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Drymeia pyrenaea Michelsen, 2011, sp. nov.
No full textDrymeia pyrenaea sp. nov. (Figs. 1, 3, 5, 7, 8.) Etymology. The specific epithet ‘pyrenaea’ alludes to the type area, the Pyrenees range of mountains. Description. Drymeia pyrenaea sp. nov. agrees in most respects so closely with D. brumalis (Rondani) that the differences between the two species are conveniently presented in table format, see Table 1. Type material examined. All deposited in ZMUC: SPAIN: GERONA: Queralps [Caralps]: 1200–1300m, holotype male, paratypes 1 male, 1 female, 13–16.vi. 1982 (Andersen, Lyneborg & Michelsen); LÉRIDA: Sorpe, 1200–1300m, paratypes 3 males, 2 females, 19–22.vi. 1982 (Andersen, Lyneborg & Michelsen). Additional material examined. All deposited in ZMUC: GERONA: S a Nuria, 2100m, 2 females, 10.vii. 1984 (V. Michelsen); Bruguera SE Ribes, 1700m, 1 male, 1 female, 18.vi. 2002 (V. Michelsen). LÉRIDA: Sorpe, 1200– 1300m, 3 females, 19–22.vi. 1982 (Andersen, Lyneborg & Michelsen); Valle de Arán, Arties, 1200–1800m, 1 male, 20.vi. 1982 (Andersen, Lyneborg & Michelsen), 1800m, 1 female, 6-7.vii. 1984 (V. Michelsen); Gavàs 6km NE Esterri d’Aneu, 3 females, 13.vi. 2002 (V. Michelsen). TERUEL: Cantavieja, 1200m, 1 female, 28.vi. 1984 (V. Michelsen). PORTUGAL: Braga, S a do Gerez, 1 female, 10.vii. 1990 (V. Michelsen). Examined Iberian material of Drymeia brumalis is as follows: SPAIN: HUESCA: 15km NE Benasque, 1 male, 2 females, 7.vii. 1997 (V. Michelsen). Distribution. Drymeia pyrenaea sp. nov. is so far only known from northern, elevated parts of the Iberian Peninsula, but the coexistence of D. brumalis in that area suggests that D. pyrenaea might even occur in the Alps.Published as part of Michelsen, Verner, 2011, A new European species of Drymeia Meigen (Diptera: Muscidae) near D. brumalis (Rondani), with observations of a unique prestomal teeth structure, pp. 51-60 in Zootaxa 3023 on page 53, DOI: 10.5281/zenodo.20774
Botanophila helviana Michelsen 1983
No full text1. Botanophila helviana Michelsen, 1983 Figs. 1, 4, 11–14. Botanophila helviana Michelsen 1983: 293; Ackland 1989: 217; Michelsen & Barták 2001: 457, 460; Michelsen & Báez 2002: 200; Michelsen 2004. Description. Size. Small, wing length 3.5–4.1mm (n= 22). Male. Frons at narrowest point slightly narrower than distance between inner margins of posterior ocelli. Parafacial at narrowest point distinctly narrower than width of postpedicel. Gena in lateral aspect strikingly narrow, only half as wide as postpedicel, genal setae arranged in two close-set rows. Arista rather longer than antenna measured from base of scape to tip of postpedicel; aristomere 3 thickened on basal quarter. Slender proboscis moderately extended: palp shorter than fore tarsomere 1; prementum longer than fore tarsomere 1 but not as long as fore tarsomeres 1–2, extensively shiny black with only a little dusting on extreme distal part. Thorax extensively covered in very dark, chocolate brown dusting; pattern of darker striping absent on mesonotum. Presutural acrostichal rows in front no farther apart than their distance to adjacent dorsocentral rows; middle pair of acrostichals moderately enlarged; several setulae present between rows. Notopleuron without accessory setulae apart from two notopleural setae. Proepimeral setae and setulae 4–8. Knob of halter bright yellow, contrasting with dark brownish infuscated wing base and calyptrae. Abdomen fairly slender, covered in dark, greyish brown dusting, in oblique posterior aspects revealing a narrow mid-dorsal black stripe and broad, laterally strongly tapering bands on tergites II–V. Terminalia (Figs. 11–13): Most similar to B. minuta (Figs. 18, 19), but cerci longer, apically with submedian prongs almost in line with lateral angles. Female. Parafacial at narrowest point distinctly narrower than width of postpedicel. Gena in lateral aspect very narrow, only about half width of postpedicel, genal setae arranged in two close-set rows. Antennal scape and pedicel ochreous brown, contrasting with black postpedicel. Arista rather longer than antenna measured from base of scape to tip of postpedicel; aristomere 3 thickened on basal quarter. Slender proboscis moderately extended: palp shorter than fore tarsomere 1; prementum longer than fore tarsomere 1 but not