123 research outputs found

    Megaselia chicheckliensis Khameneh & Khaghaninia & Disney & Maleki-Ravasan 2019, n. sp.

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    Megaselia chicheckliensis n. sp. (Figs 12–24) Material examined. Holotype male, East Azerbaijan province, Chichekli region, Arasbaran forests, open forest, 38°39.899’ N, 46°31.248’ E, 2140 m, 14.vi.2016, S. Khaghaninia (34, CUMZ—13-84). Paratypes: 3 males, locality data as the holotype (ICHMM). Description. Male. Fig. 12, whole fly. Frons as Fig. 13, with suopra-antennal bristles equally strong and dense very fine microtrichia. Cheek with 4 bristles and jowl with 2 as long and as robust. Postpecicels without SPS vesicles, but with pale spots (Fig. 14). Palps and proboscis as Fig. 15. Thorax brown, with 3 notopleural bristles and no cleft in front of these and mesopleuron with hairs and 1 bristle (Fig. 16). Scutellum with an anterior pair of small hairs and a posterior pair of bristles (Fig. 17). Abdomen as Fig. 18, the venter having hairs restricted to segments 4 and 5. Hypopygium as Figs 19–21, the penis complex including a pale process tipped with a pair of short but thick spines (Fig. 20). Hind legs brown but middle and hind legs increasingly yellowish brown. Fore tarsus with posterodorsal hair palisade on segments 1–4 (Fig. 22). Dorsal hair palisade of mid tibia extends about 0.6 times its length. Hind femur as Fig. 23. Hind tibia with about a dozen only moderately differentiated posterodorsal hairs, without anterodorsals, and spinules of apical combs simple. Wings (Fig. 24) 1.29 mm long. Costal index 0.35. Costal ratios 3.92: 1.22: 1. Costal cilia (of section 3) 0.06 mm long. No hair at base of vein 3. With 2 unequal axillary bristles the outer being 0.10 mm long. Sc not reaching R1. Haltere brown. Recognition. In the key the British species (Disney 1989) it runs to couplet 58, where neither option fits. Of five excluded Palaearctic species running to this couplet only M. tama (Schmitz) is close, but its hypopygium is clearly different. In the key of Schmitz (1958) for Abteilung III it runs to couplet 26 lead 2, to M. filamentosa Schmitz, but its palps are brown not yellow and the wings not so pale. Etymology. Named after the region of the holotype, Chicheckli region.Published as part of Khameneh, Roya Namaki, Khaghaninia, Samad, Disney, R. Henry L. & Maleki-Ravasan, Naseh, 2019, Twenty one new species of Megaselia Rondani (Diptera: Phoridae) from Iran, pp. 1-50 in Zootaxa 4711 (1) on page 4, DOI: 10.11646/zootaxa.4711.1.1, http://zenodo.org/record/357336

    Twenty one new species of Megaselia Rondani (Diptera: Phoridae) from Iran

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    Khameneh, Roya Namaki, Khaghaninia, Samad, Disney, R. Henry L., Maleki-Ravasan, Naseh (2019): Twenty one new species of Megaselia Rondani (Diptera: Phoridae) from Iran. Zootaxa 4711 (1): 1-50, DOI: https://doi.org/10.11646/zootaxa.4711.1.

    Megaselia mahabadensis Khameneh & Khaghaninia & Disney & Maleki-Ravasan 2019, n. sp.

