20,632 research outputs found
Encarsia imiza Geng & Li 2017, sp. nov.
Encarsia imiza Li & Geng, sp. nov. Figs 18–23 Type material. Holotype ♀ [on slide, NEFU]: CHINA, Liaoning Province, Anshan City, Qianshan Mountains, Gudaoguan, 19–21.IX.2015, Hui Geng, Xin-Yu Zhang, Ye Chen, YPT. Paratype. 1♀ [on slide, NEFU], CHINA, Liaoning Province, Fushun City, Yuanshuailin, 18.VI.2012, Hui Geng, Xiang-Xiang Jin, Jiang Liu, sweeping. Diagnosis. Female. Length, mesosoma plus metasoma, 0.58–0.66 mm. Head (Fig. 18) yellow with some brownish patches. Mesosoma (Fig. 20) mainly yellowish-brown with some pale yellow or brown patches. Metasoma (Fig. 20) pale yellow except petiole brown. Fore wing faintly infuscate behind submarginal and stigmal veins. Mid lobe of mesoscutum with 8 or 10 setae; placoid sensilla on scutellum moderately separated (Fig. 20). Fore wing (Fig. 21) 2.38–2.52× as long as wide, marginal fringe 0.23–0.25× as long as wing width. Mid tibial spur 0.57–0.61× as long as corresponding basitarsus. Ovipositor hardly exerted, 0.77–0.79× as long as mid tibia; third valvula 0.34× as long as second valvifer. Description. Female. Holotype. Length, mesosoma plus metasoma, 0.58 mm. Head yellow except clypeus, malar sulcus, a spot on each side of occipital foramen, and postocellar bars brown. Antenna (Fig. 19) pale yellow. Mesosoma yellowish-brown except side lobe pale yellow; pronotum, anterior margin of mid lobe, a patch on expanded part of each side lobe, inner sides of axillae, and propodeum brown. Wings (Figs 21, 22) hyaline, fore wing with a faint infuscation behind submarginal and stigmal veins, veins pale brown. Legs yellow (Fig. 23). Metasoma pale yellow except petiole brown. Head (Fig. 18) wider than mesosoma. Stemmaticum with transversely rugose sculpture. Mandible with two teeth and a truncation. Radicle (R), scape (S), pedicel (P) and 6 flagellomeres with the following ratios of length to width: R: 2.62, S: 4, P: 1.5, F1: 1.83, F2: 2.09, F3: 2.23, F4: 2.04, F5: 1.85 and F6: 1.92; relative lengths of R–F6 to length of F1: R: 1.03, S: 2.79, P: 1.03, F1: 1.00, F2: 1.33, F3: 1.42, F4: 1.48, F5: 1.45, and F6: 1.39; flagellomeres with the following numbers of longitudinal sensilla: F1: 0, F2: 1, F3: 2, F4: 3, F5: 3, F6: 3. Mesosoma 0.79× as long as metasoma (Fig. 20). Mid lobe of mesoscutum with 8 setae, each side lobe of mesoscutum with 3 setae. Distance between placoid sensilla on scutellum 4.89× maximum width of a sensillum. Distance between anterior pair of scutellar setae equal to distance between posterior pair. Fore wing (Fig. 21) 2.52× as long as wide, costal cell with 7 short setae, basal cell with 10 setae, marginal vein with 6 setae along anterior margin; marginal fringe 0.25× as long as wing width. Mid tibial spur 0.57× as long as corresponding basitarsus, and the latter 0.37× as long as mid tibia. Hind tibia 0.96× as long as mid tibia. Metasoma (Fig. 20) with petiole distinctly sculptured. T2–T7 with 1+1, 1+1, 1+1, 6, 1+4+1 (4 between cercal plates) and 4 setae, respectively. Ovipositor slightly exerted, apparently originating from base of T4, 0.79× as long as mid tibia, and 0.58× as long as mid tibia and basitarsus combined. Third valvula 0.34× as long as second valvifer. Male. Unknown. Host. Unknown. Variation. Sole paratype more infuscate on metasoma: T1 anteriorly, T5 completely, and T6 anteriorly brown; mid lobe of mesoscutum with 10 setae. Etymology. The specific name is an anagram of azimi, a similar species in this genus. Comments. Encarsia imiza resembles E. azimi Hayat in having similar body colour and the presence of internal sculpture in the cells forming the reticulation on the thoracic dorsum, but can be separated from the latter by the following characters: occiput largely yellow except for a brownish patch on each side of foramen (vs occiput completely brownish); mesosoma largely brownish-yellow (vs brown); all legs yellowish (vs fore and hind coxae brown basally); ovipositor 0.56–0.58× as long as mid tibia and basitarsus combined (vs 0.76×); gaster pale yellow (vs pale yellow with T1 across base dark brown).Published as part of Geng, Hui & Li, Cheng-De, 2017, Three new species of the Encarsia inaron (Hymenoptera: Aphelinidae) species group from China with a key to Chinese species, pp. 208-222 in Zootaxa 4306 (2) on page 215, DOI: 10.11646/zootaxa.4306.2.2, http://zenodo.org/record/84373
Encarsia dongbeiana Geng & Li 2017, sp. nov.
