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E. Neubert, La doctrine mariale de M. Cheminade, 1937
No full textRivière Jean. E. Neubert, La doctrine mariale de M. Cheminade, 1937. In: Revue des Sciences Religieuses, tome 19, fascicule 1, 1939. p. 142
E. Neubert, La doctrine mariale de M. Cheminade, 1937
No full textRivière Jean. E. Neubert, La doctrine mariale de M. Cheminade, 1937. In: Revue des Sciences Religieuses, tome 19, fascicule 1, 1939. p. 142
Model-independent properties of the B-meson distribution amplitude
No full textThe operator product expansion is used to obtain model-independent predictions for the first two moments of the renormalized B-meson light-cone distribution amplitude phi(+)(B)(omega,mu), defined with a cutoff omega <=Lambda(UV). The leading hadronic power corrections are given in terms of the parameter <(Lambda)over bar>=m(B)-m(b). From the cutoff dependence of the zeroth moment an analytical expression for the asymptotic behavior of the distribution amplitude is derived, which exhibits a negative radiation tail for omega >mu. By solving the evolution equation for the distribution amplitude, an integral representation for phi(+)(B)(omega,mu) is obtained in terms an initial function phi(+)(B)(omega,mu(0)) defined at a lower renormalization scale. A realistic model of the B-meson light-cone distribution amplitude is proposed, which satisfies the moment relations and has the correct asymptotic behavior. This model provides an estimate for the first inverse moment and the associated parameter lambda(B).
Multidentula reducta Ruud, Menkhorst & Neubert, 2016, spec. nov.
No full textMultidentula reducta spec. nov. (Fig. 19) Type locality & type specimens. – Turkey, Vilayet Sivas, Zara, 16 km towards Imranlı, 1400 m (39.8606°N 37.8765°E), H.P.M.G. Menkhorst leg., 15.vii.1988. Holotype NMBE 544681, paratypes NMBE 544682/1, HMK/12, RBA/1, RMNH/1. Diagnosis. – A middle-sized, oval Multidentula species with fine oblique striae, a missing suprapalatalis and a weakly developed or missing basalis, an oblique infrapalatalis, a non-fused subangularis, and a parietalis that is only slightly curved. Description. – Shell dextral, oval in outline, with a rather wide, open, slit-like umbilicus. The 6.0-7.2 whorls are convex with a rather deep suture. Teleoconch with irregular, sparse, oblique striae which are mostly rather fine; there are no spiral striae. Shell rather solid, slightly translucent, yellowish to horny yellowish, with a whitish band behind the peristome. The last whorl has no or only an indistinct impression and only the infrapalatalis is visible outside as a blurred white stripe. Peristome well reflected, thickened by a prominent labial callus, the columellar and palatal insertion connected by a clearly visible callus (but the callus is not extra thickened near the insertions). The subangularis is vertically pointing downwards, and is connected (but not fused) with the palatal peristome by a callus. The subangularis and parietalis are connected by a thin callus. Parietalis prominent, slightly curved and deeply recessed. A spiralis is missing. Columellaris ± oblique to the columellar peristome, deeply recessed. The basalis is absent, or present as an indistinct thickening only. Infrapalatalis well developed, oblique, and always stronger developed than the non-oblique palatalis superior. There is no suprapalatalis; a small suturalis is present. Measurements (n = 6). – H = 5.3-6.9 (mean 6.1); LWH = 3.2-3.8 (mean 3.5); MH = 2.1-2.4 (mean 2.2); LWD = 2.7-2.8 (mean 2.8); LWM = 2.7-3.0 (mean 2.8); MD = 1.7-1.9 (mean 1.8); NW = 6.0-7.2 (mean 6.6). Localities. – Known from the locus typicus only (see above). Derivatio nominis. – The name refers to the reduced apertural armature. Differentiation. – The armature of M. reducta is generally reduced: a basalis is absent or only weakly developed, a suprapalatalis is missing, and the parietalis is not as prominent and less curved as in M. squalina or M. pupoides. The palatal folds are, compared to M. squalina, also less deep in the aperture, but are placed closer to the peristome.Published as part of Ruud, A. Bank, Henk P. M. G Menkhorst & Eike Neubert, 2016, Descriptions of new and little-known land snail taxa from Turkey, and establishment of a new genus (Gastropoda, Pulmonata: Lauriidae, Enidae and Vitrinidae), pp. 5-30 in Basteria 80 (1) on page 23, DOI: 10.5281/zenodo.43974