as long as fore tarsomeres 1–2, extensively shiny black with only a little dusting on extreme distal part. Thorax covered in light greyish dusting; darker stripes not apparent on mesonotum. Presutural acrostichal rows in front no farther apart than their distance to adjacent dorsocentral rows; middle pair of acrostichals moderately enlarged; several setulae present between rows. Notopleuron without accessory setulae. Proepimeral setae and setulae 1–5. Wing base and calyptrae ochreous-yellow tinged. Knob of halter pale yellow. Abdomen covered in comparatively dense light greyish dusting only allowing a matt, dark subshine. Oviscapt (Fig. 14) similar to that of B. minuta (Fig. 20) in having slender tergite VII pieces and cerci convex along outer margin, but differs in having sternite VI distinctly wider than sternite VII. Material examined. CZECH REPUBLIC [ZMUC]: Ústí nad Labem: Bílina (50.33 N / 13.48 E), 310m, fallows + wood, 1 male 18.vi. 1994, 200m, mixed wood nr. pond, 1 male 1.vi. 1996 (M. Barták); Bílina (50.33 N / 13.50 E), 440m, mixed wood, Malaise trap, 1 male 28.v– 25.vi. 1998 (M. Barták); Bílina (50.32 N / 13.52 E), 380m, mixed wood nr. brook, Malaise trap, 1 male 13–28.v.1998, 1 male 28.v– 15. vi.1998, 1 female 15–25.vi. 1998 (M. Barták). DENMARK [ZMUC]: E Jutland: Hestehave Skov (56.17 N / 10.27 E), 1m, 1 female [paratype] 28.vi. 1974 (V. Michelsen). Møn: Høvblege (54.57 N / 12.30 E), 100m, 1 male 28.v.1983, 2 males 1 female 25.vi. 1999 (V. Michelsen). NE Zealand: Allindelille Fredskov (55.30 N / 11.46 E), 70m, 1 male [holotype] 24.v. 1981 (V. Michelsen); Gladsaxe (55.43 N / 12.28 E), 22m, 1 male 3.vi.1987, 1 male 11.vi.1987, 1 male 22.vi.1987, 1 female 8.vii.1987, 1 female 26.v. 1988 (V. Michelsen). Bornholm: Slotslyngen (55.15 N / 14.44 E), 30m, 1 female [paratype] 26-31.v. 1979 (S. Andersen & V. Michelsen). SLOVAKIA [ZMUC]: Humenné: Ruské (48.54 N / 21.10 E), 370m, sweeping undergrowth in decidious forest, 1 male 3.vi. 1985 (J. Rohácek). SPAIN [ZMUC]: Gerona: Queralbs (42.20 N / 02.09E), 12-1300m, 1 male 13-17.vi. 1982 (Andersen, Lyneborg & Michelsen). SWEDEN [MZLU]: Skåne: Kullaberg (56.17 N / 12.29 E), 1 male 6.vi. 1963 (H. Andersson). Biology. Botanophila helviana prefers sheltered, lush meadows on calcareous soils. Adults of both sexes are anthophilous, feeding on nectar and pollen. As stated earlier, it seems that the larva feed on developing achenes of Ranunculus. Distribution. Known with certainty only from temperate parts of Europe: Britain (Ackland 1989), Czech Republic (Michelsen & Barták 2001), Denmark (Michelsen 1983), France (Ackland 1989), Slovakia (Michelsen 2004), Spain (Michelsen & Báez 2002), Sweden (Michelsen 2004).Published as part of Michelsen, Verner, 2009, Revision of the Botanophila helviana species group (Diptera: Anthomyiidae) in Europe, pp. 45-61 in Zootaxa 2108 on pages 48-50, DOI: 10.5281/zenodo.27486
Spilogona maderensis Michelsen
No full textSpilogona maderensis Michelsen (Figs 14A–C) Spilogona maderensis Michelsen; Michelsen 2021b: 129 ff., figs 3A, B (female, Madeira). Material examined. MADEIRA: [MZH]. Pico Ruivo [32.76N 16.94W], holotype ♀, 27‒29.vi.1957 (H. Lindberg); Ribeira Frio [32.73N 16.88W], 1♁ 14.v.1959 (S. Panelius). Description. Male: Medium sized, wing length 5.2mm (n=1). Head and prementum grey pruinose; entire frons suffused with brown, frontal vitta barely darker than frontal triangle; parafacial plates silvery white with a golden brown tinge. Thorax covered in dense grey pruinosity except extensively dark brown on dorsum of mesonotum including scutellar disc and on upper part of anepisternum. Wing membrane rather intensely smoky brown, yet both cross-veins distinctly darker shadowed; calypteres dirty whitish with brown margins, broadly so on lower calypter; halter yellow. Legs black, femora grey pruinose. Abdomen grey pruinose except for shiny brownish black pattern as follows: entire tergite II except narrowly grey along hind margin, tergites III and IV with very large trapezoid marks narrowly separated by grey median stripe; similar but smaller marks present on tergite V. Postpedicel about 2 times as long as wide, anterior apical corner forming a very obtuse angle; arista short-pubescent, longest branches barely as long as its greatest basal diameter. Haustellum conical, short and tick. Head