Encarsia dongbeiana Li & Geng, sp. nov. Figs 2–17 Type material. Holotype. ♀ [on slide, NEFU]: CHINA, Liaoning Province, Anshan City, Qianshan Mountains, Xianrentai, 20.IX.2015, Hui Geng, Xin-Yu Zhang, Ye Chen, sweeping. Paratypes. 1♀, 1♂ [on slides, NEFU], same data as holotype; 4♀, 4♂ [on slides, NEFU], CHINA, Heilongjiang Province, Mudanjiang City, Heixiazigou, 26.VIII.2015, Hui Geng, Yan Gao, Zhi-Guang Wu, sweeping; 1♀, 2♂ [on slides, NEFU], CHINA, Jilin Province, Wangqing County, Daxingou Town, Longshui Village, 24.VIII.2015, Hui Geng, Yan Gao, Zhi-Guang Wu, sweeping. Diagnosis. Female. Length, mesosoma plus metasoma, 0.57–0.71 mm. Head mainly yellow with postocellar bars dark brown and two short oblique bands above occipital foramen brown (Figs 2, 3). Mesosoma and metasoma entirely dark yellow with only pronotum pale brown. Mid lobe of mesoscutum with 8 or 9 setae; placoid sensilla on scutellum moderately separated (Fig. 7). Fore wing (Fig. 5) 2.52–2.74× as long as wide, marginal fringe 0.26–0.3× as long as wing width. Mid tibial spur 0.54–0.62× as long as corresponding basitarsus (Fig. 8). Ovipositor slightly exerted, about as long as mid tibia; third valvula 0.40–0.46× as long as second valvifer (Fig. 9). Male. Mostly brown with some paler parts. Antenna with last two flagellomeres partly fused (Fig. 12). Genitalia 0.75–0.84× as long as mid tibia. Description. Female. Holotype. Length, mesosoma plus metasoma, 0.61 mm. Head yellow except postocellar bars dark brown and two short oblique bands above occipital foramen brown. Antenna (Fig. 4) dark yellow with radicle and scape pale yellow and distal flagellomere brown. Mesosoma and metasoma entirely dark yellow with only pronotum pale brown. Wings (Figs 5, 6) including venation hyaline. Legs completely yellow. Head (cf. Fig. 2), as wide as mesosoma in dorsal view. Ocelli forming about an obtuse triangle, POL<OOL. Stemmaticum (cf. Fig. 3) with transversely rugose sculpture. Mandible with two teeth and a truncation (cf. Fig. 2). Radicle (R), scape (S), pedicel (P), and 6 flagellomeres with the following ratios of length to width: R: 2.91, S: 3.59, P: 1.55, F1: 1.78, F2: 2.1, F3: 2.2, F4: 1.96, F5: 1.74 and F6: 1.83; relative lengths of R–F6 to length of F1: R: 1.00, S: 2.63, P: 0.88, F1: 1.00, F2: 1.25, F3: 1.38, F4: 1.38, F5: 1.25, and F6: 1.31; flagellomeres with the following numbers of longitudinal sensilla: F1: 1, F2: 2, F3: 2, F4: 3, F5: 3, F6: 3. Mesosoma 0.68× as long as metasoma. Mid lobe of mesoscutum (Fig. 7) with 9 setae, each side lobe of mesoscutum with 3 setae. Distance between placoid sensilla on scutellum approximately 4.4× maximum width of a sensillum. Distance between anterior pair of scutellar setae 1.64× distance between posterior pair. Fore wing (Fig. 5) 2.59× as long as wide, costal cell with 10 short setae, basal cell with 6 setae, marginal vein with 6 setae along anterior margin; marginal fringe 0.27 × as long as wing width. Mid tibial spur 0.58× as long as corresponding basitarsus, the latter 0.38× as long as mid tibia. Hind tibia 1.04× as long as mid tibia (Fig.8). Metasoma (Fig. 9) with petiole (Fig. 10) sculptured. T2–T7 with 1+1, 1+1, 1+1, 2+2, 1+4+1 (4 between cercal plates) and 4 setae, respectively. Ovipositor (Fig. 9) slightly exerted, apparently originating from base of T3, 1.03× as long as mid tibia, and 0.75× as long as mid tibia and basitarsus combined. Third valvula 0.46× as long as second valvifer. Male. Essentially similar to female, differing mainly in colour, antennal structure and genitalia. Head (Fig. 11) yellow except malar sulcus infuscate, a stripe at level of occipital foramen brown, and postocellar bars dark brown. Mesosoma (Fig. 13) largely yellow with pronotum, mid lobe of mesoscutum anteriorly and mediolongitudinally, axillae and propodeum brown. Metasoma brown to dark brown with T6 posteriorly and T7 yellow. Antenna (Fig. 12) dark brown with abundant longitudinal sensilla on all flagellomeres, last two flagellomeres partly fused. Genitalia (Fig. 17) 0.75–0.84× as long as mid tibia. Host. Unknown. Variation. Length, mesosoma plus metasoma, 0.57–0.71 mm. Antenna dark yellow to yellowish-brown. Mesosoma and metasoma entirely dark yellow with pronotum, cercal plates and ovipositor stylet apically brown or pale brown. F1 0.94–1.14× as long as pedicel, without or with 1 longitudinal sensillum. Fore wing 2.52– 2.74× as long as wide, marginal fringe 0.26–0.3× as long as wing width. Basal cell with 6–10 setae. Marginal vein usually with 6 or 7 (rarely 5 or 8) long setae along anterior margin. Ovipositor 0.99–1.09× as long as mid tibia, and 0.72–0.79× as long as mid tibia and basitarsus combined. Third valvula 0.4–0.46× as long as second valvifer. Etymology. dongbei = the Chinese word for northeast; and refers to the distribution of the species in the northeastern part of China. Comments. E. dongbeiana is similar to E. aferi Schmidt & Polaszek in having the head and body mostly yellowish, but differs from the latter in: mesosoma and metasoma with only pronotum brownish (vs with some brownish patches on mesosoma, petiole and faint transverse band on T1); marginal fringe of fore wing 0.26– 0.3× as long as wing width (vs 0.2–0.21×); distance between anterior pair of scutellar setae 1.4–1.64× distance between posterior pair (vs 1×); ovipositor 0.99–1.09× as long as mid tibia (vs 0.85–0.91×); third valvula 0.4– 0.46× as long as second valvifer (vs 0.3–0.33). Encarsia dongbeiana also resembles those E. melanostoma Polaszek & Hernández with a completely yellow metasoma (colouration variable in melanostoma as noted in original description), but differs from the latter in different antennal formula, 1,1,4,2 (vs 1,1,3,3), presence of internal sculpture in the cells forming reticulation on mesoscutum (vs absent), pale clypeus and malar sulcus (vs dark) and more distantly placed anterior pair of scutellar setae than that between posterior pair, 1.4–1.64× (vs 0.8–1.1×).Published as part of Geng, Hui & Li, Cheng-De, 2017, Three new species of the Encarsia inaron (Hymenoptera: Aphelinidae) species group from China with a key to Chinese species, pp. 208-222 in Zootaxa 4306 (2) on pages 210-211, DOI: 10.11646/zootaxa.4306.2.2, http://zenodo.org/record/84373
Encarsia hayati Li & Geng, sp. nov.
