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    Kanakaster plinthinos Mah 2017

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    Kanakaster plinthinos Mah 2017 Figure 17 C, D Mah 2017: 34 Comments. These specimen are nearly identical to the New Caledonia individuals described by Mah (2017) with the notable difference that the small bare patches present on the superomarginal plates in the type series are absent (Fig. 17C) and in having more than ten marginal plates per interradius rather than six to eight (Fig. 17C). The body shape and joined distal superomarginals are identical. These are the first recorded occurrences of this species from beyond the type locality. Occurrence. New Caledonia, Madagascar, Kenya, 140– 350 m. Material Examined. IE-2007-4004 Madagascar. 2608’S 4539’E, 333 m, Coll. Atimo Vatae, N / O Nosy Be 11, st. DW 3553, 5 May 2010. 1 wet spec. R=1.3 r=0.6. USNM E51268 NE of Mombasa, Kenya. 242’S, 4053 E, 140 m. Coll. R / V Anton Bruun 6 Nov. 1964. 1 wet spec. R=1.9, r=0.8.Published as part of Mah, Christopher L., 2018, New genera, species and occurrence records of Goniasteridae (Asteroidea; Echinodermata) from the Indian Ocean, pp. 1-116 in Zootaxa 4539 (1) on page 48, DOI: 10.11646/zootaxa.4539.1.1, http://zenodo.org/record/261591

    Bathyceramaster Mah 2016

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    <i>Bathyceramaster</i> Mah 2016a <p>Mah 2016a: 105</p> <p> <b>Diagnosis.</b> Abactinal plates tabulate, low to moderate in height with fasciolar grooves, variably shallow to well-developed, plates lacking stellate bases. Abactinal, marginal, actinal surfaces covered by densely arranged polygonal to round granules. Body stellate, many species with well-developed arms (R/ r=1.8–4.0).</p> <p> <b>Comments.</b> This genus was created to accommodate <i>Mediaster elegans</i> Ludwig 1905, which lacked the stellate, radiating ossicles that are present in all species belonging to the genus <i>Mediaster</i> (Fisher 1911; Mah 2016a). The tabular plates and well developed fasciolar grooves were also not as well-developed in <i>Bathyceramaster</i> as they are in <i>Mediaster</i>. Known <i>Bathyceramaster</i> spp. have only been reported from relatively deep-water settings, below 600 m, primarily between 1000–4000 m.</p> <p> Based on the occurrence of known species, <i>Bathyceramaster</i> occurs widely throughout the Pacific in a manner similar to other deep-sea Goniasteridae, such as <i>Sibogaster</i> (Mah 2016a).</p>Published as part of <i>Mah, Christopher L., 2022, New Genera, Species and Occurrences of Deep-Sea Asteroidea (Valvatacea, Forcipulatacea, Echinodermata) collected from the North Pacific Ocean by the CAPSTONE Expedition, pp. 1-75 in Zootaxa 5164 (1)</i> on page 31, DOI: 10.11646/zootaxa.5164.1.1, <a href="http://zenodo.org/record/6821026">http://zenodo.org/record/6821026</a&gt

    Cnemdiaster gilesi Mah 2007, COMB. NOV.

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    <i>CNEMDIASTER GILESI</i> (ALCOCK, 1893) COMB. NOV. <p> <i>Zoroaster gilesi</i> Alcock, 1893: 108; H.L. Clark, 1920: 101; Clark & Mah, 2001: 234; Clark & Mah, 2001: 233; Sastry, 2005: 47 143 174.</p> <p> <i>Occurrence</i>: Andaman Sea, 896–915 m.</p>Published as part of <i>Mah, Christopher, 2007, Phylogeny of the Zoroasteridae (Zorocallina; Forcipulatida): evolutionary events in deep-sea Asteroidea displaying Palaeozoic features, pp. 177-210 in Zoological Journal of the Linnean Society 150 (1)</i> on page 190, DOI: 10.1111/j.1096-3642.2007.00291.x, <a href="http://zenodo.org/record/5430876">http://zenodo.org/record/5430876</a&gt

