1,723,706 research outputs found

    Glaphyrocyclosorus X. Wan & Li Bing Zhang

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    × Glaphyrocyclosorus X.Wan & Li Bing Zhang, nothogen. nov. ≡ “ × Glaphypseudosorus erubesquirolicus Hong M.Liu, Schuettp. & H.Schneid. ” (2020: 1000), nom. inval. A hybrid genus between Glaphyropteridopsis Ching (1963: 320) and Pseudocyclosorus Ching (1963: 322). Etymology:—A combination of “ glaphyro -” (from Glaphyropteridopsis) and “- cyclosorus ” (from Pseudocyclosorus). This nothogenus contains one nothospecies.Published as part of Wan, Xia & Zhang, Li-Bing, 2022, Three new fern names of hybrid genera and four new combinations in Athyriaceae, Osmundaceae, and Thelypteridaceae, pp. 104-108 in Phytotaxa 575 (1) on pages 105-106, DOI: 10.11646/phytotaxa.575.1.8, http://zenodo.org/record/740335

    Osmunimunda X. Wan & Li Bing Zhang

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    × Osmunimunda X.Wan & Li Bing Zhang, nothogen. nov. ≡ “ × Osmuntonia Hong M.Liu, Schuettp. & H.Schneid. ” (2020: 999), nom. inval. A hybrid genus between Osmunda Linnaeus (1753: 1063) and Claytosmunda (Y.Yatabe, N.Murakami & K.Iwatsuki 2005: 127–128) Metzgar & Rouhan (2016: 594). Etymology:—A combination of “ osmun -” (from Osmunda) and “- munda ” (from Claytosmunda). This nothogenus contains two nothospecies.Published as part of Wan, Xia & Zhang, Li-Bing, 2022, Three new fern names of hybrid genera and four new combinations in Athyriaceae, Osmundaceae, and Thelypteridaceae, pp. 104-108 in Phytotaxa 575 (1) on page 105, DOI: 10.11646/phytotaxa.575.1.8, http://zenodo.org/record/740335

    Apseudathyrium X. Wan & Li Bing Zhang

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    × Apseudathyrium X.Wan & Li Bing Zhang, nothogen. nov. “ × Schnellerathyrium Hong M.Liu, Schuettp. & H.Schneid. ” (2020: 1000), nom. inval. A hybrid genus between Athyrium Roth (1800: 31, 58–59) and Pseudathyrium Newman (1851: 370–371). Etymology:—A combination of Greek “ a ”, without (from Athyrium), and “ pseudathyrium ”. This nothogenus contains one nothospecies.Published as part of Wan, Xia & Zhang, Li-Bing, 2022, Three new fern names of hybrid genera and four new combinations in Athyriaceae, Osmundaceae, and Thelypteridaceae, pp. 104-108 in Phytotaxa 575 (1) on pages 104-105, DOI: 10.11646/phytotaxa.575.1.8, http://zenodo.org/record/740335

    Polystichum uniseriale Li Bing Zhang 2012, comb. nov.

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    Polystichum uniseriale (Ching ex K.H.Shing) Li Bing Zhang, comb. nov. Basionym: Cyrtomium uniseriale Ching ex Shing (1965: 15). Type:— CHINA. Chongqing: 1957, T. Z. Liu et al. 10070 (holotype PE-00044555!, isotype PE-00044557!). This species occurs in Chongqing and southern and central Sichuan (Pingshan, Ya'an), China.Published as part of Zhang, Li-Bing, 2012, Taxonomic and nomenclatural notes on the fern genus Polystichum (Dryopteridaceae) in China, pp. 57-60 in Phytotaxa 60 (1) on page 59, DOI: 10.11646/phytotaxa.60.1.8, http://zenodo.org/record/506167

    Polystichum huashanicola Li Bing Zhang 2012, stat. nov.

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    Polystichum huashanicola (W.M.Chu & Z.R.He) Li Bing Zhang, stat. nov. Basionym:— Polystichum daguanense Ching ex L.L.Xiang var. huashanicola Chu & He (2000: 257, as " huashanicolum "). Type:— CHINA. Yunnan: Zhengxiong Xian, Huashan, in fissures of limestone under shrub forest, 1950 m, 31 August 1991, W. M . Chu & G. F. Zhang 24241 (holotype PYU!). This species is treated as a variety of Polystichum daguanense Ching ex L.L.Xiang by Zhu & He (in Kung et al. 2001), however, the two taxa are different morphologically (Zhang & Barrington, in press). They do not seem to be closely related. Polystichum huashanicola occurs in northeastern Yunnan, China. Polystichum kangdingense H.S.Kung & Li Bing Zhang, sp. nov. Type:— CHINA. Sichuan: Kangding County, 2400 m, 28 August 1981, Xian-Xu Kong (Hsian-Shiu Kung) 6189 (holotype CDBI!, isotype PE!). Based on the full Latin diagnosis in Kung & Zhang (1998: 254). This species was not validly published there since no holotype is provided. By providing the type information we hereby validate this name. Illustration: Kung & Zhang (1998: 253, pl. 3: f. 1–4.) Diagnosis: Kung & Zhang (1998: 254) ' Species affinis Polysticho submit Christ, sed laminis majoribus, bipinnatis, paleis stipitum magis latioribus, ovatis differt '.Published as part of Zhang, Li-Bing, 2012, Taxonomic and nomenclatural notes on the fern genus Polystichum (Dryopteridaceae) in China, pp. 57-60 in Phytotaxa 60 (1) on page 58, DOI: 10.11646/phytotaxa.60.1.8, http://zenodo.org/record/506167