Encarsia hayati Li & Geng, sp. nov. Figs 9–16 Type material. Holotype. ♀ [on slide], (NEFU), CHINA, Jiangxi Province, Shangrao City, Wuyishan Protection Station, 780m, 2.VII. 2013, Chao Zhang, sweeping. Paratype. 1♀ [on slide], CHINA, Hubei Province, Suizhou City, Santan, 14. VIII. 2015, Hui Geng, Yan Gao, Zhi-Guang Wu, sweeping. (NEFU). Diagnosis. Female. Length, mesosoma plus metasoma, 0.49–0.51 mm. Body pale yellow except clypeus, malar sulcus and postocellar bars brown. Antennal formula 1:1:3:3. F1 and F2 without longitudinal sensilla. Mid lobe of mesoscutum with 4 setae; each side lobe and axilla with 1 seta; placoid sensilla on scutellum narrowly separated; distance between anterior pair of scutellar setae distinctly less than that between posterior pair. Fore wing 3.35–3.49× as long as wide, sparsely setose, with a large asetose area around stigmal vein; marginal fringe 0.88–0.89× as long as wing width. Tarsal formula 5:4:5. Mid tibial spur 0.70–0.76× as long as corresponding basitarsus. Petiole with sculpture laterally. Ovipositor slightly exerted, 1.15–1.19× as long as mid tibia; third valvula 0.35–0.36× as long as second valvifer. Description. Female. Holotype. Length, mesosoma plus metasoma, 0.51 mm. Head and body including ovipositor and legs pale yellow except clypeus, malar sulcus and postocellar bars brown. Antennae yellow except last two segments slightly brown. Wings hyaline, venation pale brown. Head (Fig. 9), in frontal view, 1.77× as wide as high, and about as wide as mesosoma. Ocelli forming about an obtuse triangle, POL distinctly less than OOL. Maxillary and labial palpi 1-segmented. Mandibles with two teeth and a truncation. Eyes with fine and transparent setae. Frontovertex with short setae. Antennal formula, 1:1:3:3 (Fig. 10); radicle (R), scape (S), pedicel (P), 3 funicle segments (F1–F3) and 3 club segments (F4–F6) with the following ratios of length to width: R: 2.00, S: 4.05, P: 1.47, F1: 1.40, F2: 2.00, F3: 1.89, F4: 1.65, F5: 1.70 and F6: 2.33; relative lengths of segments R–F6 to length of F1: R: 1.15, S: 3.68, P: 1.61, F1: 1.00, F2: 1.61, F3: 1.95, F4: 1.90, F5: 1.95 and F6: 2.41; flagellum with the following numbers of longitudinal sensilla: F1: 0, F2: 0, F3: 2, F4: 3, F5: 3, F6: 3. Mesosoma (Fig. 11) 0.61× as long as metasoma. Mid lobe of mesoscutum and axillae weakly reticulate. Mid lobe of mesoscutum with 4 setae, each side lobe of mesoscutum with 1 seta. Axilla with 1 short seta. Scutellum 1.94× as wide as long, and 0.75× as long as mid lobe of mesoscutum. Distance between placoid sensilla on scutellum about own maximum width. Anterior pair of scutellar setae clearly shorter than posterior pair, and distance between anterior pair 0.7× that between posterior pair. Endophragma long and rounded at apex, extending to posterior margin of T1. Fore wing (Fig. 12) 3.49× as long as wide, sparsely setose, with a large asetose area around stigmal vein; costal cell with 4 short setae in basal half; basal cell with 2 setae, with proximal one distinctly shorter; submarginal vein with 2 setae; marginal vein 1.38× as long as submarginal vein, with 5 setae along anterior margin; marginal fringe 0.89× as long as wing width. Hind wing 8× as long as wide, marginal fringe 1.84× as long as wing width. Tarsal formula 5:4:5, mid leg with last two tarsal segments fused but indicated by a transverse suture (Fig. 13). Mid tibial spur 0.7× as long as corresponding basitarsus (Fig. 13), and the latter 0.34× as long as mid tibia. Hind tibia 0.99× as long as mid tibia. Metasoma with petiole (Fig. 15) sculptured laterally. T1–T5 with scale-like reticulation laterally. T2–T7 with 1+1, 1+1, 1+1, 2+2, 1+2+1 and 4 setae, respectively. T7 1.56× as wide as long. Ovipositor (Fig. 16) not exerted, apparently originating from middle of T3, 1.19× as long as mid tibia, and 0.89× as long as mid tibia and basitarsus combined. Third valvula 0.36× as long as second valvifer. Male. Unknown. Host. Unknown. Variation. Sole paratype with antennal F2 1.41× as long as wide, F3 1.63× as long as wide and with 2 longitudinal sensilla; fore wing with 4 setae along anterior margin; hind tibia 1.04× as long as mid tibia. Other characters the same as holotype. Etymology. This species is named in honor of Prof. Mohammad Hayat (Department of Zoology, Aligarh Muslim University, Uttar Pradesh, India) for his contributions to the study of Hymenoptera, Chalcidoidea. Comments. Mid tarsal structure of E. hayati is similar to that of the foregoing new species (see also comments under E. dianensis). Placement to species-group of this new species is uncertain. Among the species with a 5:4:5 tarsal formula and fore wing with a clear asetose area around the stigmal vein (meghalayana - and cubensis -group), E. hayati is unique by having a completely yellow body, and narrowly separated placoid sensilla on the scutellum. This species is closely related to E. dianensis based on similar structures of the mid tarsi, body color, and shape and characters of the fore wing, but can be separated from the latter by the differences given in the key and the comments under E. dianensis.Published as part of Geng, Hui & Li, Cheng-De, 2016, Five new species of genus Encarsia Förster from China (Hymenoptera: Aphelinidae), pp. 535-549 in Zootaxa 4162 (3) on pages 540-542, DOI: 10.11646/zootaxa.4162.3.7, http://zenodo.org/record/25468
Encarsia yunnana Li & Geng, sp. nov.
