488 research outputs found

    Macarangamyia Elsayed & Tokuda 2018, gen. nov.

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    Macarangamyia Elsayed & Tokuda gen. nov. Type species: Macarangamyia itiokai Elsayed &Tokuda sp. nov. Diagnosis: Macarangamyia belongs to the tribe Asphondyliini because of the following synapomorphies: the female sternite VII is distinctly larger than preceding sternites, and the gonostyli are strongly sclerotized, dorsally situated, short and compact (Tokuda 2012). The tribe Asphondyliini is divided into two subtribes: Asphondyliina and Schizomyiina (Gagné & Jaschhof 2017). The new genus, Macarangamyia, belongs to the subtribe Schizomyiina because of the presence of unfused teeth of gonostyli (Tokuda 2012). Macarangamyia can be distinguished from other genera of Schizomyiina, such as the Oriental genera Asphoxenomyia Felt and Luzonomyia Felt, by the following combination of characters: palpi four-segmented; male flagellomeres with short necks; tarsal claws untoothed; ovipositor short, membranous, protrusible, with scattered strong setae ventrally and dorsally; female cerci fused, with some blunt-tipped setae; aedeagus cylindrical, with slit developed dorsally more than ventrally; larva with bidentate spatula; a pair of spiracles present on all thoracic and abdominal segments of larva, except the terminal segment. Description. Adult: Head (Fig. 2): Compound eyes with octagonal facets. Mouthparts: palpi four-segmented; labrum triangular; labella well-developed. Antenna: flagellomeres cylindrical, with short necks; first and second flagellomeres partially fused; female flagellomeres with two connected rings of circumfila, except the terminal flagellomere with network-like circumfila, distal female flagellomeres successively shorter (Fig. 3); male flagellomeres with sinuous circumfila (Fig. 4). Thorax: Wing (Fig. 5): R1 joining C before wing midlength, arculus present, R5 joining C slightly after wing apex, C broken after the conjunction with R5; M3+4 forked with Cu. Tarsomeres I simple (Fig. 6); tarsal claws (Fig. 7) curved, untoothed on all legs; empodia shorter than claws, covered with longer setulae apically than basally; pulvilli short. Female abdomen: Tergites I–VII rectangular, with one posterior row of setae and some lateral setae; tergite VIII bare, notched laterally, posterior margin with a pair of well-developed dorsal lobs; trichoid sensilla absent from all tergites. Sternites II–VI each with several scattered setae anteriorly, one posterior row of setae; sternite VII elongated, completely sclerotized; posterior two thirds covered with scattered setae; no discernible trichoid sensilla on all sternites. Ovipositor (Figs. 8–9): protrusible, unpigmented, without microtrichiae, with scattered strong setae ventrally and dorsally; cerci tiny, fused, each bearing pair of thick blunt-tipped setae and several finer setae. Male abdomen: Tergites I–VII as in female; tergite VIII bare, with median part longer and thicker than the lateral parts. Sternites II–VI as in female; sternite VII and VIII with several scattered setae (Fig. 10). Terminalia (Fig. 11–12): Gonocoxite massive, produced ventroapically. Gonostylus with unfused denticles. Cerci separated by V-shaped emargination. Hypoproct bilobed, with one seta at tip of each lobe. Parameres well-developed. Aedeagus cylindrical, with slit developed dorsally more than ventrally. Larva: Sternal spatula bidentate. One group of lateral papillae on each side of spatula, with two setose and one asetose papillae. Two setose sternal papillae, two setose dorsal papillae, and a pair of spiracles present on all thoracic segments and abdominal segments I–VIII. Abdominal terminal segment elongated and tapered. Pupa (Figs. 14–15): Exuviae not pigmented except the antennal and facial horns and prothoracic spiracle. Antennal and facial horns well-developed, each horn bidentate; two facial horns present, pointed. Prothoracic spiracle arched, pointed apically. Scutum medially without wrinkles surrounding the ecdysal line. Abdominal tergites without dorsal spines. Abdominal spiracles present on abdominal segments II–VI. Etymology: The generic name is derived from the host-plant generic name, Macaranga, and the Greek noun myia, meaning fly. The gender is feminine.Published as part of Elsayed, Ayman Khamis, Shimizu-Kaya, Usun, Itioka, Takao, Meleng, Paulus, Yukawa, Junichi & Tokuda, Makoto, 2018, A new genus and a new species of Schizomyiina (Diptera: Cecidomyiidae: Asphondyliini) inducing petiole galls on Macaranga bancana (Miq.) in Borneo, Malaysia, pp. 188-196 in Zootaxa 4482 (1) on pages 190-193, DOI: 10.11646/zootaxa.4482.1.10, http://zenodo.org/record/144047