    Dryopteris wantsingshanica Li Bing Zhang 2012, comb. nov.

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    22. Dryopteris wantsingshanica (Ching & K.H.Shing) Li Bing Zhang, comb. nov. Basionym:— Ctenitis wantsingshanica Ching & K.H.Shing in Ching & Wang (1981: 123). Type: CHINA. Guizhou: Yinjiang, Fanjing Shan (Wantsing Shan), in forest, 2000 m, C. B. Jian et al. 31845 (holotype, PE-00044725!). Homotypic synonym:— Dryopsis wantsingshanica (Ching & K.H.Shing) Holttum & Edwards (1986: 194). This species is synonymized by Dong & Christenhusz (in press) with Dryopteris nidus. This species is insufficiently known, but it seems to have narrower scales on rhizomes and petioles. This species is only found in the type locality.Published as part of Zhang, Li-Bing, 2012, Reducing the fern genus Dryopsis to Dryopteris and the systematics and nomenclature of Dryopteris subgenus Erythrovariae section Dryopsis (Dryopteridaceae), pp. 17-27 in Phytotaxa 71 (1) on page 26, DOI: 10.11646/phytotaxa.71.1.4, http://zenodo.org/record/506633

    Dryopteris dulongensis Li Bing Zhang 2012, comb. nov.

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    5. Dryopteris dulongensis (S.K.Wu & X.Cheng) Li Bing Zhang, comb. nov. Basionym:— Ctenitis dulongensis Wu & Cheng (1985: 402, pl. 3). Type: CHINA. Yunnan: Gongshan, Dulong Jiang (Taron), in silvis semperviritibus humidis, alt. 1700 m, Qinghai-Xizang Complex Exp. 82-9568 (holotype, KUN). Homotypic synonym:— Dryopsis dulongensis (S.K.Wu & X.Cheng) S.Y. Dong (2001: 187). This species is only known from the type locality. This species has been hypothesized as a hybrid between D. apiciflora and D. heterolaena (Dong & Christenhusz, in press).Published as part of Zhang, Li-Bing, 2012, Reducing the fern genus Dryopsis to Dryopteris and the systematics and nomenclature of Dryopteris subgenus Erythrovariae section Dryopsis (Dryopteridaceae), pp. 17-27 in Phytotaxa 71 (1) on page 21, DOI: 10.11646/phytotaxa.71.1.4, http://zenodo.org/record/506633

    Whittieria engelmannii Li Bing Zhang & Liang Zhang ex Li Bing Zhang, Liang Zhang & X. Wan 2022, comb. nov.

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    Whittieria engelmannii (Prantl 1883: 351) Li Bing Zhang & Liang Zhang ex Li Bing Zhang, Liang Zhang & X.Wan, comb. nov. Basionym: Ophioglossum engelmannii Prantl (1883: 351) ≡ “ Whittieria engelmannii (Prantl) Li Bing Zhang & Liang Zhang ” (2022: 24), nom. inval. Type:— U.S.A. Texas: Comanche Spring, New Braunfels, May 1849, F.J. Lindheimer 1281 (lectotype MO201251!, here designated, isolectotypes FI003993!, K001057659!, MEXU00085077!, MEXU00000139!, TEX00348044!). Remaining syntypes: U.S.A. Texas: May 1849, F.J. Lindheimer 95 (MO251338!). U.S.A. Texas: Comanche Spring, Bexar County, May 1849, F.J. Lindheimer 53 (GH00021725!, FI003992!). Notes:— In the protologue (Prantl 1883), there were no specific gatherings cited but only “Nordamerika” (North America) was given. According to Stafleu & Cowan (1983: 379), K.A.E. Prantl’s herbarium and types are mainly at “HBG (esp. Pteridophyta)”. We searched the database of HBG (www.herbariumhamburgense.de), and only found three gatherings of Ophioglossum from Namibia and South Africa. In Tropicos (tropicos.org), four gathering are indicated as “T” (type) or “ST” (syntypes). We found three of the four gatherings in various herbaria (see above) in JSTOR (plants.jstor.org) but could not find Engelmann s.n collected before 1883. We here designated the one of the duplicates of F.J. Lindheimer 1281 at MO as the lectotype. This species has three unique features: growing in basic soils, double venation (large areoles of the sterile blade subdivided into smaller areoles; Wagner & Wagner 1994), and 71 days of spore germination time (Whittier 1981, Zhang & Zhang 2022). Distribution:— United States, Mexico, and Central America.Published as part of Wan, Xia, Zhang, Liang & Zhang, Li-Bing, 2022, Validation and lectotypification of the fern combination Whittieria engelmannii (Ophioglossaceae), pp. 205-206 in Phytotaxa 567 (2) on pages 205-206, DOI: 10.11646/phytotaxa.567.2.10, http://zenodo.org/record/714165