Encarsia yunnana Li & Geng, sp. nov. Figs 31–38 Type material. Holotype. ♀ [on slide], (NEFU), CHINA, Yunnan Province, Baoshan City, Tengchong County, Yinghe Village, 1820m, 17–19. VII. 2012, Xiang-Xiang Jin, Hui Geng, Chao Zhang, YPT. Paratypes. 3♀ [on slides], same data as holotype. (NEFU). Diagnosis. Female. Length, mesosoma plus metasoma, 0.5–0.66 mm. Ocelli forming a right to slightly obtuse triangle. POL distinctly less than OOL. Mandible with two teeth and a truncation. Maxillary and labial palpi 1- segmented. Antennal formula 1:1:4:2. Mid lobe of mesoscutum with 2 setae; each side lobe with 2 setae; axilla with 1 seta; distance between placoid sensilla on scutellum approximately 2–4× maximum width of a sensillum; distance between anterior pair of scutellar setae about equal to that between posterior pair. Fore wing 3.21–3.36× as long as wide, with an asetose area around stigmal vein; marginal fringe 0.67–0.8× as long as wing width. Tarsal formula 5:5:5. Mid tibial spur 0.88–0.97× as long as corresponding basitarsus. Petiole with sculpture laterally. T7 2.52–2.6× as wide as long. Ovipositor not or hardly exerted, 0.93–1.1× as long as mid tibia; third valvula 0.36– 0.4× as long as second valvifer. Description. Female. Holotype. Length, mesosoma plus metasoma, 0.66 mm. Head and body pale yellow except pronotum brown, petiole, T1 and T4–6 with very faint brownish bands. Ovipositor yellow. Antennae pale yellow. Fore wing hyaline, venation pale except stigmal vein mostly and base of submarginal vein slightly infuscate. Hind wing hyaline. Legs pale yellow. Head (Fig. 31) a little broader than mesosoma. Frontovertex 0.67× as broad as head width. Eyes with fine and transparent setae. Ocelli forming about a right triangle, POL distinctly less than OOL. Stemmaticum with reticulate sculpture. Mandible (Fig. 31) weakly dentate, with two teeth and a truncation. Maxillary and labial palpi 1- segmented. Antennae inserted at level of lower margin of eyes. Antennal formula, 1:1:4:2 (Fig. 32); radicle (R), scape (S), pedicel (P), 4 funicle segments (F1–F4) and 2 club segments (F5–F6) with the following ratios of length to width: R: 2.20, S: 4.13, P:1.44, F1: 1.83, F2: 2.42, F3: 1.88, F4: 1.75, F5: 1.73 and F6: 1.86; relative lengths of segments R–F6 to length of F1: R: 1.00, S: 3.00, P: 1.24, F1: 1.00, F2: 1.32, F3: 1.36, F4: 1.59, F5: 1.64 and F6: 1.90; flagellum with the following numbers of longitudinal sensilla: F1: 0, F2: 1, F3: 1, F4: 3, F5: 3, F6: 3. Mesosoma (cf. Fig. 33) 0.75× as long as metasoma. Mid lobe of mesoscutum and axillae indistinctly reticulate. Mid lobe of mesoscutum with 2 setae near posterior margin, each side lobe with 2 setae. Axilla with 1 seta. Scutellum 2.08× as wide as long, and 0.62× as long as mid lobe. Distance between placoid sensilla on scutellum approximately 4× maximum width of a sensillum. Anterior pair of scutellar setae distinctly shorter and thinner than posterior pair, and distance between anterior pair 0.92× that between posterior pair. Endophragma rounded at apex, extending to anterior margin of T2. Fore wing (Fig. 34) 3.3× as long as wide, with a small asetose area around stigmal vein; costal cell with 4 short setae in basal half; basal cell with 2 setae; submarginal vein with 2 setae; marginal vein 1.26× as long as submarginal vein, with 4 setae along anterior margin; marginal fringe 0.67× as long as wing width. Hind wing (Fig. 35) 7.69× as long as wide, marginal fringe 1.54× as long as wing width. Tarsal formula 5:5:5. Mid tibial spur 0.92× as long as corresponding basitarsus (Fig. 36), and the latter 0.28× as long as mid tibia. Hind tibia as long as mid tibia. Metasoma (Fig. 37) with petiole (Fig. 38) sculptured laterally. T1–T4 with scale-like reticulation laterally. T2– T7 with 1+1, 1+1, 1+1, 2+2, 1+2+1 and 4 setae, respectively. T7 2.59× as wide as long. Ovipositor not or hardly exerted, apparently originating from anterior margin of T4, 1.05× as long as mid tibia, and 0.83× as long as mid tibia and basitarsus combined. Third valvula 0.37× as long as second valvifer. Male. Unknown. Host. Unknown. Variation. Antennal segment F3 with 1–2 longitudinal sensilla. Distance between placoid sensilla on scutellum approximately 2–4× maximum width of a sensillum. Axillae completely yellow (slightly brownish at apex in one paratype). Marginal vein of fore wing with 4 or 5 (in one paratype with 6–8) setae along anterior margin. In one paratype, 1 seta present at apex of stigmal vein of fore wing. Etymology. The specific name is derived from the collection locality name. Comments. This new species is placed in the E. parvella- group as defined by Polaszek & Gill (2011), except for the host being unknown, in having all tarsi 5-segmented, fore wing with an asetose area around the stigmal vein, marginal fringe of the fore wing not longer than the wing width, and each side lobe of the mesoscutum with 2 setae. However, it is easily separated from all the species of the group by the following combination of characters: only 2 setae on the mid lobe of the mesoscutum, relatively densely setose fore wing disc, laterally sculptured petiole, and relatively short third valvula (less than 0.5× second valvifer). Encarsia yunnana resembles E. gerlingi Viggiani (1989) in similar body color, antennal structure, similar shape, setation, pigmentation on venation of the fore wing, and relatively short third valvulae, but is distinguished from the latter by: mid lobe of mesoscutum with 2 setae (vs 8 setae), marginal fringe of fore wing relatively long, 0.67–0.80× as long as wing width (vs shorter than half of wing width), and distance between anterior pair of scutellar setae about equal to that between posterior pair (vs distinctly shorter).Published as part of Geng, Hui & Li, Cheng-De, 2016, Five new species of genus Encarsia Förster from China (Hymenoptera: Aphelinidae), pp. 535-549 in Zootaxa 4162 (3) on pages 546-548, DOI: 10.11646/zootaxa.4162.3.7, http://zenodo.org/record/25468
