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    FIGURE. Conduplicate-spiral (CS) and cupulate (CL) stigma types of Acanthostachys, the different subgenera of Aechmea, and Karawata. A. Acanthostachys pitcairnioides (CS, Leme 483). B. Aechmea aculeatosepala (CS, Leme 3235). C–F. Aechmea subg. Aechmea. C. A. angustifolia (CS, Leme 8632). D. A. bracteata (CL, Leme 3466). E. A. phanerophlebia (CS, Leme 1472). F. A. tocantina (CS, Leme 143). G–I. Aechmea of the "Streptocalycoid complex". G. A. hoppii (CS, Leme 2339). H. A. longifolia (CS, Leme 116). I. A. vallerandii (CS, Leme 8693). J–K. Aechmea subg. Platyaechmea s.l. J. A. cucullata (CS, Leme 3462). K. A. manzanareziana (CS, Leme 3125). L. Aechmea of the "Gravisia complex": A. aquilega (CS, Leme 9516). M–N, P. Aechmea subg. Chevaliera s.l. M. A. cariocae (CS, Leme 5196). N. A. castanea (CS, Leme 1142). O. Karawata multiflora (CS, Leme 6500). P. A. ornata (CS, Leme 6760). Q–R. Aechmea subg. Macrochordion. Q. A. alba (CS, Leme 8232); R. A. bromeliifolia (CS, Leme 7951). S–T. Aechmea subg. Platyaechmea s.str. S. A. caesia (CS, Leme 3588); T. A. fasciata var. purpurea (CS, Leme 8639). U–V. Aechmea subg. Lamprococcus. U. A. capixabae (CS, Leme 6948). V. A. fulgens (CS, Leme 6209). W. Aechmea subg. Ortgiesia: A. candida (CS, Leme 9113). X. Aechmea subg. Pothuava s.str.: A. nudicaulis (CS, Leme 1436). Bars = 1 mm. in Re-evaluation of the Amazonian Hylaeaicum (Bromeliaceae: Bromelioideae) based on neglected morphological traits and molecular evidence

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    FIGURE. Conduplicate-spiral (CS) and cupulate (CL) stigma types of Acanthostachys, the different subgenera of Aechmea, and Karawata. A. Acanthostachys pitcairnioides (CS, Leme 483). B. Aechmea aculeatosepala (CS, Leme 3235). C–F. Aechmea subg. Aechmea. C. A. angustifolia (CS, Leme 8632). D. A. bracteata (CL, Leme 3466). E. A. phanerophlebia (CS, Leme 1472). F. A. tocantina (CS, Leme 143). G–I. Aechmea of the "Streptocalycoid complex". G. A. hoppii (CS, Leme 2339). H. A. longifolia (CS, Leme 116). I. A. vallerandii (CS, Leme 8693). J–K. Aechmea subg. Platyaechmea s.l. J. A. cucullata (CS, Leme 3462). K. A. manzanareziana (CS, Leme 3125). L. Aechmea of the "Gravisia complex": A. aquilega (CS, Leme 9516). M–N, P. Aechmea subg. Chevaliera s.l. M. A. cariocae (CS, Leme 5196). N. A. castanea (CS, Leme 1142). O. Karawata multiflora (CS, Leme 6500). P. A. ornata (CS, Leme 6760). Q–R. Aechmea subg. Macrochordion. Q. A. alba (CS, Leme 8232); R. A. bromeliifolia (CS, Leme 7951). S–T. Aechmea subg. Platyaechmea s.str. S. A. caesia (CS, Leme 3588); T. A. fasciata var. purpurea (CS, Leme 8639). U–V. Aechmea subg. Lamprococcus. U. A. capixabae (CS, Leme 6948). V. A. fulgens (CS, Leme 6209). W. Aechmea subg. Ortgiesia: A. candida (CS, Leme 9113). X. Aechmea subg. Pothuava s.str.: A. nudicaulis (CS, Leme 1436). Bars = 1 mm.Published as part of Leme, Elton M. C., Zizka, Georg, Paule, Juraj, Santoro, Julián Aguirre-, Heller, Sascha, Ramírez-Morillo, Ivón M., Halbritter, Heidemarie, Mariath, Jorge E. A., De, Jordano D. T. & Forzza, Rafaela C., 2021, Re-evaluation of the Amazonian Hylaeaicum (Bromeliaceae: Bromelioideae) based on neglected morphological traits and molecular evidence, pp. 1-60 in Phytotaxa 499 (1) on page 29, DOI: 10.11646/phytotaxa.499.1.1, http://zenodo.org/record/542441

