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    Eugnosta brownana Metzler & Forbes, 2012, sp. nov.

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    Eugnosta brownana, sp. nov., Metzler & Forbes (Figs 1, 2, 7, 10, 13, 22) Holotype. Adult male, pinned with labels as follows: “ USA: NM: Otero Co. White Sands Nat[ional] Mon[ument] interdune habitat 106 ° 10.84 ’ W, 32 ° 46.64 ’ N, 4,008 ' 17 May 2010 WsnmF Eric H. Metzler uv tr[a]p Accessn #: WHSA - 00131" “ HOLOTYPE USNM Eugnosta brownana Metzler & Forbes ” (USNM). Paratypes. 88 3, 39 Ƥ: USA: ARIZONA: Cochise Co: Bisbee 1600m 26.viii- 6.ix. 1998 V.O.Becker Col Col. BECKER 115297 (13). Portal, 4 mi. W. 5300 ’, Chiricahua Mts., Aug. 3 -6, 1964, D.R.Davis. (43, 1Ƥ). Carr Cyn., 5300 ’ Huachuca Mts., VII- 31-66, bl. Wagner & Powell collrs. (13). Paradise, Ariz, July 24-31 (1 Ƥ). NEW MEXICO: Eddy Co.: Carlsbad Caverns NP (hereafter CCNP), grassland habitat, 32 °06.222’N 4160 ’ 104 ° 33.759 ’W, 14 June 2007, Eric H. Metzler CCNP 1, Accsn #: CAVE-02263 (83, 4Ƥ). CCNP, Arroyo habitat, 32 ° 5.976 ’ N 4,100ft. 104 ° 33.569 ’W, 14 June 2007, Eric H. Metzler CCNP 2, Accsn #: CAVE-02263 (1 Ƥ). CCNP, Arroyo habitat, 32 ° 5.976 ’ N 4,100ft. 104 ° 33.569 ’W, 29 August 2006, Eric H. Metzler CCNP 2, Accsn #: CAVE- 0 2263 (1 Ƥ). CCNP Riparian habitat 32 °06.566’N 3638 ’ 104 ° 28.257 ’W, 14 June 2007 Eric H. Metzler CCNP 4 Accsn #: CAVE-02263 (1 Ƥ). Lincoln Co.: Valley of Fires R[ecreation]A[rea] 4 mi NW Carrizozo 19 Aug 2005 5250 ' leg. D. J. Wright (1 Ƥ). Otero Co.: White Sands National Monument (hereafter WSNM) interdune habitat 106 ° 10.84 ’ W 32 ° 46.64 ’ N 4,008 ' 17 May 2010 WsnmF Eric H. Metzler uv trp Accessn #: WHSA – 0 0 131 (83, 1Ƥ). WSNM interdune habitat 106 ° 10.84 ’ W 32 ° 46.64 ’ N 4,008 ' 17 Oct 2009 WsnmF Eric H. Metzler uv trp Accessn #: WHSA – 0 0 131 (13). WSNM interdune habitat 106 ° 10.84 ’ W 32 ° 46.64 ’ N 4,008 ' 10 Aug 2010 WsnmF Eric H. Metzler uv trp Accessn #: WHSA—00131 (13). WSNM interdune habitat 106 ° 10.84 ’ W 32 ° 46.64 ’ N 4,008 ' 20 June 2009 WsnmF Eric H. Metzler uv trp Accessn #: WHSA – 0 0 131 (13). WSNM dunes crest vegetat. 106 ° 11.42 ’W 32 ° 45.67 ’ N 4,014 ’ 3 June 2008 WSNMC Eric H. Metzler uv trp Accss # WHSA 0 0 131 (1 Ƥ). WSNM dunes crest vegetat. 106 ° 11.42 ’W 32 ° 45.67 ’ N 4,014 ’ 2 Aug. 2010 WSNMC Eric H. Metzler uv trp Accss # WHSA 0 0 131 (33). WSNM interdune vegetation 106 ° 11.38 ’W 32 ° 46.69 ’ N 4,000 ’ 10 Aug. 2010 WSNM 8 Eric H. Metzler uv trp Accss # WHSA 0 0 131 (13). WSNM interdune vegetation 106 ° 11.38 ’W 32 ° 46.69 ’ N 4,000 ’ 14 July 2010 WSNM 8 Eric H. Metzler uv trp Accss # WHSA 0 0 131 (13). WSNM interdune vegetation 106 ° 11.38 ’W 32 ° 46.69 ’ N 4,000 ’ 6 April 2008 WSNM 8 Eric H. Metzler uv trp Accss # WHSA 0 0 131 (73). WSNM interdune vegetation 106 ° 11.49 ’W 32 ° 45.60 ’ N 4,000 ’ 6 April 2008 WSNMB Eric H. Metzler uv trp Accss # WHSA 0 0 131 (93, 1Ƥ). WSNM interdune vegetation 106 ° 11.49 ’W 32 ° 45.60 ’ N 4,000 ’ 14 Sept 2009 WSNMB Eric H. Metzler uv trp Accss # WHSA 0 0 131 (13). WSNM interdune vegetation 106 ° 11.49 ’W 32 ° 45.60 ’ N 4,000 ’ 25 Aug 2009 WSNMB Eric H. Metzler uv trp Accss # WHSA 0 0 131 (13). WSNM interdune vegetation 106 ° 11.49 ’W 32 ° 45.60 ’ N 4,000 ’ 23 April 2009 WSNMB Eric H. Metzler uv trp Accss # WHSA 0 0 131 (13). WSNM edge of dunes habitat 106 ° 11.32 ’W 32 ° 45.72 ’ N 4,000 ’ 6 April 2008 WSNMB Eric H. Metzler uv trp Accss # WHSA 0 0 131 (93, 1Ƥ). WSNM edge of dunes habitat 106 ° 11.32 ’W 32 ° 45.72 ’ N 4,000 ’ 17 May 2010 WsnmB Eric H. Metzler uv trp Accss # WHSA 0 0 131 (13). WSNM edge of dunes habitat 106 ° 11.32 ’W 32 ° 45.72 ’ N 4,000 ’ 25 Aug 2009 WsnmB Eric H. Metzler uv trp Accss # WHSA 0 0 131 (23). WSNM interdunal vegetation 32 ° 45.57 ’ N 4,006 ’ 106 ° 11.59 ’W gypsum 21 August 2007 Eric H. Metzler WHSA 2 Accss#: WHSA- 0 0 131 (13). WSNM edge of dunes/basin 32 ° 45.70 ’ N 4,001 ’ 106 ° 11.24 ’W 21 August 2007 Eric H. Metzler WHSA 3 Accss#: WHSA-00131 (13). WSNM crest of dunes 106 ° 10.74 ’W 32 ° 46.66 ’ N 4,008 ’ 14 July 2010 WSNMG Eric H. Metzler uv trp Accss # WHSA 0 0 131 (33). WSNM basin grassland veg 32 ° 45.67 ’ N 4,000 ’ 106 ° 11.29 ’W gypsum 13 May 2007 Eric H. Metzler WHSA 4 Accss#: WHSA-00131 (13, 3Ƥ). TEXAS: Padre Island Nat. Seashore 17.V. 76 A&ME Blanchard. (23, 1Ƥ). Briscoe Co.: Caprock Canyons State Park 10,11- V- 96 / ECK (23, 1Ƥ). Caprock Cn.St.Pk. 10,11- V- 96 /ECK (23). Cameron Co.: San Benito Texas Aug (13, 1Ƥ). San Benito Texas (13). San Benito Texas Sept. 8-15 (33, 1Ƥ). San Benito Texas April 24-30 (13, 3Ƥ). San Benito Texas Apr 1-7 (13, 3Ƥ). San Benito Texas March 16-23 (13, 1Ƥ). San Benito Texas Mch 16-23 (1 Ƥ). San Benito Texas June 24-30 (1 Ƥ). Brownsville 22 -XI- 66 A.&M.E. Blanchard. (13, 1Ƥ). Brownsville 18 -XI- 71 A.&M.E. Blanchard. (1 Ƥ). Laguna Atascosa 9.III. 75 A.&M.E. Blanchard (1 Ƥ). Culberson Co.: Guadalupe Mts.N.P. Pine Spring 1,2- X- 91 leg.E.C.Knudson. (1 Ƥ). Guadalupe Mts.N.P. Ship on Desert 6-8 -IX- 91 leg. E.C.Knudson (1 Ƥ). Guadalupe Mts. Nat’l Pk. McKittrick Canyon 21 -IV- 90:ECK Coll. (1 Ƥ). Guadalupe Mts.N.P. Williams Ranch Rd., 2 -X- 91, leg.E.C.Knudson (1 Ƥ). Duval Co.: JctFM 1329 / 2295, 27.V 1985, J.E. Gillaspy, Collector (13). Kenedy Co.: 11 mi. E Encino, 20.IV. 1985, J.E. Gillaspy, Collector (13). Nueces Co.: 9.IX. 74 A&ME Blanchard (13). N. Padre Island 30.IX. 75 A&ME Blanchard (13). N. Padre Island 7.VI. 78 A&ME Blanchard (1 Ƥ). San Patricio Co.: R&B Welder Refuge 14–19.VII. 1985 J.E. Gillaspy Collector (13, 1Ƥ). Sinton Welder Wildlife Foundation 15.XI. 66 A.&M.E. Blanchard (13). (DJW, EHM, MSU, TLSRC, UCB, UNM, USNM). Diagnosis. Eugnosta brownana is a buff-yellow moth with transverse reddish-brown fascia. The diagnostic features are the curved reddish-brown postmedian fascia that touches the inner margin but does not reach the costa, and the gray hind wings. Superficially, E. brownana might be confused with E. bimaculana (Robinson, 1869) and E. sartana (Hübner, 1823 a). In the male genitalia, the socii of E. brownana are directed dorsally, whereas the socii of E. bimaculana and E. sartana are directed ventrally. The three species are best separated with the following key: 1. Adult moth with postmedian fascia not extending to inner margin (Figs 3, 4)............................. bimaculana - Adult moth with postmedian fascia extending to inner margin.................................................. 2 2. Adult moth with curved reddish-brown postmedian fascia ending well short of costa, gray hind wings, broadly rounded cucullus, dorsally directed socii (Figs 1, 2)......................................................... brownana - Adult moth with dark brown postmedian fascia reaching nearly to or touching costa, dark brown hind wings, nearly pointed cucullus, ventrally directed socii (Figs 5, 6)........................................................... sartana Description. Adult male (Fig. 1). Head: Front smooth, buff-yellow, closely scaled; vertex scales narrow straplike, buff-yellow; labial palpus buff-yellow, porrect, closely scaled dorsally, laterally, and mesally, longer scales form fringe ventrally, not shaggy, segment 3 closely scaled. Haustellum coiled between labial palpi. Antenna filiform, dorsally buff-yellow, closely scaled, ventrally setose, naked. Thorax: Dorsum buff-yellow, closely scaled; underside buff-yellow, closely scaled. Foreleg, closely scaled, ventrally buff-yellow, dorsally brown, tarsomere apices dirty yellow. Midleg closely scaled, ventrally buff-yellow, dorsally reddish-brown, tarsomere apices dirty yellow. Hindleg closely scaled, buff-yellow, tarsomere apices dirty yellow. Forewing length 4.2–7.1 mm, mean 5.8 mm, n = 79. Forewing buff-yellow, costa at base pale reddish-brown, postmedian fascia reddish-brown, not reaching costa, touching inner margin; subterminal fascia, reddish-brown, curved basally at mid-wing: fringe gray; underside blotchy mixture of buff-yellow