Figs 3–6. Lindholmiola lens. Fig. 3 in Revision of the genus Lindholmiola HESSE, 1931 (Gastropoda: Pulmonata: Helicodontidae)
No full textFigs 3–6. Lindholmiola lens. Fig. 3: syntype Helix lentiformis, ZMZ 508656, "Thessalie et Attique", D = 11.2 mm; Fig. 4: NMBE 515542, Attika, Methana, 500 m before village, 12.07.1996, leg. E. Neubert, D = 12.77 mm; Fig. 5: syntype Helix lens var. callojuncta, NHMG 372, D = 12.0 mm; Fig. 6: syntype Helix lens var. elia, SMF 101961, Kumani, Elis, coll. Jetschin ex Boettger, 1883, D = 11.94 mm. — All photos Neubert/Bochud, × 3.Published as part of Subai, Peter & Neubert, Eike, 2014, Revision of the genus Lindholmiola HESSE, 1931 (Gastropoda: Pulmonata: Helicodontidae), pp. 1-94 in Contributions to Natural History 23 on page 11, DOI: 10.5169/seals-787037, http://zenodo.org/record/584234
Amphiscopus sturmii subsp. marmoratus Ruud, Menkhorst & Neubert, 2016, subspec. nov.
No full textAmphiscopus sturmii marmoratus subspec. nov. (Fig. 15) Type locality & type material. – Turkey, Vilayet Kütahya, Örencik, western part, 950 m (39.4426°N 29.0662°E), B. Hausdorf leg., 21.ix.1987. Holotype NMBE 544670/1, paratypes NMBE 54467/5, HMK/2, RBA/*2, ZMH/33. Diagnosis. – A subspecies of A. sturmii characterized by the whitish/ marbled colour of the shell, the less well-developed parietal callus and the absence of an impression at the last whorl. Description. – Shell dextral, turreted, with a relative wide, open, slit-like umbilicus. The 8.3-10.1 whorls are rather convex with a moderately deep suture. Teleoconch with densely packed, rather pronounced, oblique striae; there are no spiral striae. Shell solid, not translucent, glossy, upper part of the teleoconch horny yellow coloured, the remaining part of the teleoconch whitish/ marbled coloured, but below the periphery (i.e. the part of the whorl below the insertion of the palatal peristome) again more horny yellow; there is a prominent white band behind the peristome. The last whorl does not have an impression near the palatalis superior. Peristome not or hardly reflected (with the exception of the columellar part), robustly thickened by a labial callus, the columellar and palatal insertion connected by a clearly visible callus which is often more thickened near the columellar peristome. The subangularis is tear-like and vertically pointing downwards; it is connected with the palatal peristome by a thin callus. The subangularis and parietalis are not connected. The parietalis is well developed, and only moderately deeply recessed. Palatalis superior well thickened, but not deeply recessed. There is no spiralis, infrapalatalis, basalis or columellaris. The columellar ledge reaches halfway or above the middle of the columellar side of the aperture. Measurements (n = 11). – H = 6.7-9.0 (mean 7.8); LWH = 3.0-3.5 (mean 3.3); MH = 1.9-2.2 (mean 2.1); LWD = 2.3-2.7 (mean 2.4); LWM = 2.3-2.8 (mean 2.6); MD = 1.5-1.8 (mean 1.6); NW = 8.3-10.1 (mean 9.2). Localities. – Vilayet Kütahya: type locality (see above); 4 km E. Köprüören, 1020 m (39.5115°N 29.8137°E) (ZMH/>50). Derivatio nominis. – Named after the marble-like colour of the shell. Differentiation. – Amphiscopus sturmii sturmii has a uniform dark-brown coloured shell, more convex whorls, a more prominent parietal callus, and the last whorl has a more or less faint impression at the position of the palatalis superior. Amphiscopus substurmii is sinistral and has the same colour pattern as A. sturmii sturmii.Published as part of Ruud, A. Bank, Henk P. M. G Menkhorst & Eike Neubert, 2016, Descriptions of new and little-known land snail taxa from Turkey, and establishment of a new genus (Gastropoda, Pulmonata: Lauriidae, Enidae and Vitrinidae), pp. 5-30 in Basteria 80 (1) on page 20, DOI: 10.5281/zenodo.43974
Lanistes thaytinitiensis Neubert & Damme 2012, sp. nov.