feminized with eyes widely separated by frons. Frons as well as broad frontal vitta parallel-sided; frontal triangle relatively small, extended to middle of frontal vitta. Frons and lower facial margin slightly and equally projecting. Fronto-orbital plate narrow, with 1 reclinate orbital seta and 3 inclinate frontal setae with a few interstitial setulae and bordered laterally by an irregular row of setulae. Dorsocentral setae 2 + 3; presutural acrostichal field with setulae only in irregular rows. Notopleuron without setulae.Anepisternum without interstitial setula(-e). Katepisternal setae 1 + 2, but lower posterior seta much weaker than both upper ones. Scutellum bare on lateral declivity. Vein C distal to subcostal break with extensive row of dorsal setulae; spines at subcostal break inconspicuous. Fore tibia with 1 p seta below middle. Mid femur without subapical ad seta but with a complete row of pv setae; mid tibia before apex with 2 pd setae only. Hind femur with 4 av setae in distal half and 3-4 pv setae in subbasal third; hind tibia with 1 av and 2–3 ad setae. Abdomen with discal setae confined to tergite V and lateral parts of tergite IV. Sternite I bare. Hypopygium not dissected, but exposed cerci forming a strikingly large shield-like plate with long, attenuate, narrowly separated apical prongs. Taxonomic notes. The male described above is overall very similar to and obviously conspecific with the female holotype of Spilogona maderensis, a species endemic to Madeira. It is easily separated from other species of Spilogona, including the closely related Azorean S. azorensis Michelsen (Michelsen 2021b), by the shadowed cross-veins r-m and dm-cu and the dorsal setulae on vein C. A fully feminized male head with a pair of orbital setae is an exceptional condition among Palearctic and Nearctic species of Spilogona, cf. Sorokina & Pont (2013). However, my expectation will be that the unknown male of S. azorensis likewise possesses a feminized head.Published as part of Michelsen, Verner, 2023, Macaronesian Muscidae (Diptera). IV. The genus Helina Robineau-Desvoidy, and description of the male of Spilogona maderensis Michelsen, pp. 251-276 in Zootaxa 5293 (2) on pages 274-275, DOI: 10.11646/zootaxa.5293.2.3, http://zenodo.org/record/796007
Pegomya curviphallis Michelsen, sp. nov.
No full text<i>Pegomya curviphallis</i> Michelsen, sp. nov. <p>Figs. 1–6, 8–9.</p> <p> ‘ <i>Pegomyia</i> (<i>Pegomyia</i>) <i>flavipalpis</i> (Zetterstedt, 1845) ’; Ringdahl 1938: 208, in part [Skåne, Småland provinces]. [Misidentification.]</p> <p> ‘ <i>Pegomyia flavipalpis</i> (Zetterstedt, 1845) ’; Ringdahl 1952: 172, in part [Småland]; Ringdahl 1959: 254, in part [Småland]. [Misidentifications.]</p> <p> ‘ <i>Pegomya flavoscutellata</i> (Zetterstedt, 1838) ’; Michelsen 1976: 72 [Faunal list: Anholt Is., Denmark]. [Misidentification.]</p> <p> <b>Etymology</b>. The specific epithet ‘curviphallis’ is an adjective derived from the Greek ‘curvus’ and ‘phallus’ and refers to the sickle­shaped distiphallus.</p> <p> <b>Material examined</b>. <i>Type material.</i> Holotype ɗ, SWEDEN: Småland: Bolmen, 24.vii.1942 (O. Ringdahl), MZLU. Paratypes 6ɗɗ: DENMARK: E Jutland: Anholt Is., 1ɗ 7–12.vii.1974 (V. Michelsen), ZMUC. FINLAND: Regio aboensis: Turku, Satava, 1ɗ 1.vii.2005 (K. Winqvist), ZMUC. SWEDEN: Småland: Sandreda, 2ɗɗ 4.vii.1927 (O. Ringdahl), MZLU; Bolmen, 1ɗ 24.vii.1942 (O. Ringdahl), MZLU; Linneryd, 1ɗ 25.vi–5.vii.1980 (V. Michelsen), ZMUC; Norrbotten: Råneå, Högsön, 1ɗ 9.vii.1975 (H. Andersson), MZLU.</p> <p> <i>Other material.</i> FINLAND: Nylandia: Helsingin ymp., 1Ψ 11.vii.1940 (L. Tiensuu), FMNH. SWEDEN: Skåne: Hälsingborg, 1ɗ [postabdomen missing] vi.1921 (O. Ringdahl), MZLU; Småland: Kävsjö, 1Ψ 24.vii.1976 (V. Michelsen), ZMUC; Hästhult, 1Ψ 27.vii.1976 (V. Michelsen), ZMUC; Växjö, S.Åreda, 1Ψ 5.vii.1989 (R. Danielsson), MZLU; Lammhult, Os, 3ΨΨ 13–26.vii.1991 (V. Michelsen), ZMUC; Osby, Åbrolla, 1Ψ 8–14.vii.2004 (V. Michelsen), ZMUC; Värmland: Glaskogen, 1Ψ 26.vi.1978 (V. Michelsen), ZMUC.</p> <p> <b>Description</b>. Very similar to <i>P. flavoscutellata</i> (Zetterstedt) and reliably separated in the male sex only. A differential description of both sexes is attempted in the following:</p> <p> <i>Male.</i> Colour: Pruinosity of thorax and abdomen light grey or bluish grey, sometimes with a faint yellowish stain on dorsum of thorax <i>vs.