and reddish-brown scales, some specimens with yellow scales, postmedian and subterminal fascia obscure gray silhouettes; fringe gray. Hind wing pale gray to buff-yellow, overlaid with gray scales; fringe base gray, terminally buff-yellow; underside gray and buff-yellow, strigulae reddish-brown, short, chevron-shaped, along costa, veins, and sometimes inner margin; fringe base gray, terminally pale gray. Abdomen: Dorsum closely scaled, buff-yellow; underside closely scaled, buff-yellow. Genitalia (Figs 7, 10) with tegumen widened, ventral margin excurved; uncus absent; socii long, erect, lightly setose, bent 90 ° apically; transtilla extended mesially, V shaped, erect, thin, long, two larger and several smaller stout spines on apex; saccus short, broadly U shaped; valva setose, curved upward, costa narrowly thickened, base of costa with prominent broad tooth-like dorsal projection, cucullus broadly rounded, saccular region slightly widened; juxta shielđ­shapeđ, narrοweđ at miđđle﹔ phallus sclerοtizeđ, apex curveđ 45 with blunt pοint﹔ cοrnutus stοut, slightly shorter than phallus, bifurcate basally, longitudinal suture. Adult female (Fig. 2). Head and Thorax: Similar to male. Forewing length 4.6–7.1 mm, mean 6.1 mm, n = 26. Frenulum bristle single or double. Abdomen: Genitalia (Fig. 13) with papilla analis not sclerotized, setose, long and thin; posterior apophysis extend anteriorly to posterior margin of eighth segment; anterior apophysis length similar to posterior apophysis; sterigma complex involving 9 th sternite, robust; ductus bursae, a sclerotized ring encircles 3 / 4 of ductus at posterior end, elsewhere membranous, elongate; corpus bursae elongate, dense spiculate region at posterior end, signum absent. Systematics. This new species is placed in the genus Eugnosta based on the appearance of the adult moths and the structure of the male genitalia. The sclerotized socii are attached laterally to the apex of the tegumen; the transtilla is strong, the mesal process is upturned and stout; the valva is simple; and the phallus contains a single stout cornutus. Carolella Busck, 1939 has been considered a synonym of Eugnosta or a separate genus by different authors (e.g., Pogue 1986, Brown 2005, Razowski 1998, 2011, Razowski & Becker 2007). Pogue (1986) noted the close relationship between the genera. Brown (2005) followed Pogue and listed them as separate genera. Razowski (1998, 2011) and Razowski & Becker (2007) synonymized Carolella with Eugnosta. Eugnosta brownana appears to share characters of both genera, thus we follow Razowski (2011). Distribution and biology. Eugnosta brownana occurs in Texas, New Mexico, and Arizona. It is common at White Sands National Monument and Carlsbad Caverns National Park. Its distribution in Mexico is unknown. The eggs of E. sartana were documented by Peterson (1961, 1967), and a Russian species, Eugnosta hydrargyana (Eversmann, 1842), was recorded from Jurinea cyanoides (L.) Reichenbach (Asteraceae) (Robinson et al. 2012). Etymology. The specific name of this species, brownana recognizes John W. Brown’s contributions to the study of Tortricidae. John Brown shares personal friendships with Greg Forbes and Eric Metzler that span many years. Remarks. The number of bristles of the frenulum is inconsistent. Some females have one bristle while other females have two bristles. One female has one bristle on one side and two bristles on the other side. The length of the bristles is typical. One male has one bristle on one side and two bristles on the other side. The second bristle is half the length of a typical bristle. For females, the separation of the distal ends of the papilla anales can be seen in situ in many specimens.Published as part of Metzler, Eric H. & Forbes, Gregory S., 2012, The Lepidoptera of White Sands National Monument 5: Two new species of Cochylini (Lepidoptera, Tortricidae, Tortricinae), pp. 51-60 in Zootaxa 3444 on pages 52-55, DOI: 10.5281/zenodo.28215