Encarsia polymacula Geng & Li 2017, sp. nov.
<i>Encarsia polymacula</i> Li & Geng, sp. nov. <p>Figs 31–38</p> <p> <b>Type material.</b> Holotype ♀ [on slide, NEFU]: CHINA, Liaoning Province, Huludao City, Jianchang County, 4.VII.2012, Si-Zhu Liu, Jiang Liu, sweeping.</p> <p> <b>Diagnosis.</b> <i>Female</i>. Length, mesosoma plus metasoma, 0.63 mm. Head (Fig. 31) yellow except clypeus, malar sulcus, a patch on each side of occipital foramen and postocellar bars brown. Mesosoma (Fig. 33) largely yellow with some brown to dark brown patches. Metasoma (Fig. 33) dark yellow with petiole, T1 anteriorly, sides of T1– T4 brown, T5 and T6 largely brown with middle part paler. Mid lobe of mesoscutum (Fig.34) with 11 setae; placoid sensilla on scutellum moderately separated. Fore wing (Fig. 35) 2.5× as long as wide, marginal fringe 0.24× as long as wing width. Mid tibial spur 0.42× as long as corresponding basitarsus (Fig. 37). Petiole with distinct reticulate sculpture (Fig. 38). Ovipositor slightly exerted, 0.77× as long as mid tibia; third valvula 0.4× as long as second valvifer.</p> <p> <b>Description.</b> <i>Female</i>. Holotype. Length, mesosoma plus metasoma, 0.63 mm. Head yellow except clypeus, malar sulcus, a patch on each side of occipital foramen and postocellar bars brown. Antenna (Fig. 32) dark yellow with pedicel and last flagellomere brown. Mesosoma largely yellow with pronotum, anterior margin of mid lobe, a patch on expanded part of each side lobe, apex of axillae, and propodeum laterally brown, about posterior two thirds of notauli and anterior margin of scutellum dark brown. Fore wing, including costal cell, largely infuscate below venation, and darker below base of marginal vein and stigmal vein to more or less give an appearance of 2 cross bands, veins brown. Legs yellow. Metasoma dark yellow with petiole, T1 anteriorly, sides of T1–T4 brown, T5 and T6 largely brown with middle part paler.</p> <p>Head (Fig. 31) wider than mesosoma. Mandible with three acute teeth. Radicle (R), scape (S), pedicel (P), and 6 flagellomeres with the following ratios of length to width: R: 2.43, S: 4.16, P: 1.53, F1: 2, F2: 2.35, F3: 2.22, F4: 2.26, F5: 2.08 and F6: 2.35; relative lengths of R–F6 to length of F1: R: 0.85, S: 2.8, P: 0.82, F1: 1.00, F2: 1.15, F3: 1.25, F4: 1.3, F5: 1.33, and F6: 1.33; flagellomeres with the following numbers of longitudinal sensilla: F1: 0, F2: 1, F3: 1, F4: 3, F5: 3, F6: 2.</p> <p>Mesosoma 0.95× as long as metasoma (Fig. 33). Mid lobe of mesoscutum (Fig. 34) with 11 setae, each side lobe of mesoscutum with 3 setae. Distance between placoid sensilla on scutellum 5× maximum width of a sensillum. Distance between anterior pair of scutellar setae 1.16× distance between posterior pair. Fore wing (Fig. 35) 2.61× as long as wide, costal cell with 7 short setae, basal cell with 6 setae, marginal vein with 7 setae along anterior margin; marginal fringe 0.24 × as long as wing width. Mid tibial spur 0.5× as long as corresponding basitarsus, and the latter 0.39× as long as mid tibia. Hind tibia 0.96× as long as mid tibia (Fig. 37).</p> <p>Metasoma with petiole (Fig. 38) faintly sculptured. T2–T7 with 1+1, 1+1, 1+1, 2+2, 1+4+1 (4 between cercal plates) and 4 setae, respectively. Ovipositor slightly exerted, apparently originating from base of T3, 0.77× as long as mid tibia, and 0.55× as long as mid tibia and basitarsus combined. Third valvula 0.4× as long as second valvifer.</p> <p> <i>Male</i>. Unknown.</p> <p> <b>Host.</b> Unknown.</p> <p> <b>Etymology.</b> <i>Greek</i>: <i>poly</i> = many, much, multi-; <i>Latin</i>: <i>macula</i> = spot, stain; and refers to the many brown patches on the body.</p> <p> <b>Comments.</b> <i>Encarsia polymacula</i> is similar to <i>E. levadicola</i> Polaszek & Hernández in body colour and antennal structure, but differs from the latter by having the fore wing largely infuscate below the venation (<i>vs</i> completely hyaline), F1 2× as long as wide (<i>vs</i> 1.5×), distance between anterior pair of scutellar setae 1.16× that between posterior pair (<i>vs</i> 0.8–1×), mid tibial spur 0.50× as long as corresponding basitarsus (<i>vs</i> 0.66–0.74×) and ovipositor 0.77× as long as mid tibia (<i>vs</i> 1–1.2×).</p>Published as part of <i>Geng, Hui & Li, Cheng-De, 2017, Three new species of the Encarsia inaron (Hymenoptera: Aphelinidae) species group from China with a key to Chinese species, pp. 208-222 in Zootaxa 4306 (2)</i> on page 219, DOI: 10.11646/zootaxa.4306.2.2, <a href="http://zenodo.org/record/843736">http://zenodo.org/record/843736</a>