</i> grey with a yellowish brown stain on dorsum of thorax and abdomen. Abdomen with a dark mid­dorsal stripe that is practically absent on tergites I–II, narrow on tergite III and fully developed only on tergites IV–V <i>vs.</i> extends to base of abdomen and is fully developed on tergites III–V.</p> <p> Terminalia: Distiphallus (Fig. 6) long, forming a subangular arch, long paraphallic processes slender throughout, smooth at posterior edge <i>vs.</i> distiphallus (Fig. 7) shorter, more evenly arched, paraphallic processes wider on basal two­thirds and finely serrated on posterior edge.</p> <p> <i>Female.</i> Colour: Ground colour of mesopleuron a mixture of dark and yellow <i>vs.</i> wholly dark. Pruinosity of thoracic dorsum bright yellowish <i>vs</i>. light grey, partially stained with brownish yellow.</p> <p> Terminalia: Oviscapt (Figs. 8–9) indiscernible from that of <i>P. flavoscutellata</i>.</p> <p> <b>Distribution</b>. The limited available data suggests that <i>P. curviphallis</i> replaces <i>P. flavoscutellata</i> in the lowlands of northern Europe. Still, the distribution ranges of the two species very likely overlap in some places in southern Sweden and Finland.</p>Published as part of <i>Michelsen, Verner, 2006, A new European species of Pegomya RobineauDesvoidy (Diptera: Anthomyiidae) near P. flavoscutellata (Zetterstedt), pp. 49-55 in Zootaxa 1257</i> on pages 51-52, DOI: <a href="http://zenodo.org/record/173100">10.5281/zenodo.173100</a>
Delia sileni Michelsen, 2012, sp. nov.
No full text3. Delia sileni sp. nov. Figs. 1–6, 26– 34. ‘ Delia flavifrons (Zetterstedt) ’; Hennig 1974: 701, 717, 809 (in part: text-fig. 672, plate-figs. 1014 and 1162). Misidentification. Etymology. The specific epithet ‘sileni’ is a Latin genitive derived from ‘Silene’, the generic name of its host plant. Description. The adults differ from other species of the Delia pruinosa species group as follows: Male & female. Differs from D. pruinosa in having hind tibia with apical av-seta normally developed (Fig. 6, arrow). Male. Sternite IV no wider than sternite V and sternites III–V with lateral setae shorter than in D. pruinosa (Figs. 17, 18); surstyli (Figs. 27, 28) more slender in caudal aspect than seen in other species except D. capdellae sp. nov. (Fig. 37), but in that species the surstyli are more tapering in lateral aspect (Fig. 38); distiphallus (Figs. 30, 31), as also in D. pruinosa (Figs. 13, 14), with paraphallic processes same length as postero-median extension. Female. Oviscapt (Figs. 32, 33) rather thick, decidedly longer than preabdomen when fully extended; hypoproct much longer than broad, without cuticular pubescence; cerci with several apical setae of moderate and equal strength. Oviscapt overall very similar to that of D. judicariae (Figs. 24, 25), but distinguished by having the basal membranes of segments VII and VIII covered in tiny, weakly pilose scales arranged in transverse bands (Fig. 33). Material examined. Type material: DENMARK [ZMUC]: S. Zealand: Holtug Kridtbrud, Stevns Klint, holotype 3, paratypes 93, 7 Ƥ, on Silene nutans 5.vi. 1996 (V. Michelsen). Other material: CZECH REPUBLIC [ZMUC]: Moravia: Mrsklesy, 1 Ƥ 15.v. 1974 (Barabás). FINLAND [FMNH, ZMUC]: Regio aboensis: Naantali, 23 [no date], 13 14.vi. 1936, 13 5.vii. 1955 (L. Tiensuu); Turku, 13 18.vii. 1947 (Thuneberg); Satava Is., 13 9.vi. 2005 (K. Winqvist). ITALY: South Tyrol: Stilfser Joch [= Stelvio Pass], 2757m, 2 Ƥ 12.vii. 1997 (S. Andersen). SPAIN [ZMUC]: Santander: Fuenta Dé, 13 18.vi. 1983 (R. Hall); Lerida: 1km S Túnel Viella, 1300m, 13 22.vi. 1982 (Andersen, Lyneborg & Michelsen); Gerona: Queralps, 1200–1300m, 13 13–17.vi. 1982 (Andersen, Lyneborg & Michelsen); 13 11–12.vii. 1984, 23 17–20.vi. 2002, 1Ƥ 10.vi. 2007 (V. Michelsen). TURKEY [ZMUC]: Izmir: Bergama–Kozak rd., 700m, 33, 3 Ƥ 10–12.v. 1993 (V. Michelsen). Distribution. Czech Republic, Denmark, Estonia (Hennig 1974, “Dorpat” [= Tartu]), Finland, Italy, Spain, Turkey.Published as part of Michelsen, Verner, 2012, Revision of the European Delia pruinosa species group (Diptera: Anthomyiidae) feeding as larvae in seed capsules of Silene L. (Caryophyllaceae), pp. 31-48 in Zootaxa 3434 on page 40, DOI: 10.5281/zenodo.28206
Limnophora obsignatula Michelsen 2021, sp. nov.