    Protogygia whitesandsensis Metzler & G. S. Forbes 2009, sp. n.

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    <i>Protogygia whitesandsensis</i> Metzler & Forbes, sp. n. <p>urn:lsid:zoobank.org:act: 21257498-EAF1-4AF5-9357-A3CF6A6FE47E</p> <p>Figs. 13, 17, 19, 20</p> <p> <b>Type material.</b> <b>Holotype</b>: Male: USA: NM: Otero Co. White Sands Nat [ional] Mon [ument] interdunal vegetation 32 o 46.62' N 106 o 10.82' W 4,008' 26 February 2009 Eric H. Metzler WHSAD uv tr[a]p Accession #: WHSA – 00131. (USNM). <b>Paratypes</b>: 18 males as follows: USA, NM: Otero County, White Sands National Monument (hereafter WSNM), interdunal vegetation 106 o 11.49' W 4,000' 32 o 45.60' N 15 Mar 2008 WSNMB Eric H. Metzler uv trp Accession # WHSA 00131. WSNM, interdunal vegetation, 32 o 45.57' N 4,000' 106 o 11.59' W gypsum 12 Mar 2007 WHSA 2 Eric H. Metzler Accession # WHSA 00131. WSNM, interdunal vegetation 32 o 46.62' N 106 o 10.82' W 4,008' 26 Feb 2009 Eric H. Metzler WHSAD uv trp Accession #: WHSA – 00131. WSNM, interdunal vegetation 106 o 11.59' W 32 o 45.57' N 4006' 15 Mar 2008 uv trp WSNM 2 Eric H. Metzler Accession # WHSA – 00131. WSNM, interdunal vegetation 106 o 11.33' W 32 o 45.5' N 4004' uv trp 6 Apr 2008 Eric H. Metzler WSNMA Accession # WHSA – 00131. WSNM, edge of dunes vegetation 32 o 45.72' N 106 o 11.32' W 4,000' WSNM 9 26 Feb 2009 uv trp Eric H. Metzler Accession # WHSA – 00131. WSNM, interdunal vegetation 32 o 45.60' N 106 o 11.49' W 4,000' 26 Feb 2009 Eric H. Metzler uv trp WSNMB Accession # WHSA – 00131. WSNM, 106 o 11.59' W 32 o 45.57' N 4006' 12 Mar 2008 WSNM 2 Eric H. Metzler Accesn # WHSA – 00131. Paratypes are deposited in NMSU, MSWB, MSU, EHM, CNC, USNM, and WHSA.</p> <p> <b>Etymology.</b> The specific epithet of this species, <i>whitesandsensis</i> (-ensis is a Latin suffix denoting “place, locality”) identifies the white gypsum dunes at White Sands National Monument as the type locality. The name recognizes the importance of White Sands National Monument for its unique ecosystem and accompanying biota.</p> <p> <b>Diagnosis.</b> <i>Protogygia whitesandsensis</i> is a pale silvery-gray moth without most normal transverse noctuid markings and spots. A combination of 6 characteristics distinguish <i>Protogygia whitesandsensis</i> from other species of <i>Protogygia</i>: 1) pectinations of male antenna are 9.6 × width of antenna whereas pectinations of male antenna of <i>P. pectinata</i> Lafontaine, 2004 (Fig. 14) are 7 × width of antenna; those of <i>P. biclavis</i></p> <p>(Grote, 1879) (Fig. 16) are 2 × width of antenna; those of <i>P. comstocki</i> McDunnough, 1934 (Fig. 15) are 1.5 × width of antenna; 2) fore wing color pale silvery gray; 3) fore wing with dark and white streaks from the base to the medial area, postmedial area, and outer margin; basal dashes of <i>P. pectinata</i>, <i>P. comstocki</i> and <i>P. biclavis</i> extend to antemedial area; 4) forewing with a prominent white shade in the cell between the base and the reniform spot; 5) adults fly in February and March in the gypsum dunes in the Tularosa Basin of south central New Mexico; 6) adults most closely resemble <i>P. pectinata.</i> The colors of <i>P. whitesandsensis</i> are more gray and muted than <i>P. pectinata</i>. The colors of <i>P. pectinata</i> are more brown and contrasting than <i>P. whitesandsensis</i>.</p> <p>In Lafontaine and Fauske (2004) this species easily keys out to <i>P. pectinata</i>, but <i>P. whitesandsensis</i> has wider antennae, less orange on the forewing, less contrasting veins, and a different distribution.