments; each group usually comprising 2 large ducts on either side of 1 small duct; single ducts of either size present also. Venter with most setae similar to dorsal setae but usually slightly longer. Multilocular pores each with 10 loculi, present in more-or-less single rows at posterior edges of abdominal segments V‒VII, not reaching to margins; present also posterior to vulva. Trilocular pores evenly distributed. Discoidal pores same as those on dorsum, scattered. Oral collar tubular duct groups same as those on dorsum, present mainly around margins; small ducts present across median areas of abdomen, thorax, and submedian areas of head. Distribution: This Iranian species has been recorded from Qazvin province (Moghaddam 2018). Host-plants: Dysmicoccus caspianensis has been recorded on Poaceae (Moghaddam 2018). Economic importance: None. Natural enemies: Not recorded. Comments: The accompanying illustration is reproduced from Moghaddam (2018), page 182, Fig. 1, with kind permission from the Chief Editor of Zootaxa.Published as part of MOGHADDAM, MASUMEH & WATSON, GILLIAN W., 2022, The Scale Insects Of Iran (Hemiptera: Coccomorpha) Part 2 The Mealybugs (Pseudococcidae And Rhizoecidae) And Putoidae, pp. 1-169 in Zootaxa 5126 (1) on page 48, DOI: 10.11646/zootaxa.5126.1.1, http://zenodo.org/record/646041
Phenacoccus salviacus Moghaddam
No full textPhenacoccus salviacus Moghaddam (Fig. 32) Phenacoccus salviacus Moghaddam, 2010, in Moghaddam & Alikhani, 2010. Holotype adult Ƥ, Iran, Arak, Shazand-Suraneh. DIAGNOSIS. Body of female oval. Antennae normally 9 segmented. Legs well developed; claw with a small denticle; without translucent pores on hind legs. Cerarii numbering 12–14 pairs. Anal lobe cerarii each with 2 enlarged lanceolate setae, plus 4–6 trilocular pores and 2 small setae, all situated on a weakly sclerotized area. Anterior cerarii each bearing 2 short setae and 1 or 2 trilocular pores. A dorsal median cerarius present on segment VII with 2 setae and 2 trilocular pores at base. Circulus present, divided by an intersegmental line. Ostioles well developed. Some dorsal setae each with 1 trilocular pore at base, present across all segments. Dorsal multilocular disc pores few on posterior edge of abdominal segment VII; ventrally on posterior edges of abdominal segments IV–VII, on anterior edges of segment VII, and posterior to vulva; absent from margins. Quinquelocular pores present on midline of head, thorax and anterior abdominal segments. Dorsal oral collar ducts of 1 size, across head, thorax, and abdominal segments I–VII; and ventrally with a few on margins of meso- and metathorax, and across all abdominal segments. DISTRIBUTION. Palaearctic: Iran. In Iran, P. s a lv i ac u s has only been recorded from Markazi. MATERIAL EXAMINED. Markazi: Arak, Shazand, Suraneh, 19 adult Ƥ, on Salvia bracteata (Lamiaceae), 6.iv. 2009 (M. Alikhani).Published as part of Moghaddam, Masumeh, 2013, A review of the mealybugs (Hemiptera: Coccoidea: Pseudococcidae, Putoidae and Rhizoecidae) of Iran, with descriptions of four new species and three new records for the Iranian fauna, pp. 1-107 in Zootaxa 3632 (1) on page 60, DOI: 10.11646/zootaxa.3632.1.1, http://zenodo.org/record/21761
Rhodania aeluropi Williams & Moghaddam
No full textRhodania aeluropi Williams & Moghaddam (Fig. 43) Rhodania aeluropi Williams & Moghaddam, 2007: 38. Holotype Ƥ, Khouzestan: Ahvaz, Amirabad (HMIM). DIAGNOSIS. Body of adult mounted female rotund, membranous. Anal lobes membranous, barely perceptible, each with an apical seta. Antennae 6 segmented. Legs well developed, small for size of body. Claw long, without a denticle; claw digitules pointed, surpassing claw in length. Tarsal digitules shorter than claw digitules, pointed. Fairly large translucent pores present on anterior surface of hind coxa only. Anal ring narrowly rimmed, without cells, situated either at apex or up to a distance of about twice its length from apex of abdomen; with 6 short slender setae. Cerarii represented by a pair of long flagellate setae on each anal lobe only, plus a similar seta a short distance anteriorly. Circulus absent. Ostioles absent. Dorsal surfaces with short, flagellate setae, except some marginal setae, each 15–20 μm long, on posterior abdominal segments. Disc pores present, conspicuous, with a distinct quinquelocular centre and with about 10 outer loculi usually with indistinct walls; numerous, distributed mostly in rows across anterior and posterior edges of abdominal segments, becoming more scattered on head and thorax. Dorsal oral collar ducts with membranous collar occupying about one-third length of duct; inner end of duct heavily sclerotized; not numerous, most