    Polystichum gonggashanense H. S. Kung & Li Bing Zhang ex H. He & Li Bing Zhang 2022, sp. nov.

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    Polystichum gonggashanense H.S.Kung & Li Bing Zhang ex H.He & Li Bing Zhang, sp. nov. (Figs. 1 & 2). Type:— CHINA. Sichuan: Luding County, Mt. Gongga (Minyar Gonkar), Hailuogou, #3 rd Camp to Chengmendong, elev. ca. 2950 m, 20 July 2010, Hai He 1190 (HG037) (holotype CDBI; isotypes CDBI, CTC, MO). Diagnosis:—The new species is most similar to Polystichum sinense (Christ in Baroni & Christ 1901: 289) Christ (1905: 30), but the former has an oblong-lanceolate lamina with the lower portion only slightly contracted toward the lamina base, petiole scales that are blackish brown, and rachis scales that are narrowly lanceolate. In contrast, the latter has an elliptic-lanceolate lamina with the lower portion substantially contracted toward the lamina base, petiole scales that are brown, and broad-lanceolate rachis scales. Description:—Plants 73–87 cm tall. Rhizomes erect; fronds cespitose. Petioles 24–33 cm long, 3–5 mm in diam., stramineous; large scales ovate or ovate-lanceolate often 20 mm long, 8.5 mm wide, acute at apex, ciliate or subentire on margins, the proximal ones uniformly blackish brown or brown, thick chartaceous, distally transitioning gradually to bicolorous with paler-colored margins, then to concolorous light yellow-brown and membranous; small scales linear to lanceolate. Laminae oblong-lanceolate, 49–57 cm long, 16–21 cm wide, elongated-acuminate or caudate at apex, lower portion sterile, bipinnate. Rachises ca. 2 mm in diam.; rachis scales narrow-lanceolate, brownish, long-acuminate at apex, subentire on margins, spreading or reflexed. Pinnae 22–31 pairs, spreading, alternate, short-petiolate, middle pinnae 1.5–2.5 cm apart, 4.5–10 cm long, 1.9–2.3 cm wide, lanceolate, apex acuminate or caudate, base inequilateral, 1-pinnate; basal pinnae slightly shorter, 4.5–7.0 cm long, with 7–11 pairs of free pinnules; pinna rachis with linearlanceolate small scales (these denser abaxially), ca. 5 mm long. Pinnules 13–16 pairs, sessile, spreading, herbaceous, deltoid-ovate, 12–13 mm long, 4.1–5.5 mm wide, auriculae deltate, base cuneate and slightly adnate (only the distal third of the pinna), apex acute, margins aristate-dentate, with 1.5–3.5-mm-long linear microscales on both surfaces, basal acroscopic pair the longest and lobed; veins 5–7 pairs per pinnule, dichotomous, slightly visible. Sori light brown, one row on each side of midrib, close to midrib; indusia peltate, rotund, membranous, margins subentire. Etymology:—Derived from gonggashan in Chinese pinyin, meaning Mt. Gongga (Minyar Gonkar). Vernacular name:— ūḝ山φƦ (gong ga shan er jue). Distribution:—It was found on the eastern, southeastern, and western slopes of Mt. Gongga in Jiulong, Luding, and Kangding counties, Sichuan Province, Southwest China, with elevational range of 2500–3200 m. Conservation assessment:—Based on thorough explorations by Kai-Yong Lang et al. in 1982, by L-BZ in 1989, 1990, and 1993, and by HH in 2010, only four very small populations each with a few individuals were found.All these populations are in high elevations of the same mountain. The expansion of the touristic facilities and global warming threaten the survival of Polystichum gonggashanense and the population in the first locality already disappeared. We suggest that the status of Polystichum gonggashanense be listed as CR (Critically Endangered) following IUCN criteria (IUCN 2019). Additional specimens examined (paratypes):— CHINA. Sichuan: Luding County, Mt. Gongga (Minyar Gonkar), Hailuogou, elev. ca. 2980 m, 29 October 1989, Li-Bing Zhang 169 (KUN, PYU); ibid., 1990, Li-Bing Zhang 911 (CDBI); Jiulong County, Hongba, elev. ca. 2500 m, on forest margin, October 1990, Li-Bing Zhang 2786 (CDBI, PYU); Kangding County, Liuba, elev. ca. 3200 m, July 1982, Kai-Yong Lang et al. 752 (KUN, PE); same town, 1990, Li-Bing Zhang s.n. (CDBI, PYU). Taxonomic notes:—We examined all specimens available at CTC and it appears that all sporangia were dehisced. There were fewer than 32 spores per sporangium and most sporangia had only 10–20 spores upon examination. It is possible that the new species is apomictic. In middle to high elevations of the Sino-Himalayan region species of Polystichum are common, most of which belong to P. sect. Hypopeltis (Michaux 1803: 266) T. Moore (1857: 84) taxonomically (Zhang & Barrington 2013). The taxonomy of this section is difficult partially because the species are over-described (Zhang & Barrington 2013). Between 1983 and 1993, seven “species” were described for P. sinense alone (Ching in Wu 1983, Ching & S.K.Wu in Wu 1983, Yang 1992) including P. alatawshanicum Chang Y. Yang (1992: 302), P. atuntzeense Ching in Wu (1983: 231), P. austrotibeticum Ching & S.K.Wu in Wu (1983: 233), P. decorum Ching & S.K.Wu in Wu (1983: 229), P. ellipticum Ching & S.K.Wu in Wu (1983: 233), P. lhasaense Ching in Wu (1983: 230), and P. parasinense Chang Y. Yang (1992: 302). Relevant molecular data for Polystichum so far have not yet been based on extensive taxon sampling of P. sect. Hypopeltis (Liu et al. 2007, Le Péchon et al. 2016a, 2016b). Polystichum gonggashanense has not been described so far partially because it is a rare species endemic to a small area in Mt. Gongga. Notably, Polystichum decorum, thought as conspecific with P. gonggashanense by the reviewers of Acta Botanica Yunnanica as stated in the introduction, is a synonym of P. sinense. The distinction between P. gonggashanense and P. sinense is not neglectable in the color of petiole scales, the lamina shape, and the shape of rachis scales (see above; Table 1). In addition to Polystichum sinense, P. gonggashanense is also similar to P. haleakalense Brackenridge (1854: 204) and P. kangdingense H.S.Kung & Li Bing Zhang ex Li Bing Zhang (2012: 58) in having bipinnate laminae, ovatelanceolate petiole scales, lanceolate rachis scales, and linear microscales. These four species are compared with one another in Table 1.Published as part of He, Hai & Zhang, Li-Bing, 2022, Polystichum gonggashanense (Dryopteridaceae): a new fern from Sichuan, China, pp. 207-212 in Phytotaxa 559 (2) on pages 207-210, DOI: 10.11646/phytotaxa.559.2.10, http://zenodo.org/record/702147