    Circeaster pullus Mah 2006

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    Circeaster pullus Mah 2006 Figure 4 B, C Mah 2006: 940. Feeding comments. Circeaster pullus is a widely occurring Pacific species, described from the Hawaiian Islands by Mah (2006), who also documented this species feeding on unidentified octocoral/gorgonian skeletons (Mah 2006: 948 - 499). Additional images of C. pullus include (P 5-690 -d 2-11522, P 5-690 -d 5-3819; P 5-690 -d 5-3835) which show the oral side of a goniasterid crawling along the branches of an apparent isidid octocoral (Figs 4 B, C). One image (P 5-280 -hi 8 -05129) shows what appears to be C. pullus with its disk swollen and extended above the plane of the arms and marginal plate series (Fig. 4 B) feeding on an isidid octocoral. The animal appears to be positioned above rocks with octocorals beneath it, suggesting predation. The swollen disk in C. pullu s is presumably filled with water and resembles the same type of swollen disk observed in other goniasterids such as Ceramaster granularis (Gale et al. 2013). Video observations. Keaholu Pt., Big Island: P 5-280 -hi 8 -05129. 20.6501, - 156.04301, 550 m. Twin Banks, NWHI: P 5-690 -d 5-11522, P 5-690 -d 5 -03835, P 5-690 -d 5 -03819. 23.049283, - 163.1613, 1430–1433 m.Published as part of Mah, Christopher L., 2015, New species, corallivory, in situ video observations and overview of the Goniasteridae (Valvatida, Asteroidea) in the Hawaiian Region, pp. 211-228 in Zootaxa 3926 (2) on page 222, DOI: 10.11646/zootaxa.3926.2.3, http://zenodo.org/record/23578

    Kanakaster larae Mah 2017

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    Kanakaster larae Mah 2017 Figure 17A, B Mah 2017: 32 Diagnosis. Weakly stellate/pentagonal body form (Fig. 17A). Bare patches on the superomarginal plates absent. Body surface completely covered by soft granular tegument with no bare spots on marginal plates. When underlying tegument has been removed, abactinal and marginal surface with glassy nodules (also called embossed crystalline bodies). Fifteen to sixteen marginal plates present per interradius (arm tip to arm tip) (but see comments below). Comments. Although smaller than the holotype of Kanakaster larae, this specimen shows the distinctive tegument, the wide superomarginal and inferomarginal plates and the numerous crystalline tubercles present in the abactinal, marginal and actinal plates below the tegument. It also possesses three furrow spines associated with each adambulacral plate. The abactinal plates are round and polygonal as they are in the holotype. The biggest differences between the two specimens are size-related. The specimen herein has only six marginal plates per interradius (arm tip to arm tip) (Fig. 17A, B) whereas the holotype had approximately 16 per interradius. But holotype was also substantially larger (R=3.2) than the specimen herein (R=1.5) suggesting that this could be a growth-related character. This represents the second recorded occurrence of this species. Occurrence. Port Edward, South Africa, Western Indian Ocean, NE of the Glorioso Islands, 120– 301 m. This individual was from a sample with a sponge-dominated bottom. Material Examined. IE-2013-17194, Western Indian Ocean, NE of the Glorioso Islands, 11 ° 29 'S, 47 ° 29' E to 11 ° 30 'S, 47 ° 29' E, 293–301 m. Coll. DW 4809, Jan 25, 2017. 1 wet spec. R=1.5 r=1.0.Published as part of Mah, Christopher L., 2018, New genera, species and occurrence records of Goniasteridae (Asteroidea; Echinodermata) from the Indian Ocean, pp. 1-116 in Zootaxa 4539 (1) on pages 47-48, DOI: 10.11646/zootaxa.4539.1.1, http://zenodo.org/record/261591