Encarsia huangi Li & Geng, sp. nov.
Encarsia huangi Li & Geng, sp. nov. Figs 23–30 Type material. Holotype. ♀ [on slide], (NEFU), CHINA, Yunnan Province, Xishuangbanna, Mengla County, Menglun Town, Nanxing, 13. II. 2014, Guo-Hao Zu, Zhong-Ping Xiong, sweeping. Diagnosis. Female. Length, mesosoma plus metasoma, 0.45 mm. Frontovertex 0.59× as broad as head width; ocelli forming about an obtuse triangle. Mandible with two teeth and a truncation; maxillary and labial palpi 1- segmented. Antennal formula 1:1:3:3. F1 and F2 without longitudinal sensilla. Mid lobe of mesoscutum with 4 setae; each side lobe with 2 setae; axilla with 1 seta; placoid sensilla on scutellum narrowly separated; distance between anterior pair of scutellar setae distinctly shorter than that between posterior pair. Fore wing 3.57× as long as wide, sparsely setose, with two large asetose areas, one around stigmal vein and the other at apex of wing disc; marginal fringe about as long as wing width; basal cell asetose. Tarsal formula 5:5:5. Mid tibial spur 0.83× as long as corresponding basitarsus. Petiole finely sculptured. T7 1.83× as wide as long. Ovipositor not exerted, 0.82× as long as mid tibia; third valvula 0.4× as long as second valvifer. Description. Female. Holotype. Length, mesosoma plus metasoma, 0.45 mm. Head and body (Fig. 28) including legs almost entirely pale yellow, but antennal clava a little darker and distal part of ovipositor stylet brownish. Wings mostly hyaline, except below marginal vein narrowly and apical wing margin very narrowly infuscate, venation brown. Head (Fig. 23), in frontal view, 1.56× as wide as high, and about as wide as mesosoma. Frontovertex 0.59× as broad as head width. Eyes with fine and transparent setae. Ocelli forming about an obtuse triangle. Mandible with two teeth and a truncation. Maxillary and labial palpi 1-segmented. Antennae inserted at level of lower margin of eyes. Antennal formula, 1:1:3:3 (Fig. 24); radicle (R), scape (S), pedicel (P), 3 funicle segments (F1–F3) and 3 club segments (F4–F6) with the following ratios of length to width: R: 2.13, S: 3.25, P: 1.60, F1: 1.33, F2: 1.54, F3: 1.50, F4: 1.52, F5: 1.40 and F6: 1.68; relative lengths of segments R–F6 to length of F1: R: 1.25, S: 3.25, P: 1.50, F1: 1.00, F2: 1.25, F3: 1.50, F4: 1.75, F5: 1.75, and F6: 2.00; flagellum with the following numbers of longitudinal sensilla: F1: 0, F2: 0, F3: 1, F4: 3, F5: 3, F6: 3. Mesosoma (Fig. 25) 0.64× as long as metasoma. Mid lobe of mesoscutum and axillae weakly reticulate. Mid lobe of mesoscutum with 4 setae, each side lobe of mesoscutum with 2 setae. Axilla with 1 short seta. Scutellum 2.48× as wide as long, and 0.53× as long as mid lobe of mesoscutum. Placoid sensilla on scutellum separated slightly, by less than own maximum width. Anterior pair of scutellar setae clearly shorter than posterior pair, and distance between anterior pair 0.52× that between posterior pair. Endophragma long and rounded at apex, extending to posterior margin of T2. Fore wing (Fig. 26) 3.57× as long as wide, sparsely setose, with two large asetose areas, one around stigmal vein and the other at apex of wing disc; costal cell with 4 short setae in basal half; basal cell asetose; submarginal vein with 2 setae; marginal vein 1.38× as long as submarginal vein, with 5 setae along anterior margin; marginal fringe as long as wing width. Tarsal formula 5:5:5. Mid tibial spur 0.83× as long as corresponding basitarsus (Fig. 27), and the latter 0.28× as long as mid tibia. Hind tibia 0.93× as long as mid tibia. Metasoma (Fig. 28) with petiole (Fig. 29) finely sculptured. T1–T5 with scale-like reticulation laterally. T2–T7 with 1+1, 1+1, 1+1, 2+2, 1+2+1 and 4 setae, respectively. T7 1.83× as wide as long. Ovipositor (Fig. 30) not exerted, apparently originating from posterior margin of T4, 0.82× as long as mid tibia, and 0.64× as long as mid tibia and basitarsus combined. Third valvula 0.40× as long as second valvifer. Male. Unknown. Host. Unknown. Etymology. This species is named in honor of Prof. Huang Jian (Plant Protection College of Fujian Agriculture and Forestry University, China) for his contributions to the study of Hymenoptera, Aphelinidae. Comments. This new species is placed in the E. citrina- group (Viggiani & Mazzone 1979; Hayat 1989) based on the shape of the fore wing, longer marginal fringe, presence of an asetose area around the stigmal vein, mid lobe of mesoscutum with 4 setae, petiole with fine sculpture, submarginal vein with 2 setae, and mid tarsi 5-segmented, but differs from all the other species of this group (citrina, lounsburyi, curtifuniculata, gracilens,?flava,?fusca,?latipennis) mainly by the narrowly separated placoid sensilla on the scutellum, and entirely pale yellow body. Further differences are listed in the key.Published as part of Geng, Hui & Li, Cheng-De, 2016, Five new species of genus Encarsia Förster from China (Hymenoptera: Aphelinidae), pp. 535-549 in Zootaxa 4162 (3) on pages 544-546, DOI: 10.11646/zootaxa.4162.3.7, http://zenodo.org/record/25468
Encarsia dianensis Li & Geng, sp. nov.