No full text<i>Lanistes thaytinitiensis</i> sp. nov. (Fig. 7) <p>Type specimens: Holotype NMBE 5018966; paratypes NMBE 5018967–5018969, MNHN/2, SMF 340185 /2.</p> <p>Type locality: Thaytiniti, Dhofar, Oman.</p> <p>Stratum typicum: Paludal biomicritic limestones of the Zalumah Formation.</p> <p>Age: Late Priabonian.</p> <p>Material: This species was only recovered from the Zalumah exposures at Thaytiniti. Collected by M. Pickford during one of the palaeontological missions led by H. Thomas in 1988–1990.</p> <p>Etymology: named after the type locality.</p> <p> Diagnosis: A palaeogene species of <i>Lanistes</i> characterized by the discoid shape and flattened upper side.</p> <p> Description: Medium sized <i>Lanistes</i>; discoid shell consisting of 4 <i>½</i> whorls with virtually no exserted spire, apex pointed; upper part of the gradually increasing whorls completely flattened, lower part convex, forming a peripheral angle accentuated by a ridge; at the base the periphery of the whorls form a second angle with the flattened walls of the umbilicus, in which the whorls wind downwards (or upwards being hyperstrophic) in a regular spiral; aperture horizontal at the top, the outer margin convex, the upper part of the inner margin, connected to the shell, concave and the free lower part straight, basal margin pointed; growth lines regular and fine, no spiral sculpture; operculum unknown.</p> <p>Measurements: Holotype (Fig. 7): H = 12.5 mm; W = 38.2 mm; h = 12.0 mm; w = 11.5 mm. Paratype (NMBE 5018967): H = 11.2 mm; W = 32.2 mm; h = 11.2 mm; w = 10.3 mm.</p> <p> Remarks: We did not find any fossils of this species in the Salalah region, while in the Zalumah exposures at Thaytiniti is seems to be the most common of the two <i>Lanistes</i>. This indicates either a difference in age or in environmental conditions between these two sites. <i>Lanistes thaytinitiensis</i> sp. nov. is the only flattened discoid <i>Lanistes</i> species known.</p>Published as part of <i>Neubert, Eike & Damme, Dirk van, 2012, Palaeogene continental molluscs of Oman, pp. 1-28 in Contributions to Natural History 20</i> on pages 9-10, DOI: 10.5169/seals-787080, <a href="http://zenodo.org/record/5838544">http://zenodo.org/record/5838544</a>
Helix borealis sensu Neubert 2014
No full textAPPENDIX 2. SYSTEMATICS, DISTRIBUTION, VARIABILITY AND ECOLOGY OF HELIX BOREALIS Helix borealis is generally poorly known, including conchological variability and its potential taxonomic significance. Most of the published accounts are in taxonomic compilations, often repeating earlier information, supplemented by faunistic data (see below for an overview of the literature). The biology and ecology of this taxon has been mostly neglected. Notable exceptions to this pattern are Hesse (1920), providing information on soft body morphology, and Welter-Schultes (1998a), who discussed its phenology and past distribution in Crete and on nearby islands. Kobelt (1895) distinguished, besides typical H. borealis, two more Greek taxa currently considered its synonyms. According to his description, Helix thiesseana Kobelt, 1878, described from Evvia, should have a more globular shell with darker palatal callus than the nominotypical form and almost missing spiral sculpture. The name H. thiesseana has been sometimes used not only for populations from Evvia, but also for individuals with darker apertures from the Peloponnese peninsula. Helix aetolica Kobelt, 1892 from Aetolia, western Greece (name invalid due to primary homonymy), should be larger with broader shell, darker coloration and less developed spiral sculpture than the nominotypical subspecies. The Turkish populations were never formally described. SYNONYMY OF HELIX BOREALIS Helix cincta – d’Audebard de Férussac AEJPJF, 1821– 1822: 29 (quarto edition) [partim: La Gréce, l’Archipel; l’ile de Zante] Helix cincta – Deshayes, 1835: 160 [partim: de Morée] Helix ambigua Mousson, 