No full textLimnophora obsignatula sp. nov. (Figs 5A‒C, 6A‒C, 8D‒F, 12C) “ Spilogaster notata (Fallén) ”; Macquart 1839: 115 (Canary Is.). Misidentification. “ Limnophora obsignata (Rondani) ”; Becker 1908a: 102 (Tenerife); Frey 1936: 132 p.p. (Gran Canaria, Tenerife, La Gomera, La Palma); Hennig 1959: 376 (Canary Is. only); Chandler 1979: 120 (La Palma); Santos Abréu 1976: 94 (La Palma); Báez 1981: 628 p. p. (La Gomera?, La Palma). Misidentifications. “ Limnophora notata (Fallén) ”; Santos Abréu 1976: 96 (La Palma). Misidentification. The Canarian records listed above may for the most part represent misidentifications of Limnophora obsignatula sp. nov., an endemic Canarian species replacing the very similar and widespread L. obsignata (see above) in the western islands of Gran Canaria, Tenerife, La Gomera and La Palma. However, the material recorded by Frey (1936) on the authority of the muscid specialist J. Villeneuve even contains specimens of L. quaterna (see below), and the records of L. obsignata by Báez (1981) from La Palma, La Gomera and Fuerteventura refer in part or entirely to L. paneliusi (see below) as evidenced by his illustrations of the male body pattern (pl. I, figs 4‒5) and genitalia (pl. II, figs 1, 3). Material examined. CANARY ISLANDS: Type material [NHMD]: Holotype ♂, TENERIFE: Carrizal Alto, 28°19´N 16°51´W, 640‒730m, 30.i‒3.ii.2016 (V. Michelsen). Paratypes: LA PALMA: Bco. Gallegos, 28°48´N 17°49´W, 470m, 1♂ 17.i.2013 (V. Michelsen). LA GOMERA: La Laja, 28°06´N 17°11´W, 480‒600m, 7♂, 2♀ 23.xi.2002, 4♂, 2♀ 22‒29.i.2016 (V. Michelsen); Los Loros, 28°09´N 17°16´W, 520‒770m, 6♀ 1‒7.ii.2008 (V. Michelsen); Bco. Arure, 28°06´N 17°19´W, 200‒265m, 3♀ 4.ii.2008 (V. Michelsen); Bco de Aguajilva, Lomito Fragoso, 28°07´N 17°10´W, 220‒250m, 6♂, 6♀ 19‒24.i.2019 (V. Michelsen). TENERIFE: Los Carrizales, 28°19´N 16°51´W, c. 645m, 3♂, 2♀ 8‒10.ii.2008 (V. Michelsen); Carrizal Alto, 28°19´N 16°51´W, 640‒730m, ♀ 30.i‒ 3.ii.2016 (V. Michelsen). Other material [MZH, NHMD]: GRAN CANARIA: Los Lagunetas, 3♂, 2♀ 1931 (R. Storå); Moya, 2♂ 1931 (R. Storå). TENERIFE: Agua Garcia, 1♀ 1931 (R. Frey); Las Mercedes, 1♀ 1931 (R. Storå); Valle de Masca, 1♂, 2♀ 12‒13.v.1947 (H. Lindberg). LA GOMERA: Cumbre, 5♂, 6♀ 1931 (R. Frey); Hermigua, 2♂, 5♀ 1931 (R. Frey, R. Storå); Bco de Aguajilva, Lomito Fragoso, 28°07´N 17°10´W, 220‒250m, 3♀ 24‒29.i.2020 (V. Michelsen). LA PALMA: La Caldera, 2♂ 1931 (R. Storå), 1♂, 1♀ 7.viii.1931 (R. Frey); Santa Cruz, 1♂ 1931 (R. Frey). Sixteen specimens (8♂, 8♀) from totally 123 specimens examined showed fungal infection by Stigmatomyces sp., presumably S. limnophorae Thaxt. (Laboulbeniales). Etymology. The specific epithet hints at its probable status as sister species of L. obsignata. Description. Medium sized, wing length 3.9‒5.8mm. Male. Arista (Fig. 6B) short-plumose, longest branches ca. 0.75 times as long as width of postpedicel. Head and body (Figs 5A, B) black, covered in dense, mainly whitish grey pruinosity with a contrasting black “ Anthomyia -pattern” on dorsum of thorax and abdomen. Head (Fig. 6A) more or less brownish pruinose on frons, parafacialia and face; brownish black on upper half of occiput. Frons at narrowest point 0.9‒1.2 times as wide as ocellar tubercle, frontal vitta at this point 1.6‒2.0 times as wide as the fronto-orbital plates; fronto-orbital plate with 1‒2 short reclinate orbital setae immediately above 5‒7 inclinate frontal setae. Frons lying slightly behind moderately produced facial margin. Mesonotal black pattern with pair of large, squarish presutural marks with narrow, wedge-shaped separation disappearing on occipital declivity; band immediately behind the suture very broad, its hind margin almost straight, lying distinctly behind the insertion of third pair of postsutural dorsocentral setae; a narrow median stripe from the postsutural band sometimes extends to the black band on basal two-thirds of scutellum. Dorsocentral setae 4 behind the suture; presutural acrostichal