frequent on midline, usually on abdominal segments III–VI, and around margins, and with only a few present in submarginal and submedial areas. Ventral surface with setae similar to those on dorsum. Disc pores as on dorsum also with a similar distribution but frequent and more scattered on head and thorax. Ventral oral collar ducts as on dorsum, present around margins and with a few present across middle of posterior abdominal segments. DISTRIBUTION. Palaearctic: Iran. In Iran, R. aeluropi occurs in Khouzestan. MATERIAL EXAMINED. Khouzestan: Ahvaz, Amirabad, 2 adult Ƥ, on Aeloropus sp. (Poaceae), 18.x. 2005 (A. Narrehei). COMMENTS. R. aeluropi was collected in sugarcane fields (Williams & Moghaddam, 2007). The accompanying illustration is reproduced from Williams & Moghaddam (2007).Published as part of Moghaddam, Masumeh, 2013, A review of the mealybugs (Hemiptera: Coccoidea: Pseudococcidae, Putoidae and Rhizoecidae) of Iran, with descriptions of four new species and three new records for the Iranian fauna, pp. 1-107 in Zootaxa 3632 (1) on pages 77-79, DOI: 10.11646/zootaxa.3632.1.1, http://zenodo.org/record/21761
Phenacoccus bromi Moghaddam
No full textPhenacoccus bromi Moghaddam (Fig. 38, Plate 2 B, distribution map Fig. 69 A) Phenacoccus bromi Moghaddam, 2018: 185. Field characteristics: Body broadly oval, greyish, covered with white powdery wax, with 1 pair of short waxy filaments at posterior end of body; body segmentation apparent. Microscopic diagnosis: Slide-mounted adult female oval to broadly oval. Anal lobes moderately developed. Antennae each with 9 segments. Legs well developed; tarsal and claw digitules knobbed; translucent pores difficult to observe, present on hind tibia; claw with a distinct denticle; tarsal digitules pointed at tip and shorter than claw. Cerarii numbering 18 pairs; anal lobe cerarii (C 18) each with 5 lanceolate setae, 4 or 5 spine-like auxiliary setae and 12‒18 trilocular pores, all situated on a sclerotized area; penultimate cerarii (C 17) each with 3 or 4 lanceolate setae and 4 or 5 trilocular pores; anterior cerarii each with 2 lanceolate setae and 2 trilocular pores, except for frontal and ocular cerarii (C 1 and C 3) each with 3 setae. Circulus oval, sclerotized, situated between abdominal segments III and IV, divided by an intersegmental line. Ostioles well developed. Anal ring with 2 rows pores and 6 setae. Dorsum with lanceolate setae, some with a trilocular pore situated near setal collar; some minute setae present. Trilocular pores and minute discoidal pores scattered throughout. Oral collar tubular ducts and multilocular disc pores absent. Venter with normal flagellate setae, except for some short lanceolate setae present on margins. Multilocular disc pores and quinquelocular pores absent. Trilocular pores and minute discoidal pores scattered throughout. Oral collar tubular ducts few, numbering 1‒3 on abdominal segments VI‒VIII. Distribution: Phenacoccus bromi is known only from Iran, Markazi province (Moghaddam 2018). Host-plants: The species has been recorded on Bromus sp. (Poaceae) (Moghaddam 2018). Economic importance: None. Natural enemies: None recorded. Comments: Phenacoccus bromi is an unusual species of Phenacoccus in that it lacks multilocular and quinquelocular pores, and has tarsal digitules that are capitate. The accompanying illustration is reproduced from Moghaddam (2018), page 186, Fig. 4, with kind permission from the Chief Editor of Zootaxa.Published as part of MOGHADDAM, MASUMEH & WATSON, GILLIAN W., 2022, The Scale Insects Of Iran (Hemiptera: Coccomorpha) Part 2 The Mealybugs (Pseudococcidae And Rhizoecidae) And Putoidae, pp. 1-169 in Zootaxa 5126 (1) on pages 91-92, DOI: 10.11646/zootaxa.5126.1.1, http://zenodo.org/record/646041
Phenacoccus betae Moghaddam
No full textPhenacoccus betae Moghaddam (Fig. 26) Phenacoccus betae Moghaddam, 2010 a: 65 –67. Holotype adult Ƥ, Iran, Kermanshah (HMIM). DIAGNOSIS. Body of female broadly oval. Antennae normally 9 segmented. Legs well developed, translucent pores small and difficult to observe, present on hind femur and tibia; claws with a small but distinct denticle. Cerarii numbering 7 pairs, 4 on head (C 1 –C 4), and 3 on last 3 abdominal segments (C 16 –C 18). Anal lobe cerarii each with a pair of lanceolate setae, and 4 or 5 trilocular pores. Anterior cerarii with 2 small lanceolate setae and 2 or 3 trilocular pores. Preocular cerarii (C 3) with 3 lanceolate setae. Circulus absent. Ostioles well developed. Dorsal multilocular disc pores present across posterior edges of abdominal segments V −VII, and ventrally in 2 or 3 rows across posterior edges of abdominal segments IV −VII, on anterior edge of segment VII, on margin of