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    Megaselia mahabadensis n. sp. (Figs 205–218) Material examined. Holotype male, West Azerbaijan province, Mahabad city, Ghalate-shah region, 36°46.01’N, 45°22.37’E, 1605 m, grassland, 19.VI.2017, S. Khaghaninia (54, CUMZ—13-97). Description. Male. Whole fly as Fig. 205. Frons as Fig. 206, with very fine microtrichia (Fig. 207). Cheek with 5 bristles and jowl with 2 that are longer and more robust. Postpedicels, which lack SPS vesicles, palps and proboscis as Fig. 208, the palps having many small pits (Fig. 209).Thorax brown, with 3 notopleural bristles and no cleft in front of these. Mesopleuron with many hairs and a single short bristle (Fig. 210). Scutellum with an anterior pair of small hairs and a posterior pair of bristles. Abdominal tergites brown with relatively few small hairs except for longer more robust hairs laterally on T6 (Fig. 211). Venter dark gray, and with hairs on segments 3–6, but those on 3–5 a small and few in number. Hypopygium as Figs 211–213), the hypandrial lobes lacking. Legs brown. Fore tarsus with posterodorsal hair palisades on segments 1–4 with 5 clearly longer than 4 (Fig. 214) and basitarsurs vetrally with rows of small spinules (Fig. 215). Dorsal hair palisade of mid tibia extends about 0.54 times its length. Hairs below basal half of hind femur subequal to or slightly longer than those of anteroventral row of outer half (Fig. 216). Hind tibia with 9–10 moderately differentiated posterodorsal hairs, without anterodorsals, and spinules of apical combs simple. Wings (Figs 217 & 218) 1.5–1.6 mm long mm long. Costal index 0.42. Costal ratios 3.41: 1.59: 1. Costal cilia (of section 3) 0.09 mm long. Vein 3 hair lacking. 3 axillary bristles, the outermost being 0.09 mm long. Sc not reaching R1 and vein 7 obscure. Haltere brown. Recognition. In the key to the Megaselia males of the British Isles (Disney 1989) it runs to couplet 28. As the bristle on the mesopleuron is relatively short one could take either lead, but both take one on to couplet 60 and then onto couplet 91, where the hypopygium differs from either option. In Schmitz’s (1958) key to Abteilung III it runs to couplets 17 or 27, but the hypopygium differs from all the options. In the key of Schmitz & Delage (1974) to Abteilung V it runs to couplet 7 lead 1, where again the hypopygium rules this option. Etymology. Named after Mahabad city.Published as part of Khameneh, Roya Namaki, Khaghaninia, Samad, Disney, R. Henry L. & Maleki-Ravasan, Naseh, 2019, Twenty one new species of Megaselia Rondani (Diptera: Phoridae) from Iran, pp. 1-50 in Zootaxa 4711 (1) on pages 34-35, DOI: 10.11646/zootaxa.4711.1.1, http://zenodo.org/record/357336

    Megaselia ajabshirensis Khameneh & Khaghaninia & Disney & Maleki-Ravasan 2019, n. sp.

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    Megaselia ajabshirensis n. sp. (Figs 1–11) Material examined. Holotype male, East Azerbaijan province, Ajabshir region, 37°31.978’ N, 46°07.716’ E, 1662 m, grassland, 16.Vii.2010, S. Khaghaninia (50, UCMZ—13-99). Description. Male. Fig. 1, whole fly. Head as Fig. 2, with the frons lacking microtrichia except at its edges. Cheek with 5 bristles and jowl with 2 that are longer and more robust. Postpedicels, which lack SPS vesicles, palps and proboscis from below as Fig. 3. Thorax brown, with 3 notopleural bristles and no cleft in front of these, and mesopleuron with hairs (Fig. 4). Scutellum with an anterior pair of hairs and a posterior pair of bristles. Abdominal tergites and venter as Fig. 5. Hypopygium as Figs 6–8, with left hypandrial lobe having microtrichia projecting from its tip (Fig. 7). The right lobe is vestigial. Fore tarsus with posterodorsal hair palisade on segments 1–4 (Fig. 9) and its basitarsus with rows of spinules ventrally (Fig. 10). Dorsal hair palisade of mid tibia extends about 0.63 times its length. Hind femur as Fig. 11. Hind tibia with 15–16 moderately differentiated posterodorsal hairs, without anterodorsals, and spinules of apical combs simple. Wings 1.36 mm long, light gray with thick veins light brown and thin veins pale gray. Costal index 0.39. Costal ratios 4.1: 1.1: 1. Costal cilia (of section 3) 0.08 mm long. No hair at base of vein 3. With 2 axillary bristles, the outer one being 0.09 mm long. Sc not reaching R1. Haltere knob brown. Recognition. In the key the British species (Disney 1989) it runs to couplet 87. The hypopygium of M. conformis (Wood) has short hairs on the epandrium. The palps of M. nigripalpis (Lundbeck) are brown and its cosrtal index exceeds 0.45. A further 8 Palaearctic species will run to the same couplet. Most are excluded by having more than 2 axillary bristles, brown palps and/or costal index exceeding 0.45. Of the rest the dorsal edge of the epandrium is sub-equal in length to the ventral edge in contrast to our new species. Non Palaearctic species are likewise excluded. Etymology. Named after the region of the holotype.Published as part of Khameneh, Roya Namaki, Khaghaninia, Samad, Disney, R. Henry L. & Maleki-Ravasan, Naseh, 2019, Twenty one new species of Megaselia Rondani (Diptera: Phoridae) from Iran, pp. 1-50 in Zootaxa 4711 (1) on pages 3-4, DOI: 10.11646/zootaxa.4711.1.1, http://zenodo.org/record/357336