1859: 15, non Helix ambigua Linnaeus, 1758 (presently Fossarus ambiguus) [de la Grèce et de la Thessalie …de Corfou et de Céfalonie se retrouve avec toutes ses particularités sur toute la côte de l’Epire, tant à Sayades qua’à Prevesa] Helix ambigua var. borealis Mousson, 1859: 16 [Ile de Corfou…das les broussailles des rochers de la citadelle] Helix cyrtolena Bourguignat, 1860: 165 [nom. nov. for Helix ambigua Mousson, 1859] Helix (Pomatia) Thiesseana Kobelt, 1878: 320 [bei Chalcis auf Euböa] Helix Thiesseana – Kobelt W, 1879–80: 1, pl. 179, figs 1805–1806 [bei Chalcis auf Euböa] Helix cincta – Westerlund & Blanc, 1879: 79 [Pylos à Navarin, Pyrgos en Elide et dans l’Arcadie] Helix cincta Var. ambigua – Westerlund & Blanc, 1879: 79 [Iles de Céfalonie et d’Ithaque] Helix thiesseana – Westerlund & Blanc, 1879: 80 [Ile d’Eubée partie boréale] Helix Thiesseana – Godet, 1880: 25 [Chalcis (Euboea)] Helix ambigua – Hesse, 1882: 322 [auf Corfu an der strasse nach Castrades, und auf Zante an der Citadelle] Helix (Helicogena) ambigua – Boettger, 1883: 317 [Corfu] Helix (Helicogena) ambigua var. Thiesseae – Boettger, 1883: 329 [Patras; incorrect subsequent spelling of Helix thiesseana Kobelt, 1878] Helix (Helicogena) ambigua var. Thiesseae – Boettger, 1885: 118 [Achaia, Santameri] Helix [Pomatia] ambigua – Tryon & Pilsbry, 1888: 244, pl. 69, fig. 30 [Corfu, Cephalonia] H e l i x [Po m a t i a] a m b i g u a Va r. t h i e s s e a n a – Tryon & Pilsbry, 1888: 244, pl. 69, fig. 31 [Achaia; Chalcis, Euboea] Helix ambigua – Westerlund, 1889: 458 [Griechenland auf Corfu, Cephalonia, Zante, Ithaka]. Helix ambigua Forma clathrata Westerlund, 1889: 459 [Corfu u. Epirus] Helix thiesseana – Westerlund, 1889: 459 [Griechenland bei Chalkis auf Euboea]. Helix (Pomatia) ambigua – Kobelt W, 1891–92: 24, pl. 127, fig. 766 [in Griechenland und Epirus, sowie auf den jonischen Inseln] Helix (Pomatia) ambigua var. aetolica Kobelt W, 1891 –92: 106, pl. 146, figs. 936–937, non Helix (Macularia) Codringtoni var. Aetolica O. Boettger, 1888 (currently Codringtonia parnassia Roth, 1855) [Vrachori in Aetolien; on the plate erroneously labelled as Helix ambigua var. acarnanica] Helix ambigua – Schuberth, 1892: 51, pl. 5, fig. 18 [Corfu] Helix (Pomatia) ambigua – Kobelt W, 1893–97: 778, pl. 215, figs 1–2 [auf der jonischen Inseln, in Epirus und Nordgriechenland] Helix (Pomatia) ambigua var. aetolica – Kobelt W, 1893–97: 778, pl. 215, figs 2 [Vrachori] Helix (Pomatia) thiesseana – Kobelt W, 1893–97: 779, pl. 215, figs 3–4 [bei Chalkis auf Euböa] Helix (Pomatia) thiesseana – Sturany, 1902: 405 [Kalavryta, 800 m Höhe] Helix (Helicogena) ambigua – Kobelt W, 1902–06: 117, pl. 323, figs 1–4 Helix (Helicogena) ambigua thiesseana – Kobelt W, 1902–06: 118, pl. 215, figs 3–4 Helix (Pomatia) ambigua – Sangiorgi, 1903: 94 [Cefalonia, comune] Helix (Helicogena) ambigua thiesseana – Hesse, 1920: 183, pl. 654, figs 6–7 [Patras, Cerigo] Helix (Helicogena) ambigua – Hesse 1915-1920: 251 [Griechenland, Jon. Inseln, Kreta] Helicogena cincta subsp. ambigua – Käufel, 1930: 185 [Korfu, Levkas, Kephalonia] Helix cincta Rasse ambigua – Knipper, 1939: 369 Helix (Helicogena) cincta ambigua – Käufel & Fuchs, 1941: 201 [Zante: Maries, Keri, Skopos] Helix cincta ambigua – Zilch, 1952: 150 Helix (cincta thiesseana) – Zilch, 1952: 150 Helix cincta ambigua – Klemm, 1962: 255 [Levkas: Frini, Olivenhain] Helix (Helix) cincta ambigua – Rähle, 1980: 218 [Kephallinia, Zakynthos] Helix (Helix) cincta ambigua – Liebegott, 1986: 22 [Skopelos, Kira Panagia, Giura, Piperi] Helix cincta ambigua – Rähle, 1986: 6 [Ithaki] Helix (Helix) cincta borealis – Hausdorf, 1993: 45 Helx (Helix) cincta ambigua – Frank, 1997: 141 Helix cincta – Facorellis et al., 1998: 965 [mesolithic: Gioura] Helix (Helix) cincta – Welter-Schultes, 1998a: 100 [Crete: Anopoli, Alikampos; Gavdos, Gavdopoula] Helix cincta borealis – Neubert et al., 2000: 113 [Turkey: between Kaş