</p> <p> <b>Description. Adult male</b> (Fig. 13): <b>Head</b> – vertex, narrow and narrow hair-like scales, white, occasional black scales, erect; front, narrow hair-like scales, white, a narrow band of scales across the front between the eyes, black, hair-like; labial palpus white with black scales, blackened laterally; basal and medial segments erect, hair-like scales, shaggy; apical segment angled anteriorly, closely scaled dorsally and anteriorly, white; haustellum coiled between labial palpi with more than four complete loops; antenna broadly bipectinate, dorsally white, scattered black scales, pectinations and ventral surface naked, dark brown. <b>Thorax</b> – color white, scattered black scales; tegula white, scattered black scales, laterally and mesally lined with black; dorsum white, scattered black scales, mid-dorsal black stripe; underside white, mixed with black scales, hair-like, long erect, shaggy appearance. Legs: coxa and femur white mixed with black giving a dirty appearance, closely scaled with long hair-like scales on ventral surface forming a shaggy fringe, fore tibia and mid-tibia black terminus, tarsomeres similarly white with black scales, each tarsomere terminus ringed with black. Fore wing: length 14-17 mm, mean 15.2 mm, n = 14. pale silvery gray; basal line, antemedial line, medial shade, postmedial line, claviform spot, and orbicular spot absent; prominent sub costal white shade in cell from base to reniform spot; basal dash, 3 elements, black, prominent, from base to reniform spot, white, not prominent, from base to medial area, black, not prominent, from base to antemedial area; subterminal line black, contrasting, zig zag from costa to inner margin; radial, medial, cubital, and anal veins lined with white; reniform lower and upper lobes lined with white; terminal line vaguely white; fringe, base marked with black scales, otherwise white; underside pale silvery gray, dusky shade in cell, veins variously lined with white, fringe concolorous. Hind wing: pale gray; veins variously lined with white or darker gray scales; discal spot dark gray; fringe, base concolorous, outer half white; underside concolorous; discal spot absent; fringe concolorous. <b>Abdomen</b> – dorsum light gray mixed with black scales, closely scaled, overlaid with long hair-like scales; underside of segments 3, 4, and 5 black and white contrasting, gray elsewhere with admixture of black scales. <b>Genitalia</b> (Fig. 17) – tegumen widened laterally, narrowed dorsally; uncus widened mesally, abruptly down-turned and pointed apically, long hairs dorsally; vinculum V shaped; valve strap-like slightly wider in saccular area, apex with dense setae, directed basad on mesial surface; clavus length 3 × width; clasper sinuous, tear drop shaped, ending in drawn-out point apically; aedeagus slightly bent at middle; vesica bent to right, base of vesica with 6 or 7 stout setae; subbasal diverticulum short with one stout terminal seta.</p> <p> <b>Adult female</b> : unknown.</p> <p> <b>Remarks.</b> This new species is placed in the genus <i>Protogygia</i> based on the male genitalia, and the adult male’s close resemblance to <i>P. pectinata</i>.</p> <p> <b>Distribution and biology.</b> <i>Protogygia whitesandsensis</i> occurs in White Sands National Monument, Otero County, New Mexico (Figs. 19, 20). Adults were collected in black light traps placed within the gypsum dunes. Females and the immature stages are unknown.</p>Published as part of <i>Metzler, Eric, Bustos, David & Forbes, Gregory, 2009, The Lepidoptera of White Sands National Monument, Otero County, New Mexico, USA 1. Two new species of Noctuidae (Lepidoptera, Noctuinae, Agrotini), pp. 47-62 in ZooKeys 9 (9)</i> on pages 54-57, DOI: 10.3897/zookeys.9.182, <a href="http://zenodo.org/record/576451">http://zenodo.org/record/576451</a&gt