    Megaselia ghalateshahensis Khameneh & Khaghaninia & Disney & Maleki-Ravasan 2019, n. sp.

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    Megaselia ghalateshahensis n. sp. (Figs 123–135) Material examined. Holotype male, West Azerbaijan province, Mahabad city, Ghalate-shah region, 36°46.01’N, 45°22.37’E, 1605 m, grassland, 19.VI.2017, S. Khaghaninia (56, CUMZ—13-97). Description. Male. Whole fly as Fig. 123. Frons as Fig. 124, largely devoid of microtrichia (Fig. 125). Cheek with 3 bristles and jowl with a single longer and more robust bristle. Postpedicels, which lack SPS vesicles, palps and proboscis as Fig. 126, the labella without short spinules below. Thorax brown. Mesopleuron with hairs and a single long bristle, and with 3 notopleural bristles and no cleft in front of these (Fig. 127). Scutellum with an anterior pair of small hairs and a posterior pair of bristles. Abdominal as Fig. 128. Venter brown, and with hairs on segments 5 and 6). Hypopygium as Figs 129 & 130). Legs brown. Fore tarsus Fig. 131, with some small spinules below the basitarsus (Fig. 132). Dorsal hair palisade of mid tibia extends about 0.60 times its length. Hairs below basal half of hind femur longer than those of anteroventral row (Fig. 133). Hind tibia with a dozen moderately differentiated posterodorsal hairs, without anterodorsals, and spinules of apical combs simple. Wings (Figs 134 & 135) 1.39 mm long. Costal index 0.45. Costal ratios 3.13: 1: 1. Costal cilia (of section 3) 0.07 mm long. Vein 3 without a hair. 2 unequal axillary bristles, the outermost being 0.07 mm long. Sc not reaching R1. Haltere brown (Fig. 134). Recognition. In the key to the Megaselia males of the British Isles (Disney 1989) it runs to couplets 47 and 48, but none of the options fit. A dozen omitted or subsequently described species run to these couplets, differences in the hypopygia, costal indexes and/or the colour of the legs and/or palps exclude all but M. globicornis Schmitz. However, the latter has the hairs below the basal half of the hind femur clearly shorter than those of the anteroventral row (Fig. 139) and the similar hypopygium (Figs 136–138) differs in detail, such its possession of a long, fingerlike, left hypandrial lobe (Fig. 137). In Schnitz’s (1957) key to the Palaearctic Abteilung II it runs to couplet 20 where the hypopygia of both options are clearly different. It fails to run down for the rest of the world. Etymology. Named after the region of the holotype.Published as part of Khameneh, Roya Namaki, Khaghaninia, Samad, Disney, R. Henry L. & Maleki-Ravasan, Naseh, 2019, Twenty one new species of Megaselia Rondani (Diptera: Phoridae) from Iran, pp. 1-50 in Zootaxa 4711 (1) on pages 19-23, DOI: 10.11646/zootaxa.4711.1.1, http://zenodo.org/record/357336

    FIGURES 213–218. Megaselia mahabadensis n in Twenty one new species of Megaselia Rondani (Diptera: Phoridae) from Iran