and Demre] Helix cincta – Triantis et al., 2008: 475 [island group of Skyros] Helix cincta – Welter-Schultes, 2012: 611 [partim] Helix cincta cincta – Psonis et al., 2015: 383 [partim: Crete: Anopoli, Alikampos; Skyros; Peloponnese: Skollis mountain; Kerkyra: Pantokratoras mountain] Helix borealis – Neubert, 2014: 98 Helix borealis – Korábek et al., 2015 Helix borealis – Neubert & Korábek, 2015 DISTRIBUTION OF HELIX BOREALIS Helix borealis has a fragmented distribution. Its main range lies in the Peloponnese, south-western Pindus and the Ionian coast and Islands. On Evvia it is restricted to the north of the island (Neubert, 2014), unless it also lives near Chalkis, as stated in the original description (Kobelt, 1878). It was collected live on Skopelos and Skyros in the northern Sporades. Liebegott (1986) reports it only subfossil from Kyra Panagia, Gioura and Piperi in the northern Sporades; also Facorellis et al. (1998) refers to old shells (older than 7700 BC) from Gioura. On Crete, the species is apparently rare; only two confirmed localities and one probable were reported (Welter-Schultes, 1998a; Psonis et al., 2015). On the islands of Gavdos and Gavdopoula, south of Crete, it is most likely extinct. Shells dated to c. 6000–25 000 BC by radiocarbon analysis demonstrate the autochthonous origin of the species on these islands. However, they also indicate that the species may have been extinct for a long time already (Welter-Schultes, 1998a). In Anatolia, the distribution is broader than indicated by Neubert (2014). The species has been found in the province of Antalya between Kaş, Finike and Kemer (Neubert et al., 2000; own data); the known sites are listed in Table 2. PHENOTYPIC DIVERSITY OF H. BOREALIS The variation in conchological characters exhibits some geographic structure, which can be partly identified with the taxa distinguished by Kobelt (1895), partly the varieties have not been formally described. On Corfu and in the vicinity of Igoumenitsa lives a form identifiable with typical H. borealis (Supporting Information, Fig. S1D). Usually, the three upper bands are separated and well visible on the top of the shell, and the aperture has an ochre, rather than brown, colour. Towards the south (Acarnania and the islands of Lefkada, Kefalonia, Zakynthos) the shell colour is often pale without bands, but with a slightly darker upper half of the shell (Fig. 1C; Supporting Information, Fig. S1E). The colour of the aperture is similar and the coloration is often well developed only on the palatum and upper columella. These two forms were not distinguished from each other by the mitochondrial phylogeny, but together form a distinctive clade sister to the next form (Fig. 3). Snails similar to the syntypes of H. aetolica live in the northern Peloponnese, Aetolia, Evrytania, and marginally also in western Thessaly (Fig. 1E; Supporting Information, Fig. S1F). They usually have a regularly rounded shell with developed, but often inconspicuous, bands. The upper three usually fuse and their colour does not contrast much with that of the background. The aperture margins are completely dark-coloured. Younger individuals are sometimes covered by brown periostracum, which peels off rapidly, but the periostracum seems to be well developed only at more humid sites. Spiral sculpture varies but is often developed and relatively coarse. To the south of the Peloponnese, the shell surface coloration is often paler, with the background more whitish, and there is a tendency to have better separated and more contrasting bands (Supporting Information, Fig. S1G). However, the bands may also be reduced (Fig. 1D; Supporting Information, Fig. S1H). This and the previous form are not clearly separable and there are intermediates, but there is also a corresponding phylogenetic south–north divide (Fig. 4). Populations from Evvia (Fig. 1B; Supporting Information, Fig. S1B) are characterized