    Sthenaster emmae Mah 2010

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    Sthenaster emmae Mah et al. 2010 Figure 17 A–D In situ images of this species were identified based on the distinctive arrangement of abactinal plates as well as the wide superomarginal interradial plates. This is contrasted with those in Gilbertaster which are more elongate. Color in life of this species is white on the disk surface with darker orange plates. Arms are a more solid colored light orange. A second morphotype, of what is interpreted as a smaller individual was a uniform pink-brown in color. New observations of this species in conjunction with the extensive survey observations of Okeanos Explorer throughout the Gulf of Mexico and adjacent regions show occurrence to be limited to the southeastern coastal region of North America, specifically, the Savannah Banks and Central Plateau Scarp region. Feeding & Other Observations Imagery collected by the Okeanos Explorer are the first clear observations of this species in situ and apparently feeding on prey. Initial determination of the predatory status of Sthenaster emmae was based primarily on gut contents of the holotype, which included spicules from Eunicella modesta (Verrill 1883) as well as unclear video. Video observations captured one small individual on a denuded stalk projecting from dead Lophelia. Other imagery captured Sthenaster with a swollen disk possibly hunched over an unidentified prey item. Occurrence: Savannah Banks and off the coast of Jacksonville, FL, Central Plateau Scarp and Richardson’s Jellyfish, 252– 874 m. Images Examined Central Plateau Scarp, 30.924592, -78.088036, 874 m, EX 1903L2_IMG_20190629 T 165108Z_ ROVHD.jpg (small individual) Central Plateau Scarp, 30.925962, -78.089989, 865 m, EX 1903L2_IMG_20190629 T 184639Z_ ROVHD.jpg (swollen) Central Plateau scarp, 30.924906, -78.088884, 870 m EX 1903L2_IMG_20190629 T 173638Z_ ROVHD.jpg (swollen) Richardson’s Jellyfish, 30.924993, -78.089565, 865 m, EX 1903L2_IMG_20190701 T 180214 Z_ ROVHD.jpgPublished as part of Mah, Christopher L., 2020, New species, occurrence records and observations of predation by deep-sea Asteroidea (Echinodermata) from the North Atlantic by NOAA ship Okeanos Explorer, pp. 201-260 in Zootaxa 4766 (2) on pages 237-238, DOI: 10.11646/zootaxa.4766.2.1, http://zenodo.org/record/376401

    Neoferdina antigorum Mah 2017

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    <i>Neoferdina antigorum</i> Mah, 2017 <p>FIGURE 11A–E</p> <p>Mah 2018: 44</p> Diagnosis <p>This species is characterized by the presence of very few to absent strongly convex, bald abactinal plates but with a distinct round, bald patch on each superomarginal plate (Figs.11A–C), which forms a regular, homogeneous series along the arm (Fig. 11C). A continuous granular tegument covers most of the body surface, including abactinal, marginal and actinal plates (Figs 11B, D–E). Abactinal plates are mostly homogeneous (Fig. 11A–B). This species has a minority of convex abactinal plates with bare patches and spinelets on distalmost inferomarginal plates (Fig. 11A–B), bald patches absent from the proximalmost inferomarginal plates (Fig. 11A).</p> Comments <p> <i>Neoferdina antigorum</i> was described from mesophotic depths in the Philippines (Mah 2017) and further specimens from mesophotic settings, as reported herein, suggest that it may be widespread across the tropical Pacific. This species has not been recorded above the mosphotic zone.</p> Occurrence <p>Okinawa, Japan, 46– 53 m.</p> <p>Outside Japan. Philippines, 80– 200 m.</p> Material Examined <p>USNM E45964 3km WSW of Yuhi Misaki, Okinawa Island, Ryukyu Islands, 26.8467 127.253, 52–53 m. Coll. RF Bolland, 14 Nov 1987, 1 dry spec. R=1.7 r=0.6.</p> <p>USNM 1579916 1 km WNW of Onna Village, Horseshoe Cliffs, Okinawa Island, Ryukyu Islands, 26.5 127.848, 46–52 m. Coll. R.F. Bolland, 8 Feb 1990. 1 wet spec. R=1.4 r=0.4.</p>Published as part of <i>Mah, Christopher L., Kogure, Yoichi, Fujita, Toshihiko & Higashiji, Takuo, 2024, New Taxa and Occurrences of Mesophotic and Deep-sea Goniasteridae (Valvatida, Asteroidea) from Okinawa and adjacent regions, pp. 1-41 in Zootaxa 5403 (1)</i> on page 29, DOI: 10.11646/zootaxa.5403.1.1, <a href="http://zenodo.org/record/10561570">http://zenodo.org/record/10561570</a&gt