    Glaphyrocyclosorus erubesquirolicus X. Wan & Li Bing Zhang 2022, comb. nov.

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    × Glaphyrocyclosorus erubesquirolicus (W.C.Shieh & J.L.Tsai 1987: 7) X.Wan & Li Bing Zhang, comb. nov. Basionym: Thelypteris × erubesquirolica W.C.Shieh & J.L.Tsai (1987: 7) ≡ Cyclosorus × erubesquirolicus (W.C.Shieh & J.L.Tsai) J. Mazumdar (2013: 398) ≡ “ × Glaphypseudosorus erubesquirolicus (W.C.Shieh & J.L.Tsai) Hong M.Liu, Schuettp. & H.Schneid. ” (2020: 1000), nom. inval. Type:— Taiwan Island: Nantou Hsien, Wanghsiang, 17 December 1984, J.-L .Tsai T10379a-d (TCF). Notes: This nothospecies is a hybrid between Glaphyropteridopsis erubescens (Wallich ex Hooker 1863: 236) Ching (1963: 320) and Pseudocyclosorus esquirolii (Christ 1907: 144–145) Ching (1963: 324). This hybrid is an allotetraploid with 72 bivalent chromosomes, which has intermediate morphology of the parents (Shieh & Tsai 1987). Distribution: Taiwan Island.Published as part of Wan, Xia & Zhang, Li-Bing, 2022, Three new fern names of hybrid genera and four new combinations in Athyriaceae, Osmundaceae, and Thelypteridaceae, pp. 104-108 in Phytotaxa 575 (1) on page 106, DOI: 10.11646/phytotaxa.575.1.8, http://zenodo.org/record/740335
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