abdominal segment III, and posterior to vulva. Quinquelocular pores few around mouthparts. Trilocular pores present, evenly dispersed. Discoidal pores minute, scattered. Dorsal oral collar ducts of 1 size, scattered on head, thorax and abdominal segments, except segment VIII. Ventral ducts of 2 sizes: large duct as on dorsum, distributed on head, thorax and margins of abdominal segments, and small ducts present medially on posterior abdominal segments. DISTRIBUTION. Palaearctic: Iranian species, P. betulae occurs in Kermanshah and Markazi. MATERIAL EXAMINED. Kermanshah: Kermanshah, 4 adult Ƥ, on Beta vulgaris (Betulaceae), May, 1973. Markazi: Arak, Toureh, 2 adult Ƥ, on Amaranthus blitoides (Amaranthaceae), 22.vi. 2010, 1850 m. (M. Alikhani). COMMENTS. P. b et ae is similar to P. p u m i l u s in having: (i) dorsal multilocular disc pores present on posterior abdominal segments, (ii) circulus absent, (iii) and cerarii present on head. P. b e t a e differs from P. pumilus in having (character states on P. pumilus in brackets): (i) quinquelocular pores present on venter around mouthparts (quinquelocular pores absent from venter).Published as part of Moghaddam, Masumeh, 2013, A review of the mealybugs (Hemiptera: Coccoidea: Pseudococcidae, Putoidae and Rhizoecidae) of Iran, with descriptions of four new species and three new records for the Iranian fauna, pp. 1-107 in Zootaxa 3632 (1) on page 49, DOI: 10.11646/zootaxa.3632.1.1, http://zenodo.org/record/21761
Hemiberlesia uramanica Moghaddam 2021, sp. n.
No full textHemiberlesia uramanica Moghaddam sp. n. (Fig. 3) Field characters: Adult female appearance in life is not recorded. Material examined. Holotype, adult female, IRAN, left label: 1739 / Kurdistan / Sanandaj / Shoysheh / 3.vii.2007 / Alt. 1437 m. / N35˚19′26.5″ / E46˚35′46.1″; right label: Holotype / Hemiberlesia uramanica Moghaddam sp. n. on Cotoneaster vulgaris / (Rosaceae) / coll. M. Moghaddam (HMIM). Paratypes: IRAN, 2 adult females together on 1 slide, with same data on left label as holotype slide; right label: Paratype / Hemiberlesia uramanica / Moghaddam sp. n. (HMIM). Slide-mounted adult female (Fig. 3), n = 3. Body pyriform to circular (0.72–0.92 mm long and 0.64– 0.60 mm wide). Cuticle membranous, except for parts of pygidium. Antennae each with 1 long seta. Anterior and posterior spiracles each without associated disc pores. Pygidium with 3 pairs of lobes; median lobes (L 1) well developed, each almost rectangular, with 1 or 2 notches at apex. Second lobes (L 2) smaller than L 1, dentate at apex. Third lobes (L 3) reduced, each represented by a sclerotized prominence. Plates distinct, dentate at apex, never longer than a median lobe; plates numbering 2 between median lobes, 2 between L 1 & L 2 and 3 between L 2 & L 3; absent from margins lateral to third lobes.Anal opening located 0.82‒0.88 (0.88) times length of anal opening from base of median lobes, anal opening 17‒18 (18) µm long. Perivulvar pores absent. Dorsum. Macroducts 1-barred, each relatively long and slender; 2 or 3 macroducts present between median lobes; 3 or 4 between L 1 & L 2; 3 or 4 between L 2 & L 3; and present in oblique rows on segment V. Ducts on prepygidial segments smaller than pygidial ducts, present on submarginal and submedian areas of abdominal segments III and IV; also on submarginal areas of segments I and II, and singly on each side of metathorax. Paraphyses small, present on inner edges of L 1, L 2 and L 3. Venter. Microducts present mainly near margins of pygidium, and near margins and in submedian areas of prepygidial segments; microducts also present on medial areas and near margins of meso- and metathorax. Sclerotized apophyses distinctly present on pygidium. Perivulvar pores absent. Comments. The adult female of H. uramanica sp. n. is similar to that of H. rapax (Comstock) in having: (i) no perivulvar pores, and (ii) antennae each with a single seta, but differs in having (character-states for H. rapax in brackets) (i) 2 or 3 median ducts between median lobes (ducts between median lobes absent); and (ii) numerous dorsal submarginal and submedian macroducts, present in distinct groups on abdominal segments III and II (segments III and II with sparse ducts, scattered on submargins). Etymology. The name is based on the locality name, Uraman (a famous area in Kurdistan province), and the Latin suffix ‘ -ica ’, denoting origin.Published as part of Moghaddam, Masumeh, 2021, Three new species of armoured scale insect (Hemiptera: Coccomorpha Diaspididae) from Iran, pp. 371-379 in Zootaxa 4991 (2) on pages 376-378, DOI: 10.11646/zootaxa.4991.2.10, http://zenodo.org/record/503037
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