by pale shells with completely reduced or highly vestigial banding; aperture margins are dark. The upper half of the shell is usually slightly darker than the lower half. The shells often lack spiral sculpture. Columella and aperture margins are relatively slim. This form corresponds fully to H. thiesseana. The analysed individual from Skopelos island (north of Evvia) was also of this type. The examined shells from Crete (Supporting Information, Fig. S1C) were small (around 3 cm), had a dark aperture and pale, but developed, banding and vestigial spiral sculpture. The Turkish populations (Supporting Information, Fig. S1A) are similar to H. thiesseana, and most of the differences may stem from differences in size. The shells are smaller, more compact, and with a smoother surface and finer transverse ribs. Also, unlike H. thiesseana, the Turkish snails often have narrow brown longitudinal bands on the older whorls. Typically, the middle band is the darkest one and is aligned with the suture. The foot is greyish brown with darker, grey or brown back (Fig. 1). The morphology of the genital organs was described by Hesse (1915–20: 183, pl. 654) from specimens from Patra and Kythira Island. Earlier, Schuberth (1892: 51) mentioned a specimen from Corfu as having a short stem of mucous glands and a curved love dart. Neubert (2014: 101) dissected an individual from south-western Anatolia. We examined three individuals from Evvia, 12 from six localities of the ‘ H. aetolica ’ morphotype, two individuals from Kefalonia and one typical H. borealis from Corfu. The genital system (Supporting Information, Fig. S2) does not differ substantially from that of related species (Neubert, 2014) nor are there consistent differences between mitochondrial clades. Short and weakly ramified mucous glands are characteristic. They are usually markedly shorter than the dart sac, but may also be as long as the sac. The love dart is curved towards its tip, with two high and sharp blades in the plane of the curve and two low blunt blades on the sides (the state is unknown for the Cretan-Turkish lineage). The diverticulum branches off in the proximal third to half of the combined pedunculus and bursa stem length. It is usually thick, but its length varies greatly from short (Supporting Information, Fig. S2E, individuals from a locality near Kalavryta, Peloponnese) to almost reaching the length of the bursa stem; sometimes, it is aligned with the bursa stem. The interior of the penis (Supporting Information, Fig. S3) also does not provide characters to differentiate between the H. borealis clades. The atrial stimulator is a knob of varying size. ECOLOGY OF HELIX BOREALIS The habitats where we found H. borealis varied greatly. On Evvia, the snails were attached to high limestone cliffs. Near Sparti, living specimens were also found on bare exposed limestone rocks. In the ruins of ancient Messene, we found it in the grass between the remains of the buildings. In the western Peloponnese, it is often found on Plio-Pleistocene sand and gravel deposits; on this substrate, we found shells on margins of pine forests. On the western coast, we found it in numbers on sand dunes covered with shrubs, a few hundred meters from the shore. Individuals of the ‘ H. aetolica ’ form with closely related haplotypes were found in a grazed phrygana at low elevation (Supporting Information, Fig. S4A), fir growths near Karpenisi at c. 1200 m a.s.l. (Supporting Information, Fig. S4B), and even at a small junk heap under Platanus L. trees in a village. In the north, the species is not limited to limestone. The phenology of the species likely strongly differs between regions. In the south, it is already largely inactive, at least by mid-April, in contrast to the northern part of the range in the mountains in Aetolia. Welter-Schultes (1998a) reports that on Crete the species allegedly emerges after the first October or November