field with setulae only, arranged in four rows separated from adjacent dorsocentral setae by a bare stripe. Calypteres brownish infuscated at the margins; wing membrane faintly brownish black tinged; halter pale yellow; vein M 1 distinctly curved forward in distal part. Mid femur with 2 subapical pd setae and 3‒4 pv setae on basal third; mid tibia with 2(‒3) p setae inserted above and one below middle. Hind femur with 2(‒3) av setae distally, without pv setae. Abdomen (Fig. 5C) with paired blackish marks on tergites I‒II, III and IV and a median brown mark on tergite V; marks on tergites III and IV large, trapezoid, as long as respective tergites and laterally extended approximately to dorsally visible margin of the abdomen. Sternite I bare. Sternite V and hypopygium as in Figs 8D‒F; distinguished from male L. obsignata by the shape of the surstyli. Female. Head (Fig. 6C) on upper two-thirds of frons with a band of brown pruinosity contrasting with light grey vertex and silvery grey lower fronto-orbital and parafacial plates. Fronto-orbital plate with 2 reclinate orbital setae and 5 inclinate frontal setae. Mesonotum with separation between presutural black marks broader, wedge-shaped, posteriorly as wide as acrostichal field. Calypteres paler, dirty whitish. Mid femur basally with 1 pv seta only. Abdomen with paired black marks on tergites III and IV large, trapezoid. Ovipositor is in Fig. 12C, inseparable from that of L. obsignata. See Table 1 for ovipositor characters. Distribution. Apparently endemic to the western Canary Islands. Known so far from Gran Canaria, Tenerife, La Gomera and La Palma.Published as part of Michelsen, Verner, 2021, Macaronesian Muscidae (Diptera). II. The genus Limnophora Robineau-Desvoidy with description of a new Canarian endemic species, pp. 101-127 in Zootaxa 4952 (1) on pages 111-113, DOI: 10.11646/zootaxa.4952.1.6, http://zenodo.org/record/467180
Pegomya prisca Michelsen, sp. nov.
No full textPegomya prisca Michelsen, sp. nov. Figs. 1 –7, 14, 16. ‘ Pegomyia fulviceps (Zetterstedt) ’; Stein 1906: 49 [key], 59 [description]; Stein 1907: 703 in part [catalogue]. Misidentifications. Etymology. The specific epithet ‘prisca’ is a Latin adjective meaning pristine or oldfashioned and refers to the unspecialized female oviscapt. Material examined. Type material. Holotype ♂, SWEDEN: Halland: Simlungsdalen, Fröböke, 17–29.vii. 1994 (V. Michelsen), ZMUC. Paratypes 2 ♂♂, 2 ΨΨ: SWEDEN: Småland, Bolmen, 1 ♂ 20.viii. 1957 (O. Ringdahl), MZLU, 1 ♂ 6.ix. 1958 (O. Ringdahl), MZLU; Gnosjö, 1 Ψ 18.x. 1981 (V. Michelsen), ZMUC; NW Lake Bolmen, 1 Ψ 22.ix. 2002 (V. Michelsen), ZMUC. Other material. CZECH RUPUBLIC: Horní Lomná, 49.32 N 18.37 E, 650m, fly trap, 2 ΨΨ viii–ix. 1986 (Barták), ZMUC; Šumava, Malá Niva, peat bog, 48.55 N 13.49 E, 780m, 1 Ψ 26.ix. 1992 (Barták), ZMUC. SLOVAKIA: TatryTatranský Kotlina, 1km E, meadow, 49.14 N 20.16 E, 800m, 1 Ψ 22.x. 1985 (Barták), ZMUC. Description. Overall very similar to P. testacea (De Geer), but differs as follows: Male. Smaller, wing length (in mm) 6.2–6.9 (n = 3) vs. 6.8–8.2 (n = 41). Colour: Antennal postpedicel black vs. fuscous yellow. Ground colour of scutum yellowish brown, contrasting with yellow scutellum vs. scutum and scutellum concolorous, bright yellow to fuscous greyish yellow. Mid and hind femora yellow, narrowly darkened at tips vs. wholly yellow. Tarsi brown, distinctly darker than yellow femora and tibiae vs. yellow, same color as femora and tibiae. Tergites II–IV yellow, narrowly banded with black at hind margins vs. wholly yellow. Wing: Vein C distal to subcostal break with a regular row of fine dorsal setulae vs. fine dorsal setulae of vein C very few and irregular. Abdomen: Short, depressed abdomen parallel sided vs. ovoid. Terminalia (Figs 1–7 vs. 8–13): Sternite V (Fig. 1 vs. 8) less broad basally, overall more sparsely setose and with pointed apex of lobes membranous rather than sclerotized. Outline of epandrium in caudal view rounded (Fig. 