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    FIGURES 213–218. Megaselia mahabadensis n. sp. male. 213, right face of hypopygium; 214, front tarsus; 215, front basitarsus; 216, hind femur to basitarsus; 217, wing, 218, base of wing. Scale bars 20 μm.Published as part of Khameneh, Roya Namaki, Khaghaninia, Samad, Disney, R. Henry L. & Maleki-Ravasan, Naseh, 2019, Twenty one new species of Megaselia Rondani (Diptera: Phoridae) from Iran, pp. 1-50 in Zootaxa 4711 (1) on page 37, DOI: 10.11646/zootaxa.4711.1.1, http://zenodo.org/record/357336

    FIGURES 148–151. Megaselia haddadi n in Twenty one new species of Megaselia Rondani (Diptera: Phoridae) from Iran

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    FIGURES 148–151. Megaselia haddadi n. sp. male. 148, hypopygium; 149, front tarsus; 150, hind femur; 151, wing. Scale bars 20 μm.Published as part of Khameneh, Roya Namaki, Khaghaninia, Samad, Disney, R. Henry L. & Maleki-Ravasan, Naseh, 2019, Twenty one new species of Megaselia Rondani (Diptera: Phoridae) from Iran, pp. 1-50 in Zootaxa 4711 (1) on page 26, DOI: 10.11646/zootaxa.4711.1.1, http://zenodo.org/record/357336

    Megaselia shabestarensis Khameneh & Khaghaninia & Disney & Maleki-Ravasan 2019, n. sp.

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    Megaselia shabestarensis n. sp. (Figs 70–78) Material examined. Holotype male, East Azerbaijan province, Shabestar city—Shanejan region, 38°13.85’ N, 45°42.93’ E, 1664 m, grassland, 18.vi.2014, R. Namaki Khameneh (20, UCMZ 13-94). Paratypes: 37 males, locality data as the holotype (5 UCZM, 32 ICHMM). Description. Male. Whole fly as Fig. 70. Head as Fig. 71, with the frons lacking microtrichia except at its edges. Cheek with 4 bristles and jowl with 2 that are longer and more robust. Postpedicels Fig. 72, without SPS vesicles but with pale spots. Palps with small pits (Fig. 72). Three notopleural bristles, with no cleft in front of these, and mesopleuron bare Fig. 73. Scutellum with an anterior pair of hairs and a posterior pair of bristles. Abdominal tergites brown and venter gray, the latter with hairs on segments 5 and6 only (Fig. 74). Hypopygium as Figs 74 & 75. Legs brown. Fore tarsus (Fig. 76) with posterodorsal hair palisade on segments 1–4 and 5 clearly longer than 4. Dorsal hair palisade of mid tibia extends about 0.6 times its length. Hairs below basal half of hind femur longer than those of anteroventral row of outer half (Fig. 77). Hind tibia with about 20 weakly differentiated posterodorsal hairs, without anterodorsals, and spinules of apical combs simple. Wings (Fig. 78) 0.9–1.6 mm long. Costal index 0.30–0.35. Costal ratios 7.8–7.9: 1.5–1.7:1. Costal cilia 0.09 mm long. Hair at base of vein 3 0.02 mm long. With 2 unequal axillary bristles, the outermost being 0.09 mm long. Vein Sc not reaching vein 1. Recognition. In the key the British species (Disney 1989) it runs to couplets176 and 179. This complex of species has since been revised (Disney 2014). In the revised key to this group it runs to M. devia Schmitz and M. hirticaudata (Wood). It differs from both in having the hairs of the abdominal venter restricted to segments 5 and 6, its costal index being less than 0.4, and from M. devia having dusky as opposed to yellow palps. In the rest of the world fauna it most closely resembles the Nerarctic M. pygmaeola Borgmeier (1966) but its costal ratios are 4: 1: 1, as opposed to section 2 being clearly longer than 3, and its abdominal tergites are much more sparsely haired. Etymology. Named after the type locality Shabestar City.Published as part of Khameneh, Roya Namaki, Khaghaninia, Samad, Disney, R. Henry L. & Maleki-Ravasan, Naseh, 2019, Twenty one new species of Megaselia Rondani (Diptera: Phoridae) from Iran, pp. 1-50 in Zootaxa 4711 (1) on page 11, DOI: 10.11646/zootaxa.4711.1.1, http://zenodo.org/record/357336