    Chionodes bustosorum Metzler, sp. nov.

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    <i>Chionodes bustosorum</i> Metzler, sp. nov. <p>BOLD:ACS7412</p> <p>Figs 1–3</p> <p> <b>Holotype</b> ♂, pinned, double-mounted, with labels as follows: “New Mexico: Otero Co.,White Sands Nat[ional] Mon[ument]; Interdune vegetation; 106˚11.38'W; 32˚46.60' N 4,000 '; 11 June 2010 WSNM 8; Eric H. Metzler uv tr[a]p; Accss # White Sands National Monument 00131.” [blue label] “Barcode of Life Project Leg(s), DNA extracted”; “USNMENT 01142737 ”; [green label] “Genitalia slide by EHMetzler, ♂ USNM 146317” [red label] “ HOLOTYPE USNM; <i>Chionodes bustosorum</i> Metzler 2016 ”. Deposited in USNM.</p> <p> <b>Paratype</b> ♂, pinned, double-mounted, same locality/date as holotype, [blue label] “Barcode of Life Project Leg(s) removed, DNA extracted”; “USNMENT 01142738 ” [green label] Genitalia slide by EHMetzler, ♂ USNM 146318.” Deposited in USNM.</p> <p> <b>Description</b>. Adult male (Fig. 1). Head: Front and vertex scales broadly spatulate, erect, cream-white; front smooth, scales spatulate, directed forward and ventrally, cream-white; palpi upturned, basal segment scales appressed, mid-segment = ½ length of palpi, slightly shaggy, scales spatulate, apical segment = 1/3 length of palpi, divergent apically, scales appressed, cream-white; haustellum base densely scaled, cream-white. Antenna, each segment basally ringed with semi-erect cream-white scales. Thorax: dorsal and ventral surfaces with appressed, cream-white scales. Legs with appressed, cream-white scales. Forewing: Length 5.4 mm, mean 5.4 mm, n = 2; Uniformly cream-white, including fringe, apex rounded; underside pale yellow with concolorous fringe. Hindwing mirror-like reflective-white with concolorous, long fringe, apex slightly produced; underside white, male with patch of pale yellow sex scales arising from wing base and extending along inner margin. Abdomen: Scales appressed, pale yellow. T8 (Fig. 2d) with lateral sides parallel, posteriorly broadly convex, anteriorly broadly concave. Male genitalia (Fig. 2) (2 preparations examined) with uncus broad, spoon shaped, setose laterally; culcitula absent; gnathos with base sclerotized, lobed laterally, lobes extending 1/3 length of tegumen, each lobe with a posteriorly directed finger-like projection enclosing a diamond-shaped lateral process; gnathos sharply curved at 0.2x length, gently curved most of length, apex sharply recurved; tegumen broadly A-shaped, excavated to 1/3, robust suture separating pedunculi, each pedunculus narrowing to junction with vinculum, base of each pedunculus twisted 180˚ at junction with vinculum; vinculum = 0.8x length of tegumen, abruptly narrowed immediately anterior of juncture with tegumen, distal 2/3 trough-shaped, sides parallel extended to blunt rounded apex; posterolateral lobe from vinculum an extension of twisted pedunculus, sclerotized, trough-shaped, apex bluntly rounded, length = width of base of pedunculus of tegumen; saccus not differentiated from vinculum; valvae asymmetrical, unequal in length and dissimilarly shaped: right valva, maximally extended to 1/2 length of tegumen, then strongly recurved mesially at 2/3 length, robust where recurved, apex weakly bifurcate, posterior projection thorn-like, robust anterior projection pointed, beak-like; left valva narrow, curved, bow-like mesially or laterally, extended to middle of uncus, terminal 1/10 bent approximately 90˚, apex doubly bifurcate, posterior projection robust pointed, beak-like, anterior projection bifurcate, not robust. Phallus with distal part sculpting complex, caecum approximately 5x length of distal part, longitudinal sclerotized bar at anterior end.</p> <p>Adult female. Unknown.