2) vs. subangular (Fig. 9). Cercal plate (Fig. 3 vs. 10) less drawn out apically. Pre and postgonites (Figs. 5–6 vs. 11–12) and phallus (Fig. 7 vs. 13) differently shaped in profile view. Female. Smaller, wing length (in mm) 6.0– 7.5 (n = 6) vs. 6.2–8.6 (n = 76). Colour: Ground colour of scutum yellowish brown, contrasting with yellow scutellum vs. scutum and scutellum concolorous, yellow or (late season only) dark brownish infuscated. Mid and hind femora yellow, narrowly darkened at tips vs. wholly yellow. Tarsi yellowish brown, darker than femora and tibiae vs. yellow, concolorous with femora and tibiae. Tergites II–IV yellow, narrowly banded with black at hind margins vs. wholly yellow or vaguely darker at hind margins (late season only). Wing: Crossvein dmcu rather upright and weakly sigmoid vs. oblique and strongly sigmoid. Terminalia: Oviscapt (Fig. 14 vs. 15) relatively shorter: c. 2.5 mm from fore margin of tergite VI to tip of cerci when wing length = 6.4 mm vs. c. 3.2 mm when wing length = 6.8 mm; sclerotized parts dark brownish vs. light brownish yellow. Distribution. The few existing records of P. p r i s c a from southern Sweden, Czech Republic, Slovakia and? Austria (Stein 1906) lie within the range of P. testacea.Published as part of Michelsen, Verner, 2006, A new European species of Pegomya RobineauDesvoidy (Diptera: Anthomyiidae) near P. t e s t a c e a (De Geer), pp. 37-46 in Zootaxa 1260 on pages 39-42, DOI: 10.5281/zenodo.17317
Helina grancanariae Michelsen 2023, sp. nov.
No full text4. <i>Helina grancanariae</i> sp. nov. <p>(Figs 2D–F, 3D–F)</p> <p> <i>Helina obscurisquama</i> (Stein); Baéz 1983: 180 ff. (p.p., Gran Canaria). Misidentification.</p> <p> <b>Material examined.</b> <b>CANARY ISLANDS:</b> <i>Type material</i> (holotype ♁, paratypes 5♁, 8♀) [NHMD]: GRAN CANARIA: Holotype ♁, paratypes 5♁ 1♀, El Valle, 27°54´N 15°32´W, 730‒935m, 19‒24.i.2017 (V. Michelsen); paratype ♀, 2km W of Soria, 27°54´N 15°40´W, 800m, 28.xi.2014 (V. Michelsen); paratypes 4 ♀, Tasarte, 27°54´N 15°46´W, 270m, 28‒30.xi.2014 (V. Michelsen); paratypes 2♀, Sagrado Corazón, 28°02´N 15°33´W, 700‒825m, 12‒17.i.2017 (V. Michelsen). <i>Other material</i> [MZH]: GRAN CANARIA: Los Lagunetas, 1♀ 26.vi.1931, 2♀ 22.viii.1931 (R. Storå).</p> <p> <b>Description.</b> Medium to large sized, wing length 5.5–8.0 mm. The present species matches in most respects the description of <i>Helina obscurisquama</i> s. str. given above. Accordingly, the following description gives only differences from that species. <i>Male:</i> Abdomen (Fig. 2E) with bluish grey abdominal pruinosity even sparser, largely obscuring the usual paired dark marks on tergites III and IV. Eyes more widely separated by frons (Fig. 2D), distance at narrowest point 2.5‒3 times diameter of anterior ocellus; frontal vitta at this point 1‒2 times as wide as fronto-orbital plate (in <i>H. obscuriscuama</i> frons at this point with contiguous fronto-orbital plates and distance between eyes equals the diameter of anterior ocellus). Hind tibia (Fig. 2F) with 5‒7 long and curved <i>av</i> and 2‒7 similar <i>ad</i> setae supplemented by finer setae and longish setulae in anterior position; preapical dorsal seta erect, often longer than second hind tarsomere. Abdomen (Fig. 2E) relatively slender, ovoid; sternite V and hypopygium (Figs 3D, E) more prominent due to larger hypopygium. <i>Female:</i> Differs from the very similar female of <i>H. obscurisquama</i> by the bluish rather than brownish tinge of the thorax, making it concolorous with the abdomen.</p> <p> <b>Distribution.</b> Endemic to Gran Canaria.</p>Published as part of <i>Michelsen, Verner, 2023, Macaronesian Muscidae (Diptera). IV. The genus Helina Robineau-Desvoidy, and description of the male of Spilogona maderensis Michelsen, pp. 251-276 in Zootaxa 5293 (2)</i> on pages 257-258, DOI: 10.11646/zootaxa.5293.2.3, <a href="http://zenodo.org/record/7960070">http://zenodo.org/record/7960070</a>
Helina santosabreui Michelsen 2023, sp. n.