rains, only to disappear a few days later. Although there seems to be substantial plasticity, individual lineages may be more restricted in their tolerances. In fact, broadly tolerant is the ‘ H. aetolica ’ form from Peloponnese, Aetolia-Acarnania, Phocis and Evrytania, which appears common in parts of its range. Some other lineages, such as ‘ H. thiesseana ’ and the Cretan lineage, have restricted distributions and probably narrower (realized) niches. At several sites from Lefkada to the Albanian frontier, we have found only old-looking empty shells in April 2016, but it is impossible to say whether this reflects the season or a recent decline. It is also well possible that most of the mortality occurs when the snails are buried in the soil.Published as part of Korábek, Ondřej, Kosová, Tereza, Dolejš, Petr, Petrusek, Adam, Neubert, Eike & Juřičková, Lucie, 2021, Geographic isolation and human-assisted dispersal in land snails: a Mediterranean story of Helix borealis and its relatives (Gastropoda: Stylommatophora: Helicidae), pp. 1310-1335 in Zoological Journal of the Linnean Society 193 (4) on pages 1332-1335, DOI: 10.1093/zoolinnean/zlaa186, http://zenodo.org/record/576147
Amphiscopus moolenbeeki Ruud, Menkhorst & Neubert, 2016, spec. nov.
No full textAmphiscopus moolenbeeki spec. nov. (Fig. 14) Type locality & type material. – Turkey, Vilayet Tokat, Reşadiye, 8 km towards Çekerek, 770 m (40.1523°N 35.6320°E), H.P.M.G. Menkhorst leg., 19.vii.1990. Holotype NMBE 544668, paratypes NMBE 544669/2, HMK/>50, RBA/3, RMNH/3, ZMH/3. Diagnosis. – A large, highly turreted Amphiscopus species without a parietalis or palatalis superior. Description. – Shell dextral, highly turreted, with an open, slit-like umbilicus. The 10.4-13.2 whorls are convex with a moderately deep suture. Teleoconch with widely spaced, fine, oblique striae; there are no spiral striae. Shell solid, not translucent, glossy, uniformly horny yellow coloured, with a prominent whitish band behind the peristome. Peristome not or hardly reflected (with the exception of the columellar part), thickened by a labial callus, the columellar and palatal insertion connected by a clearly visible callus which is quite thickened near the columellar peristome. The edge of the palatal insertion forms a wide arc. The dot-like to tear-like subangularis is connected with the palatal peristome by a thin callus. There is no parietalis, spiralis, palatalis superior, infrapalatalis, basalis or columellaris. The columellar ledge reaches halfway or above the middle of the columellar side of the aperture. Measurements (n = 8). – H = 10.0-14.4 (mean 12.3); LWH = 4.0-4.8 (mean 4.3); MH = 2.6-2.9 (mean 2.7); LWD = 3.0-3.3 (mean 3.1); LWM = 2.9-3.3 (mean 3.1); MD = 1.9-2.2 (mean 2.1); NW = 10.4-13.2 (mean 11.9). Localities. – Vilayet Tokat: type locality (see above); Reşadiye, 7.5 km towards Çekerek (HMK/1). Derivatio nominis. - Named after our friend Rob(ert) Moolenbeek, who made numerous important malacological contributions and who curated the molluscan collection of the formerly Zoologisch Museum Amsterdam for decades (now integrated in the Naturalis Biodiversity Center, Leiden). Remarks. – This taxon has been assigned to Amphiscopus because of the overall shape of the shell, the small aperture and the thickened peristome. So far, only the species sturmii and substurmii has been assigned Amphiscopus; therefore moolenbeeki is now the third known Amphiscopus species. It can be easily recognized from A. sturmii and A. substurmii by the characters mentioned in the diagnosis.Published as part of Ruud, A. Bank, Henk P. M. G Menkhorst & Eike Neubert, 2016, Descriptions of new and little-known land snail taxa from Turkey, and establishment of a new genus (Gastropoda, Pulmonata: Lauriidae, Enidae and Vitrinidae), pp. 5-30 in Basteria 80 (1) on pages 20-22, DOI: 10.5281/zenodo.43974
Euchondrus adwani Eike Neubert & Zuhair Amr 2016, n. sp.