    FIGURES 26–28 in Two new species of Schizomyia (Diptera: Cecidomyiidae) from Japan, with an updated key to larval, pupal and adult Schizomyia in Japan

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    FIGURES 26–28. Larva of Schizomyia uechiae n. sp. 26. Ventral view of prothoracic segment. 27. Terminal larval segments dorsally. 28. Ventral view of abdominal segment VIII and anus. Scale bars = 50 µm.Published as part of Elsayed, Ayman Khamis, Yukawa, Junichi & Tokuda, Makoto, 2019, Two new species of Schizomyia (Diptera: Cecidomyiidae) from Japan, with an updated key to larval, pupal and adult Schizomyia in Japan, pp. 348-360 in Zootaxa 4688 (3) on page 357, DOI: 10.11646/zootaxa.4688.3.2, http://zenodo.org/record/351678

    Discovery of a new host plant of Pseudasphondylia tominagai Elsayed & Tokuda, 2019 (Diptera, Cecidomyiidae)

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    Barely visible flower bud galls of Deutzia floribunda Nakai (Hydrangeaceae) were found in Kumamoto Prefecture, Kyushu, Japan. Morphological and molecular analyses indicated that the gall inducer is Pseudasphondylia tominagai Elsayed & Tokuda, 2019 (Diptera, Cecidomyiidae), previously known only to form galls on Eleutherococcus spinosus (L.f.) S.Y.Hu (Araliaceae) in Honshu, Japan. This study marks the first report of a species of Pseudasphondylia Monzen, 1955 on Hydrangeaceae, having host plants from different families and establishing D. floribunda as a new host record for P. tominagai

    FIGURE 21 in Description of a new and redescriptions of two known species of Tor ym u s (Hymenoptera: Torymidae) in Taiwan with a key to Taiwanese species

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    FIGURE 21. Female of Torymus orientalis.Published as part of Matsuo, Kazunori, Yang, Man - Miao, Tung, Gene - Sheng, Tokuda, Makoto & Yukawa, Junichi, 2012, Description of a new and redescriptions of two known species of Tor ym u s (Hymenoptera: Torymidae) in Taiwan with a key to Taiwanese species, pp. 47-57 in Zootaxa 3409 on page 54, DOI: 10.5281/zenodo.28188

    Illiciomyia Tokuda,a New Genus for Illiciomyia yukawai sp.n.(Diptera : Cecidomyiidae : Asphondyliini) Inducing Leaf Galls on Illicium anisatum (Illiciaceae) in Japan

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    A new genus, Illiciomyia Tokuda, is erected for Illiciomyia yukawai sp. n. that induces leaf galls on illicium anisatum (Illiciaceae) in Japan. The new genus belongs to the subtribe Asphondyliina (Diptera: Cecidomyiidae :Asphondyliini) and can be distinguished from other genera by the lack of an apical spur on the first tarsomere on all legs and the shallow construction of male flagellomeres. In the subtribe Asphondyliina, the genus is considered to have a close relation to the genera Bruggmanniella and Pseudasphondylia by having common characters such as two separate teeth of gonostylus and transverse rows of rather long spines on the anterior half of dorsal surface of pupal abdominal segment. However, the paramere-like structure of Illiciomyia is selerotized, while paramere of Bruggmanniella and Pseudasphondylia is not sclerotized. These differences indicate that Illiciomyia is phylogenetically apart from Bruggmanniella and Pseudasphondylia

    FIGURES 10–12. Macarangamyia itiokai n in A new genus and a new species of Schizomyiina (Diptera: Cecidomyiidae: Asphondyliini) inducing petiole galls on Macaranga bancana (Miq.) in Borneo, Malaysia

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    FIGURES 10–12. Macarangamyia itiokai n. sp. 10. Terminal part of male abdomen, excluding male terminalia. 11. Male terminalia. 12. Ventral view of aedeagus. Scale bars = 50 µm.Published as part of Elsayed, Ayman Khamis, Shimizu-Kaya, Usun, Itioka, Takao, Meleng, Paulus, Yukawa, Junichi & Tokuda, Makoto, 2018, A new genus and a new species of Schizomyiina (Diptera: Cecidomyiidae: Asphondyliini) inducing petiole galls on Macaranga bancana (Miq.) in Borneo, Malaysia, pp. 188-196 in Zootaxa 4482 (1) on page 192, DOI: 10.11646/zootaxa.4482.1.10, http://zenodo.org/record/144047