</p> <p> <b>Diagnosis</b>. Ron Hodges (personal communication to EHM) considered <i>C. bustosorum</i> to be undescribed based on the diagnosis in combination with the details of the description. <i>Chionodes bustosorum</i> (Fig. 1) is a small (forewing length 5.4 mm) creamy white moth with no discernable markings on the surface. Two nearly identical specimens were captured in black light traps with Scythrididae of similar size and appearance. The apex of the forewing of <i>C. bustosorum</i> is rounded, whereas the forewing of Scythrididae terminate in an acute apex. The apex of the hindwing of <i>C. bustosorum</i> is rounded and produced, a wing shape that is typical for most Gelechiidae. The apex of the hindwing of Scythrididae is acute. The male genitalia of <i>C. bustosorum</i> are typical for the genus <i>Chionodes</i> (Fig. 2) with a broad prominent spoon-shaped uncus. <i>Chionodes bustosorum</i> keys out to couplet 6, <i>obscurusella</i> or <i>formosella</i> groups, in the Hodges (1999: 25) key. The male sex scales of <i>C. bustosorum</i> are on the ventral surface of the hindwing, similar to species in the <i>obscurusella</i> group. Based on characters of the male genitalia as figured by Hodges, <i>C. bustosorum</i> appears to be in the <i>abella</i> complex of the <i>formosella</i> group, however, <i>C. bustosorum</i> fails to match any features in the early couplets in the key to species of the <i>formosella</i> group (see Hodges 1999: 33). The diagnostic features of the male genitalia of <i>C. bustosorum</i> are the markedly asymmetrical valvae. The left valva is long and narrow, whereas the right valva is shorter, more robust, and recurved mesially like the cutting blade of a hand-held scythe. In a comparison of male genitalia, based on the photographs in Hodges (1999), the genital capsule of <i>C. bustosorum</i> is most similar to a lateral mirror image of <i>C. abella</i> as illustrated on p. 261, fig. D-1 of that work. The angle of the recurved right valva of <i>C. bustosorum</i> is broad, like a hand-held scythe, whereas the angle of the recurved left valva of <i>C. abella</i> is acute. The area between the pedunculi of the tegumen of <i>C. bustosorum</i> is not sclerotized, whereas in <i>C. abella</i> it is lightly sclerotized. The lateral sides of the uncus of <i>C. abella</i> appear to be nearly parallel; the uncus of <i>C. bustororum</i> is oval. The genital capsule also resembles a mirror image of that of <i>C. abdominella</i> as illustrated on p. 263, fig. F-20 of Hodges (1999).</p> <p> <b>Remarks</b>. This new species is placed in the genus <i>Chionodes</i> based on the presence of the caecum on the aedeagus as defined by Hodges (1999). The holotype and paratype are identical in habitus and genital structure. The paratype imago is illustrated because it possesses complete antennae. The holotype was selected because it is the specimen that yielded a nearly complete barcode sequence.</p> <p> <b>Etymology.</b> The specific name of this species, <i>bustosorum</i>, a noun in the genitive case, recognizes David Bustos, Chief of Resources at White Sands National Monument since 2007, and his wife, Andrea. David aggressively pursues research; his efforts contributed greatly to the immense accumulation of scientific data during his years at White Sands National Monument. Metzler knows from personal experience that David works long and crazy hours in support of the Monument. He is enthusiastic about the research on moths, to wit, he often sends emails to Metzler from home, long after Metzler is in bed. David is the recipient of the U.S. National Park Service’s 2014 Director's Trish Patterson Student Conservation Association Award for Natural Resource Management in a Small Park. His efforts would not be possible without the support and encouragement of his lovely wife Andrea. Metzler takes great pleasure in naming this moth in honor of David and Andrea Bustos.</p> <p> <b>Distribution and biology.</b> <i>Chionodes bustosorum</i> occurs in White Sands National Monument, Otero County, New Mexico (Fig. 3). The immature stages and host plant are unknown.</p>Published as part of <i>Metzler, Eric H. & Landry, Jean-François, 2016, The Lepidoptera of White Sands National Monument, Otero County, New Mexico, USA 10. A remarkable new white species of Chionodes Hübner (Gelechiidae), pp. 372-380 in Zootaxa 4109 (3)</i> on pages 373-377, DOI: 10.11646/zootaxa.4109.3.7, <a href="http://zenodo.org/record/262007">http://zenodo.org/record/262007</a&gt