Encarsia dianensis Li & Geng, sp. nov. Figs 1–8 Type material. Holotype. ♀ [on slide], (NEFU), CHINA, Yunnan Province, Ruili City, Nanjingli Village, 26–27. IV. 2013, Xiang-Xiang Jin, Guo-Hao Zu, Chao Zhang, YPT. Paratypes. 5♀ [on slides], CHINA, Yunnan Province, Xishuangbanna, Mengla County, Menglun Town, Nanxing, 12–14. II. 2014, Guo-Hao Zu, Zhong-Ping Xiong, YPT; 2♀ [on slides], CHINA, Yunnan Province, Puer City, Manxieba Village, 17–18. IV. 2013, Xiang-Xiang Jin, Guo-Hao Zu, Chao Zhang, YPT. (NEFU). Diagnosis. Female. Length, mesosoma plus metasoma, 0.42–0.52 mm. Body entirely yellow to pale yellow, sometimes pronotum, petiole and T1 slightly brown. Antennal formula 1:1:3:3. Mid lobe of mesoscutum with 4 or 5 setae; each side lobe and axilla with 1 seta; placoid sensilla on scutellum widely separated; distance between anterior pair of scutellar setae distinctly greater than that between posterior pair. Fore wing 3.36–3.67× as long as wide, sparsely setose, with a large asetose area around stigmal vein; marginal fringe 0.77–0.97× as long as wing width. Tarsal formula 5:4:5. Mid tibial spur 0.79–0.84× as long as corresponding basitarsus. Petiole smooth. T7 2.2–2.38× as wide as long. Ovipositor slightly exerted, 0.58–0.7× as long as mid tibia; third valvula 0.5–0.6× as long as second valvifer. Description. Female. Holotype. Length, mesosoma plus metasoma, 0.49 mm. Head and body, including ovipositor and legs entirely pale yellow; pedicel and flagellum pale brown, distal segments slightly darker. Wings hyaline, venation pale brown. Head (Fig. 1), in frontal view, 1.52× as wide as high, a little broader than mesosoma. Frontovertex 0.64× as broad as head width. Eyes with fine and transparent setae. Ocelli forming about an obtuse triangle, POL distinctly less than OOL. Stemmaticum reticulate. Mandible with two teeth and a truncation. Maxillary and labial palpi 1- segmented. Antennae inserted at level of lower margin of eyes. Distance between toruli about 0.5× distance from torulus to eye margin. Antennal formula, 1:1:3:3 (cf. Fig. 2); pedicel (P), 3 funicle segments (F1–F3) and 3 club segments (F4–F6) with the following ratios of length to width: P: 1.50, F1: 1.25, F2: 1.24, F3: 1.20, F4: 1.21, F5: 1.23 and F6: 1.57; relative lengths of segments P–F6 to length of F1: P: 1.50, F1: 1.00, F2: 1.05, F3: 1.20, F4: 1.45, F5: 1.60, and F6: 2.00; flagellum with the following numbers of longitudinal sensilla: F1: 0, F2: 0, F3: 1, F4: 2, F5: 3, F6: 3. Mesosoma (cf. Fig. 3) 0.7× as long as metasoma. Mid lobe of mesoscutum and axillae reticulate. Mid lobe of mesoscutum with 4 setae, each side lobe of mesoscutum with 1 seta. Axilla with 1 short seta. Scutellum 2.19× as wide as long, and 0.64× as long as mid lobe. Distance between placoid sensilla on scutellum approximately 7× maximum width of a sensillum. Anterior pair of scutellar setae distinctly shorter than posterior pair, and distance between anterior pair 1.48× that between posterior pair. Endophragma long and rounded at apex, extending to middle of T2. Fore wing (Fig. 4) 3.51× as long as wide, sparsely setose, with a large asetose area around stigmal vein; costal cell with 4 short setae in basal half; basal cell with 1 seta; submarginal vein with 2 setae; marginal vein 1.26× as long as submarginal vein, with 4 setae along anterior margin; marginal fringe 0.88 × as long as wing width. Tarsal formula 5:4:5, mid leg with last two tarsal segments fused but indicated by a transverse suture (Fig. 5). Mid tibial spur 0.82× as long as corresponding basitarsus (Fig. 5), and the latter 0.28× as long as mid tibia. Hind tibia 0.95× as long as mid tibia. Metasoma (Fig. 8) with petiole smooth. T1–T5 with scale-like reticulation laterally. T2–T7 with 1+1, 1+1, 1+1, 2+2, 1+2+1 and 4 setae, respectively. T7 2.38× as wide as long. Ovipositor (Fig. 8) slightly exerted, apparently originating from middle of T5, 0.64× as long as mid tibia, and 0.5× as long as mid tibia and basitarsus combined. Third valvula 0.5× as long as second valvifer. Male. Unknown. Host. Unknown. Variation. Length, mesosoma plus metasoma, 0.42–0.52 mm. Body entirely yellow to pale yellow, occasionally pronotum, petiole and T1 slightly pale brown. F1 1.13–1.46× as long as wide, and 0.64–0.95× as long as F2. F6 1.57–2.09× as long as wide. Number of longitudinal sensilla on F1–F 6 in two paratypes from Puer: F1: 0, F2: 1, F3: 2, F4: 3, F5: 3, F6: 3. Fore wing 3.36–3.67× as long as wide, marginal fringe 0.77–0.97× as long as wing width. Mid lobe of mesoscutum with 4 setae (5 in one paratype). Mid tibial spur 