    A taxonomic revision and molecular phylogeny of the eastern Palearctic species of the genera Schizomyia Kieffer and Asteralobia Kovalev (Diptera, Cecidomyiidae, Asphondyliini), with descriptions of five new species of Schizomyia from Japan

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    The genus Asteralobia (Diptera, Cecidomyiidae, Asphondyliini, Schizomyiina) was erected by Kovalev (1964) based on the presence of constrictions on the cylindrical male flagellomeres. In the present study, we examine the morphological features of Asteralobia and Schizomyia and found that the male flagellomeres are constricted also in Schizomyia galiorum, the type species of Schizomyia. Because no further characters clearly separating Asteralobia from Schizomyia were observed, we synonymize Asteralobia under Schizomyia. Molecular phylogenetic analysis strongly supports our taxonomic treatment. We describe five new species of Schizomyia from Japan, S. achyranthesae Elsayed & Tokuda, sp. n., S. diplocyclosae Elsayed & Tokuda, sp. n., S. castanopsisae Elsayed & Tokuda, sp. n., S. usubai Elsayed & Tokuda, sp. n., and S. paederiae Elsayed & Tokuda, sp. n., and redescribe three species, S. galiorum Kieffer, S. patriniae Shinji, and S. asteris Kovalev. A taxonomic key to the Japanese Schizomyia species is provided

    FIGURES 46–52. Ametrodiplosis stellariae n in Three new species of Ametrodiplosis (Diptera: Cecidomyiidae) from Japan, with a key to the Japanese species and a molecular phylogenetic analysis

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    FIGURES 46–52. Ametrodiplosis stellariae n. sp. 46. Head (left paplus is incomplete). 47. Ventral view of mouthparts. 48. Female antennal flagellomeres. 49. Male antennal flagellomeres. 50. Dorsal view of female flagellomere V. 51. Ventral view of male flagellomere V. 52. Tarsomere V and acromere. Scale bars = 50 µm.Published as part of Elsayed, Ayman Khamis, Yukawa, Junichi, Mochizuki, Ko, Tokuda, Makoto & Kawakita, Atsushi, 2021, Three new species of Ametrodiplosis (Diptera: Cecidomyiidae) from Japan, with a key to the Japanese species and a molecular phylogenetic analysis, pp. 151-172 in Zootaxa 4942 (2) on page 166, DOI: 10.11646/zootaxa.4942.2.1, http://zenodo.org/record/460039

    Illiciomyia Tokuda,a New Genus for Illiciomyia yukawai sp.n.(Diptera : Cecidomyiidae : Asphondyliini) Inducing Leaf Galls on Illicium anisatum (Illiciaceae) in Japan

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    A new genus, Illiciomyia Tokuda, is erected for Illiciomyia yukawai sp. n. that induces leaf galls on illicium anisatum (Illiciaceae) in Japan. The new genus belongs to the subtribe Asphondyliina (Diptera: Cecidomyiidae :Asphondyliini) and can be distinguished from other genera by the lack of an apical spur on the first tarsomere on all legs and the shallow construction of male flagellomeres. In the subtribe Asphondyliina, the genus is considered to have a close relation to the genera Bruggmanniella and Pseudasphondylia by having common characters such as two separate teeth of gonostylus and transverse rows of rather long spines on the anterior half of dorsal surface of pupal abdominal segment. However, the paramere-like structure of Illiciomyia is selerotized, while paramere of Bruggmanniella and Pseudasphondylia is not sclerotized. These differences indicate that Illiciomyia is phylogenetically apart from Bruggmanniella and Pseudasphondylia

    FIGURE 63. A in Three new species of Ametrodiplosis (Diptera: Cecidomyiidae) from Japan, with a key to the Japanese species and a molecular phylogenetic analysis

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    FIGURE 63. A Maximum Likelihood phylogenetic tree based on concatenated COI, 16S and 28S partial sequences. Terminal labels for Ametrodiplosis spp. include the locality name. All Ametrodiplosis aeroradicis individuals were collected from Trachelospermum asiaticum, except one specimen from T. gracilipes var. liukiuense in Kagoshima (indicated by an asterisk).Published as part of Elsayed, Ayman Khamis, Yukawa, Junichi, Mochizuki, Ko, Tokuda, Makoto & Kawakita, Atsushi, 2021, Three new species of Ametrodiplosis (Diptera: Cecidomyiidae) from Japan, with a key to the Japanese species and a molecular phylogenetic analysis, pp. 151-172 in Zootaxa 4942 (2) on page 169, DOI: 10.11646/zootaxa.4942.2.1, http://zenodo.org/record/460039
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