    Ryukyuaster onnae Mah 2007

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    <i>Ryukyuaster onnae</i> Mah 2007 <p>FIGURE 13C–D</p> <p>Mah 2007: 320</p> Diagnosis <p>Body form stellate.Approximately 16 marginals per interradius. Lateral edge of marginal plates with granular cover. Tubercular granules, granular spinelets on abactinal, marginal plate surface (Fig. 13C). Intermarginal pits present between central two to three interradial superomarginal–inferomarginal plate pairs. Pits bare, flanked by clavate spines, 3–4. Actinal intermediate plates covered by polygonal granules at R = 1.5 cm (Fig. 13D). Furrow spines three to five. Terminal plates enlarged.</p> Comments <p>Nearly all of the material for this species from Mah (2007) were collected from mesophoitc depths (40–60 m) with only two individuals from shallow water (2–4 m).</p> <p>The new specimen from the Sulu Archipelago suggests that this species is much more widespread than was previously realized. Its mesophotic occurrence (51 m) and small size might explain why it has been recognized only recently.</p> Occurrence <p>Okinawa, Ryukyu Islands, Japan. 40–51.2 m.</p> <p> Outside Japan. <b>Sulu Archipelago (extension).</b> 3– 67 m.</p> Material Examined <p>USNM E46165 Seragaki Beach, 1.3 km ENE of Maeki-zaki, Okinawa, Ryuku Islands, Japan. 45.75–48.8 m Coll. R.F. Bolland 16 Aug 1985. (1 dry spec. R=0.9, r=0.5)</p> <p>USNM E53611 Horseshoe Cliffs, 1km WNW of Onna Village, Okinawa, Ryuku Islands, Japan. 26º30’N, 127º50’54E, 40–46 m (130–150 ft), Coll. R.F. Bolland 16 March 1985, 1 dry spec. R=1.1, r= 0.6 cm.</p> <p>USNM E53706 Horseshoe Cliffs, 1km WNW of Onna Village, Okinawa, Ryuku Islands, Japan. 26º30’N, 127º50’ 54E, 47 m (155 ft). Coll. R. F. Bolland 20 Jan 1985, 1 dry spec. R=1.3, r=1.0.</p> <p> WAM 1629.74 (1) SE of Capui Island, Sulu Archipelago, sand, <i>Lithothamnion</i>, 51.2 m. Coll. B.R. Wilson, Feb 19, 1964. 1 dry spec. R=1.7 r=1.0 cm.</p>Published as part of <i>Mah, Christopher L., Kogure, Yoichi, Fujita, Toshihiko & Higashiji, Takuo, 2024, New Taxa and Occurrences of Mesophotic and Deep-sea Goniasteridae (Valvatida, Asteroidea) from Okinawa and adjacent regions, pp. 1-41 in Zootaxa 5403 (1)</i> on pages 34-35, DOI: 10.11646/zootaxa.5403.1.1, <a href="http://zenodo.org/record/10561570">http://zenodo.org/record/10561570</a&gt

    Sagenaster Mah 2007, GEN. NOV.