0.79–0.84× as long as corresponding basitarsus, and the latter 0.28–0.33× as long as mid tibia. Hind tibia 0.87–0.95× as long as mid tibia. Ovipositor 0.58–0.7× as long as mid tibia, and 0.45–0.55× as long as mid tibia and basitarsus combined. Third valvula 0.5–0.6× as long as second valvifer. Etymology. Chinese: dian = Yunnan Province, which refers to the distribution of the species in the Yunnan Province of China. Comments. In situations where the mid tarsi have the last two segments partly fused, but indicated either by a transverse suture or a distinct constriction, most (if not all) authors usually regarded the tarsi as 4-segmented. This includes Hill (1970) for E. africana (in his fig. 6), Polaszek et al. (2004) for E. dispersa (in their fig. 9B), Myartseva (2007) for E. flaviceps (in his fig. 13), Myartseva & Evans (2008) for E. florena (in their fig. 128), and Myartseva et al. (2012) for E. xilitla (in their fig. 3). Females of E. dianensis also have the last two tarsal segments of the mid leg partly fused and indicated by a transverse suture (Figs 5, 6, 7), so we regarded it as 4-segmented. Encarsia dianensis is placed in the E. cubensis -group (Evans & Polaszek 1998) based on a 5:4:5 tarsal formula, fore wing with large asetose area around the stigmal vein, marginal fringe of the fore wing not longer than the maximum wing width, widely separated placoid sensilla on the scutellum, smooth petiole, 3-segmented antennal clava, F1 subquadrate and shorter than the pedicel, and a short and slightly exerted ovipositor. The E. cubensis -group currently contains 8 species worldwide (Evans & Polaszek 1998; Myartseva et al. 2013). Our new species is easily separated from all the species of this group and related species by its entirely yellow head and pale body. In all other species of the group, the head and at least the anterior half of the mesoscutum are dark. This species is closely related to E. hayati, but can be distinguished from the latter by the following combination of characters: antennal flagellum stout, as in Fig. 3 (vs slender, as in Fig. 10); distance between anterior pair of scutellar setae distinctly greater than that between posterior pair (vs distinctly less), placoid sensilla on scutellum widely separated (vs narrowly separated); ovipositor distinctly shorter than mid tibia (vs longer than), third valvula 0.5–0.6× as long as second valvifer (vs 0.35–0.36×). If E. dianensis is regarded as having 5-segmented mid tarsi, it could be related to E. flavescens Huang & Polaszek because of having similar body color, fore wing with a bare strip along the wing margin beginning from stigmal vein and ending at about distal end of retinaculum and similar numbers of setae on the mid and side lobes of the mesoscutum, but can be distinguished from the latter by: marginal fringe of fore wing long, 0.77–0.97× as long as disc width (vs 0.47×); placoid sensilla on scutellum separated by 7× maximum width of a sensillum (vs 3– 4×); ovipositor very short, 0.45–0.55× as long as mid tibia and basitarsus combined (vs 1.07×).Published as part of Geng, Hui & Li, Cheng-De, 2016, Five new species of genus Encarsia Förster from China (Hymenoptera: Aphelinidae), pp. 535-549 in Zootaxa 4162 (3) on pages 536-539, DOI: 10.11646/zootaxa.4162.3.7, http://zenodo.org/record/25468
Two new species and a new record of the Encarsia longifasciata-group (Hymenoptera, Aphelinidae) from Malaysia and China
The genus Encarsia Förster, 1878, which is the largest genus of the family Aphelinidae, contains 453 valid species worldwide. Most species of Encarsia with known biology are primary endoparasitoids of Aleyrodidae and Diaspididae.
Species of the Encarsia longifasciata-group from Malaysia and China are reviewed. This is the first record of this group from Malaysia. Two new species, E. borneensis Geng & Li sp. n. and E. pauroseta Geng & Li sp. n., are described and illustrated. Encarsia longifasciata is newly recorded from Malaysia (Borneo). An updated key to the longifasciata-group species (females) worldwide is provided.Copyright © Geng H et al. This is an open access article distributed under the terms of the Creative Commons Attribution License (CC BY 4.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited. The attached file is the published version of the article.NHM Repositor
The expression of tachykinin receptors in the human lower esophageal sphincter
Abstract not availableKe Zhang, Que T. Chen, Jing H. Li, Xian Geng, Jun F. Liu, He F. Li, Yong Feng, Jia L. Li, Paul A. Dre
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