    The Lepidoptera of White Sands National Monument, Otero County, New Mexico, USA 10. A remarkable new white species of Chionodes Hübner (Gelechiidae)

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    Metzler, Eric H., Landry, Jean-François (2016): The Lepidoptera of White Sands National Monument, Otero County, New Mexico, USA 10. A remarkable new white species of Chionodes Hübner (Gelechiidae). Zootaxa 4109 (3): 372-380, DOI: 10.11646/zootaxa.4109.3.

    FIGURE 1 in The Lepidoptera of White Sands National Monument, Otero County, New Mexico, USA 10. A remarkable new white species of Chionodes Hübner (Gelechiidae)

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    FIGURE 1. Chionodes bustosorum adult male paratype (scale bar = 1 mm).Published as part of Metzler, Eric H. & Landry, Jean-François, 2016, The Lepidoptera of White Sands National Monument, Otero County, New Mexico, USA 10. A remarkable new white species of Chionodes Hübner (Gelechiidae), pp. 372-380 in Zootaxa 4109 (3) on page 374, DOI: 10.11646/zootaxa.4109.3.7, http://zenodo.org/record/26200

    FIGURE 22–23. Distribution maps. 22 in The Lepidoptera of White Sands National Monument 5: Two new species of Cochylini (Lepidoptera, Tortricidae, Tortricinae)

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    FIGURE 22–23. Distribution maps. 22. Eugnosta brownana distribution map; 23. Cochylis yinyangana distribution map.Published as part of Metzler, Eric H. & Forbes, Gregory S., 2012, The Lepidoptera of White Sands National Monument 5: Two new species of Cochylini (Lepidoptera, Tortricidae, Tortricinae), pp. 51-60 in Zootaxa 3444 on page 58, DOI: 10.5281/zenodo.28215

    FIGURES 13–15. Eugnosta female genitalia. 13. E in The Lepidoptera of White Sands National Monument 5: Two new species of Cochylini (Lepidoptera, Tortricidae, Tortricinae)

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    FIGURES 13–15. Eugnosta female genitalia. 13. E. brownana female genitalia paratype; 14. E. bimaculana female genitalia; 15. E. sartana female genitalia.Published as part of Metzler, Eric H. & Forbes, Gregory S., 2012, The Lepidoptera of White Sands National Monument 5: Two new species of Cochylini (Lepidoptera, Tortricidae, Tortricinae), pp. 51-60 in Zootaxa 3444 on page 55, DOI: 10.5281/zenodo.28215

    FIGURES 7–12. Eugnosta male genitalia. 7. E in The Lepidoptera of White Sands National Monument 5: Two new species of Cochylini (Lepidoptera, Tortricidae, Tortricinae)

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    FIGURES 7–12. Eugnosta male genitalia. 7. E. brownana male genitalia paratype; 8. E. bimaculana male genitalia; 9. E. sartana male genitalia; 10. E. brownana male phallus paratype; 11. E. bimaculana male phallus; 12. E. sartana male phallus.Published as part of Metzler, Eric H. & Forbes, Gregory S., 2012, The Lepidoptera of White Sands National Monument 5: Two new species of Cochylini (Lepidoptera, Tortricidae, Tortricinae), pp. 51-60 in Zootaxa 3444 on page 54, DOI: 10.5281/zenodo.28215

    Figures 17-18 in The Lepidoptera of White Sands National Monument, Otero County, New Mexico, USA 1. Two new species of Noctuidae (Lepidoptera, Noctuinae, Agrotini)

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    Figures 17-18. Male genitalia of Protogygia species: a-valves, b-aedeagus with vesica everted. 17. P. whitesandsensis, paratype. 18. P. pectinata.Published as part of Metzler, Eric, Bustos, David & Forbes, Gregory, 2009, The Lepidoptera of White Sands National Monument, Otero County, New Mexico, USA 1. Two new species of Noctuidae (Lepidoptera, Noctuinae, Agrotini), pp. 47-62 in ZooKeys 9 (9) on page 56, DOI: 10.3897/zookeys.9.182, http://zenodo.org/record/57645

    ROSENTHAL, Eric Inventory of documents

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    COVERAGE 1904; 1 File; 011 metre.Private papers of Eric Rosenthal, author, journalist and broadcaster
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