No full textEuchondrus adwani n. sp. (Figure 1) Material: Holotype NMBE 539263; paratype NMBE 539264 /1. Type locality: Syria, surrounding of the monastery of Deir Moussa, 34.0219°N 36.8423°E, 1300 m a.s.l., 11.iii. 2010, leg. Adwan Shehab. *Corresponding author. Email: [email protected] © 2016 Taylor & Francis Measurements (holotype): Height = 11.04 mm; diameter = 4.13 mm; peristome height = 4.03 mm; peristome diameter = 2.97 mm; number of whorls = 8. Diagnosis. Euchondrus adwani n. sp. differs from the widespread E. septemdentatus by its conical shell (broadly oval in E. septemdentatus), its flat suture and teleoconch whorls (suture deeper, whorls much more rounded in E. septemdentatus), the heavy palatal labial callus (weaker in E. septemdentatus), the bar-like subangularis (weaker in E. septemdentatus), and the keeled last whorl (rounded in E. septemdentatus). Description: Shell solid, dextral, cylindrical, upper part cylindro-conical in outline; shell pale brownish to horny yellow coloured; 8 rather flat-sided teleoconch whorls, suture flat with a distinct white sutural thread; teleoconch smooth, glossy, last whorl with fine, straight and irregularly spaced striae; aperture subtriangular, peristome strongly thickened by a labial callus, moderately reflected, with a rich dentition (description clockwise): palatum with a small suturalis followed by a conical palatalis superior and a broad infrapalatalis with the latter two denticles placed on a thick callus; columellar side with a basalis and a straight columellaris; parietum with a strong and long parietalis, bordered by a small spiralis, subangularis large, bar-like, left side of the parietum with another small denticle at the attachment site of the peristome; last teleoconch whorl dorsally compressed forming a distinct blunt ridge (arrows); umbilicus slit-like open, periomphalum large, dish-like. Remarks: This species shows some superficial similarities with E. desertorum Rochanaburananda in Forcart, 1981 (Figure 2), which is endemic to the Negev Desert (Heller, 2009). Both species have a straight conical shell, but E. desertorum is considerably larger than E. adwani n. sp. and its aperture is rounded and not subtriangular. It also differs in the formation of the dentition: in E. desertorum, the infrapalatalis is bifid (simple in E. adwani), the spiralis is large and connected to the parietalis (small and disconnected in E. adwani), and the subangularis is weaker (very strong in E. adwani). The last whorl of E. adwani displays a distinct keel with an enlarged periomphalum, while in E. desertorum the dorsum is rounded, and the periomphalum is much smaller. Etymology: This species is named in honour of Dr. Adwan Shawabi, who was a keen collector of molluscs from Syria, and a personal friend, and who was killed in February 2015 in the Syrian civil war (Amr, 2015).Published as part of Eike Neubert & Zuhair Amr, 2016, On a new species of Euchondrus Boettger, 1883 from Syria (Pulmonata: Enidae), pp. 58-60 in Zoology in the Middle East 62 on pages 58-60, DOI: 10.1080/09397140.2015.1132564, http://zenodo.org/record/88703
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