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    <i>SAGENASTER</i> MAH GEN. NOV. <p> <i>Etymology</i>: Latin (<i>sagena</i>) for fish-net, referring to the reticulate or net-like skeleton in this genus.</p> <p> <i>Type species</i>: <i>Zoroaster evermanni</i> Fisher, 1905.</p> <p> <i>Diagnosis</i>: Disc and arm skeleton reticulate. Plates cruciform, forming open papular fields. Primary spines present on all plates. Internal buttress absent.</p> <p> <i>SAGENASTER EVERMANNI</i> (FISHER, 1905) (W/SYNONYM <i>EVERMANNI MORDAX</i> FISHER, 1919)</p> <p>(FIGS 2E, F)</p> <p> <i>Zoroaster</i> (<i>Myxoderma) evermanni</i> Fisher, 1905: 317; not H.L. Clark, 1913: 198 (= <i>Myxoderma sacculatum</i>).</p> <p> <i>Zoroaster evermanni</i> Fisher, 1919a: 390; H.L. Clark, 1920: 100; 1923: 152; Fisher, 1928: 40, pl. 13, fig. 3, pl. 14, figs 1.1a, 1b, pl. 18, fig. 4; pl. 19, fig. 1; pl. 20, figs 3, 4; Alton, 1966: 1709; Carey, 1972: 41; Lambert, 1978a: 23; 1978b: 64; Maluf, 1988: 44, 125; Nybakken <i>et al</i>., 1998: 1777; Clark & Mah, 2001: 233.</p> <p> <i>Zoroaster evermanni mordax</i> Fisher, 1919a: 391, 1928: 34, 43, pl. 13, fig. 4; pl. 14, fig. 2; pl. 19, fig. 2.</p> <p> <i>Occurrence</i>: Aleutian Islands (Alaska), Queen Charlotte Island (Canada), Oregon, Washington to S. California, Mexico, 100–2710 m.</p> <p> <i>Material</i>: Alaska: CASIZ 115898, Aleutian Islands, Unalaska Island, 54.00°21.50″N, 167.00°47.50″W, 905–910 m, Coll. 14.vi.1979 (wet spec. <i>R</i> = 17.2, <i>r</i> = 1.6). Oregon: NMNH E 10356, SW of mouth of Columbia River, 45°51.5′N, 124°42′W. 823 m, Coll. 6.iii.1962. (1 dry spec. <i>R</i> = 12.6, <i>r</i> = 0.9 cm) NMNH E 10359 SW of mouth of Columbia River, 45°52.3′N, 124°52′W. 823 m, Coll. 13.v.1963. (1 dry spec. <i>R</i> = 9.1, <i>r</i> = 0.9 cm) NMNH E16023, SW of mouth of Columbia River, 45°52′N, 124°54′W. 823 m, Coll. 27.v.1962 (1 dry spec. <i>R</i> = 9.2, <i>r</i> = 0.9 cm); NMNH E 10360, SW of mouth of Columbia River, 45°40.5′N, 124°55′W, 1372 m, Coll. 2.viii.1963. (1 dry spec. <i>R</i> = 3.2, <i>r</i> = 0.35 cm) NMNH E 10357, SW of mouth of Columbia River, 45°52′N, 124°54′W. 732 m, Coll. 23.i.1963. (1 dry spec. <i>R</i> = 14.2, <i>r</i> = 0.8 cm) NMNH E 16019, SW of mouth of Columbia River, 45°56′N, 124°51′W. 682 m, Coll. 12.iii.1962. [1 dry spec. <i>R</i> = ∼8.2, <i>r</i> = 0.8 cm (arm tips broken)] NMNH E 10358, SW of mouth of Columbia River, 45°57.3′N, 124°48.7′W. 594 m, Coll. 10.v.1962. [1 dry spec. <i>R</i> = ∼16, <i>r</i> = 0.9 cm (arm tips broken)] NMNH E 16007, SW of mouth of Columbia River, 46°14′N, 124°44′W. 979 m, Coll. 13.iii.1962. [2 dry specs. <i>R</i> = ∼12.5, <i>r</i> = 1.0 cm, <i>R</i> = 7.2, <i>r</i> = 0.8 cm (arm tips broken)] CASIZ 122302, Off Oregon coast, 44°20′N, 125°5′W, 823–914.0 m, Coll. 10.xii.1961 (1 wet spec. <i>R</i> = 11.8, <i>r</i> = 0.7). CASIZ 121506 Cascadia Plain, 45°55.50′N, 125°38.8′W, 2030.0 m, Coll. 20.iii.1970. (5 wet specs. <i>R</i> = 2.0, <i>r</i> = 0.3, <i>R</i> = 1.1, <i>r</i> = 0.2, <i>R</i> = 0.8, <i>r</i> = 0.2; <i>R</i> = 1.1, <i>r</i> = 0.2, <i>R</i> = 0.4, <i>r</i> = 0.1) California: CASIZ 113317 Humboldt County on mud, 704–841 m, Coll. vi.1977. (3 dry spec. <i>R</i> = 8.8, <i>r</i> = 1.2, <i>R</i> = 12.2, <i>r</i> = 1.1; <i>R</i> = 9.7, <i>r</i> = 0.8) CASIZ 113318 Eureka, Humboldt County on soft green-black mud, 100–1005 m, Coll. i.1977. (2 dry specs. <i>R</i> = 10.6, <i>r</i> = 0.8, <i>R</i> = 11.2, <i>r</i> = 0.8) CASIZ 11319, Eureka, Humboldt County on soft green-black mud, 823 m [2 dry specs. <i>R</i> = 10.5, <i>r</i> = 1.1, <i>R</i> = ∼15, <i>r</i> = 1.1 (arm tips broken)] CASIZ 115950 Off Point Loma Lighthouse, San Diego on green mud, fine sand, 294–933 m, Coll. 5.iii.1904. [1 wet spec. <i>R</i> = 13.9, <i>r</i> = 1.1 (arm tips broken)] CASIZ 115910 Gulf of Farallones, 913–1000 m, Coll. 16.xii.1985. (2 wet specs., <i>R</i> = 3.1, <i>r</i> = 0.4, <i>R</i> = 2.6, <i>r</i> = 0.6) CASIZ 115925 Off Bodega head, Sonoma County, 549–567.0 m Coll. 16.iii.1965. [1 wet spec. <i>R</i> = ∼14.8, <i>r</i> = 1.0 (arms broken)]</p>Published as part of <i>Mah, Christopher, 2007, Phylogeny of the Zoroasteridae (Zorocallina; Forcipulatida): evolutionary events in deep-sea Asteroidea displaying Palaeozoic features, pp. 177-210 in Zoological Journal of the Linnean Society 150 (1)</i> on page 194, DOI: 10.1111/j.1096-3642.2007.00291.x, <a href="http://zenodo.org/record/5430876">http://zenodo.org/record/5430876</a&gt
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