No full text10. Helina santosabreui sp. n. (Figs 9E, F) Mydaea (Spilogaster) duplicata var. tagojana Santos Abréu; Santos Abréu 1976: 115 ff., fig. 17 (La Palma). [Unavailable infrasubspecific name.] “ Helina quadrum (Fabricius) ”; Báez 1983: 183 ff., pl. 1 fig. 6, pl. 2 figs 1, 7 (La Gomera, La Palma). Misidentification. Material examined. CANARY ISLANDS: Type material (holotype ♁, paratypes 5♀) [NHMD]: LA GOMERA: El Rincón, 1♀ 20.xi.2002 (V. Michelsen); Los Loros, 28°09´N 17°16´W, 500‒770m, 1♀ 1.ii.2008 (V. Michelsen); LA PALMA: Tigalate, 28°32´N 17°48´W, 680m, 2♀ 10‒21.i.2013 (V. Michelsen); Las Tricias, 28°47´N 17°58´W, 660m, holotype ♁, 2♀ 11‒16.i.2013 (V. Michelsen). Other material [MZH]. LA PALMA: supra El Paso, 600m, 1♀ 4.iv.1950 (H. Lindberg). Description. Medium sized, wing length 6.0–7.0mm (n = 8). Male: Head and body mainly dark, densely greyish pruinose, in posterior view with four brownish-black stripes on anterior two-thirds of scutum (interrupted at the suture); abdomen (Fig. 9E) broad-ovoid with paired blackish marks on tergites III and IV in shape of large, almost full-length rectangles; linear brown marks further present in the middle of tergites III–V; cross-veins, or at least r-m, faintly shadowed. Legs dark with ochre-brown mid- and hind tibiae. Eyes bare. Arista plumose, longest branches about same length as width of postpedicel. Frons at narrowest point narrower than ocellar tubercle; fronto-orbital plates at this point narrowly separated by frontal vitta. Dorsocentral setae 2 + 4; presutural acrostical field with setulae only arranged in 4–5 irregular rows; prealar seta very short; katepisternal setae 2 + 2. Fore tibia with 1 p seta; mid tibia with 0 ad and 2–3 p setae; hind femur without pv setae; hind tibia with 2–3 av, 3 ad setae, 1 short pd seta in upper third and in middle third with inconspicuous comb of fine p setae. Terminalia: not examined in the absence of available males other than the holotype, but see Báez (1983, pl. 2, figs 1, 7, as “ H. quadrum ”), showing the cerci forming a cordiform plate as also seen in H. reversio. Female: apart from usual sexual differences of the head and abdomen different from the male by the abdomen (Fig. 9F) having rectangular dark marks on tergites III–IV not quite reaching hind margin of tergites, legs ochre yellow even on fore tibia and tips of mid- and hind femora and hind tibia without row of fine p setae. Distribution. Endemic to the western Canary Islands. Known so far from La Gomera and La Palma. Taxonomic notes. Helina santosabreui seems closely related to the common, widely distributed H. reversio with which it co-exists on La Gomera and La Palma. It differs in both sexes by the slightly stouter body outline, by the paired dark marks on tergites III and IV being larger, rectangular, and by the presence of a p seta on fore tibia.Published as part of Michelsen, Verner, 2023, Macaronesian Muscidae (Diptera). IV. The genus Helina Robineau-Desvoidy, and description of the male of Spilogona maderensis Michelsen, pp. 251-276 in Zootaxa 5293 (2) on page 267, DOI: 10.11646/zootaxa.5293.2.3, http://zenodo.org/record/796007
Pegomya skulei Michelsen, sp. nov.
No full text<i>Pegomya skulei</i> Michelsen, sp. nov. <p>Figs. 32–37.</p> <p> <b>Etymology.</b> The present species is named after Bjarne Skule, a Danish lepidopterist who collected the only known specimen.</p> <p> <b>Description.</b> <i>Male.</i> Small (WL 3.7mm; n=1). Externally indiscernible from <i>P. terminalis</i> except for wholly blackish ground colour of head, body and appendages. Even terminalia (Figs. 32–37) in most respects very similar to <i>P. terminalis</i> apart from shape and vestiture of surstyli which are diagnostic in respect to very short and sharply pointed meso-distal process (Fig. 34) and, as seen in profile (Fig. 35), in shape of mesobasal laminate process, and fewer but stronger ventro-basal setulae.</p> <p> <i>Female.</i> Unknown.</p> <p> <b>Material examined.</b> GREECE [holotype, ZMUC]: Laconia: SW Monemvassia, 1 male 1–7.iv.1983 (B. Skule). The holotype shows signs of having been retrieved from a light trap.</p> <p> <b>Distribution.</b> Known only by the holotype from southern Peloponnese.</p>Published as part of <i>Michelsen, Verner, 2008, The Palaearctic species of the Pegomya terminalis species group (Diptera: Anthomyiidae), with descriptions of two new species, pp. 31-46 in Zootaxa 1781</i> on page 45, DOI: <a href="http://zenodo.org/record/274299">10.5281/zenodo.274299</a>
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