    FIGURES 219–222. Megaselia miandoabensis n in Twenty one new species of Megaselia Rondani (Diptera: Phoridae) from Iran

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    FIGURES 219–222. Megaselia miandoabensis n. sp. male. 219, whole fly; 220, frons; 221, detail of frons; 222, postpedicel. Scale bars 20 μm.Published as part of Khameneh, Roya Namaki, Khaghaninia, Samad, Disney, R. Henry L. & Maleki-Ravasan, Naseh, 2019, Twenty one new species of Megaselia Rondani (Diptera: Phoridae) from Iran, pp. 1-50 in Zootaxa 4711 (1) on page 38, DOI: 10.11646/zootaxa.4711.1.1, http://zenodo.org/record/357336

    Megaselia haddadi Khameneh & Khaghaninia & Disney & Maleki-Ravasan 2019, n. sp.

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    Megaselia haddadi n. sp. (Figs 140–151) Material examined. Holotype male, West Azerbaijan province, Khoy city, Evogli region, 38°42.436’N, 45°12.246’E, 968 m, Malaise trap, 26.ix.2013, S. Khaghaninia (63, CUMZ—13-90). Paratype: 1 male, locality data as the holotype (ICHMM). Description. Male. Whole fly as Fig. 140. Frons as Fig. 141, without microtrichia except at edges (Fig. 142). Cheek with 2 bristles and jowl with 2 that are longer and more robust. Antennae, palps and proboscis as Fig. 143, the postpedicels being without SPS vesicles. Mesopleuron bare and with 2 notopleural bristles and no cleft in front of these. Scutellum with an anterior pair of small hairs and a posterior pair of bristles.Abdominal tergites brown and venter gray with fine hairs on segments 3–6. Hypopygium as Figs 144–148, with at least hypandrial lobe (L in Fig. 147) but no right lobe. Legs yellow. Fore tarsus with posterodorsal hair palisade on segments 1–4 and 5 a little longer than 4 (Fig. 149). Dorsal hair palisade of mid tibia extends about 0.73 times its length. Hairs below basal half of hind femur clearly longer than those of anteroventral row of outer half (Fig. 150). Hind tibia with a dozen well differentiated posterodorsal hairs, without anterodorsals, and spinules of apical combs simple. Wings (Fig. 151) 1.48 mm long. Costal index 0.44. Costal ratios 4.22: 2.04: 1. Costal cilia (of section 3) 0.09 mm long. Vein 3 hair 0.07 mm long. 2 unequal axillary bristles, the outermost being 0.10 mm long. Sc not reaching R1. Haltere knob yellow. Recognition. In the key to the Megaselia males of the British Isles (Disney 1989) it runs to couplet 224 lead 2 M. hayleyensis Disney, which was subsequently synonymised with M. quadriseta (Schmitz), which had been described from the female only (Buck & Disney 2001). However, M. quadriseta has a longer costal index and costal cilia, front tarsi with posterodorsal hair palisades on all five segments and the postpedicels with SPS vesicles. The subsequently described M. citrinella Buck runs to this couplet but it has an orange thorax, and also has a longer costal index and costal cilia. In Borgmeier’s (1966) to Nearctic Group VII it runs to couplet 12, but its costal index is too short. The subsequently described M. creasoni Hartop, Brown & Disney (2015) runs to the same couplet, but it has 3 notopleural bristles and the bristles at the tip of the proctor are clearly longer and more robust than those on cerci. It fails to fit any species from the rest of the world. Etymology. Named after Professor Karim Haddad Irani-nejad. (Retired professor of Department of Plant Protection, Tabriz, Iran).Published as part of Khameneh, Roya Namaki, Khaghaninia, Samad, Disney, R. Henry L. & Maleki-Ravasan, Naseh, 2019, Twenty one new species of Megaselia Rondani (Diptera: Phoridae) from Iran, pp. 1-50 in Zootaxa 4711 (1) on pages 23-24, DOI: 10.11646/zootaxa.4711.1.1, http://zenodo.org/record/357336
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