    FIGURE. Fruits of the genera and subgenera of Bromelioideae. A. Acanthostachys pitcairnioides (Leme 483). B. Acanthostachys strobilacea (Leme 8306). C. Aechmea subg. Platyaechmea s.str.: A. caesia (Leme 3588). D. Aechmea subg. Lamprococcus: A capixabae (Leme 6948). E. Aechmea of the "Streptocalycoid complex": A. nidularioides (Leme 2150). F. Wittmackia lingulatoides (Leme 9585). G–H. Aechmea of the "Gravisia complex". G. A. aquilega (Leme 9516). H. A. blanchetiana (Leme 8890). I–L. Aechmea subg. Chevaliera s.l. I. A. castanea (Leme 6940). J. A. digitata (Leme 4019). K. A. fernandae (Leme 4492). L. A. ornata (Leme 6760). M. Greigia stenolepis (Leme 9738). N. Bromelia antiacantha (Leme 258). O. Bromelia aff. reversacantha (Leme 4609). P–Q. Billbergia subg. Billbergia. P. B. amoena var. amoena (Leme 6168). Q. B. amoena var. stolonifera (Leme 215-B). R. Billbergia subg. Helicodea: B. zebrina (Leme 128). S. Cryptanthus tabuleiricola (Leme 8014). T. Billbergia (Pseudaechmea) viridiflora (SEL 99-026). U. Portea petropolitana var. noetiigii (Leme 5277). V. Hohenbergia lanata (Leme 9625). W. Quesnelia quesneliana (Leme 107). X. Fernseea bocainensis (Leme 1422). Y. Disteganthus lateralis (Leme 9390). Z. Neoglaziovia variegata (Leme 9631). a. Aechmea subg. Pothuava s.l.: A. roberto-seidelii (Leme 1203). b. Aechmea subg. Macrochordion: A. bruggeri (Leme 4765). c. Araeococcus flagellifolius (Leme 9501). Bars = 10 mm. in Re-evaluation of the Amazonian Hylaeaicum (Bromeliaceae: Bromelioideae) based on neglected morphological traits and molecular evidence

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    FIGURE. Fruits of the genera and subgenera of Bromelioideae. A. Acanthostachys pitcairnioides (Leme 483). B. Acanthostachys strobilacea (Leme 8306). C. Aechmea subg. Platyaechmea s.str.: A. caesia (Leme 3588). D. Aechmea subg. Lamprococcus: A capixabae (Leme 6948). E. Aechmea of the "Streptocalycoid complex": A. nidularioides (Leme 2150). F. Wittmackia lingulatoides (Leme 9585). G–H. Aechmea of the "Gravisia complex". G. A. aquilega (Leme 9516). H. A. blanchetiana (Leme 8890). I–L. Aechmea subg. Chevaliera s.l. I. A. castanea (Leme 6940). J. A. digitata (Leme 4019). K. A. fernandae (Leme 4492). L. A. ornata (Leme 6760). M. Greigia stenolepis (Leme 9738). N. Bromelia antiacantha (Leme 258). O. Bromelia aff. reversacantha (Leme 4609). P–Q. Billbergia subg. Billbergia. P. B. amoena var. amoena (Leme 6168). Q. B. amoena var. stolonifera (Leme 215-B). R. Billbergia subg. Helicodea: B. zebrina (Leme 128). S. Cryptanthus tabuleiricola (Leme 8014). T. Billbergia (Pseudaechmea) viridiflora (SEL 99-026). U. Portea petropolitana var. noetiigii (Leme 5277). V. Hohenbergia lanata (Leme 9625). W. Quesnelia quesneliana (Leme 107). X. Fernseea bocainensis (Leme 1422). Y. Disteganthus lateralis (Leme 9390). Z. Neoglaziovia variegata (Leme 9631). a. Aechmea subg. Pothuava s.l.: A. roberto-seidelii (Leme 1203). b. Aechmea subg. Macrochordion: A. bruggeri (Leme 4765). c. Araeococcus flagellifolius (Leme 9501). Bars = 10 mm.Published as part of Leme, Elton M. C., Zizka, Georg, Paule, Juraj, Santoro, Julián Aguirre-, Heller, Sascha, Ramírez-Morillo, Ivón M., Halbritter, Heidemarie, Mariath, Jorge E. A., De, Jordano D. T. & Forzza, Rafaela C., 2021, Re-evaluation of the Amazonian Hylaeaicum (Bromeliaceae: Bromelioideae) based on neglected morphological traits and molecular evidence, pp. 1-60 in Phytotaxa 499 (1) on page 36, DOI: 10.11646/phytotaxa.499.1.1, http://zenodo.org/record/542441

    FIGURE. Flowers and longitudinal/cross sections of ovaries of the genera of the "Nidularioid complex" and Aechmea alliance. A. Neoregelia leucophoea (Leme 1107). B–C. Neoregelia subg. Protoregelia: N. longisepala (Leme 3049). D–E. Canistropsis burchellii (Leme 1301). F–G. Canistropsis microps (Leme 9017). H–J. Canistrum subg. Canistrum: C. aurantiacum (Leme 5028). K–L. Canistrum subg. Cucullatanthus: C. auratum (Leme 5803). M–N. Edmundoa ambigua (Leme 1073). O–P. Eduandrea selloana (Leme 7630). Q–R. Nidularium camposportoi var. robustum (Leme 3772). S–T. Nidularium rutilans (Leme 1072). U–V. Wittrockia superba (Leme 4751). W– X. Wittrockia gigantea (Leme 1289). Y–a. Aechmea aculeatosepala (Leme 3235). b–c. Aechmea castelnavii (Leme 4168). d–e. Aechmea hoppii (Leme 2339). Bars = 5 mm. in Re-evaluation of the Amazonian Hylaeaicum (Bromeliaceae: Bromelioideae) based on neglected morphological traits and molecular evidence

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    FIGURE. Flowers and longitudinal/cross sections of ovaries of the genera of the "Nidularioid complex" and Aechmea alliance. A. Neoregelia leucophoea (Leme 1107). B–C. Neoregelia subg. Protoregelia: N. longisepala (Leme 3049). D–E. Canistropsis burchellii (Leme 1301). F–G. Canistropsis microps (Leme 9017). H–J. Canistrum subg. Canistrum: C. aurantiacum (Leme 5028). K–L. Canistrum subg. Cucullatanthus: C. auratum (Leme 5803). M–N. Edmundoa ambigua (Leme 1073). O–P. Eduandrea selloana (Leme 7630). Q–R. Nidularium camposportoi var. robustum (Leme 3772). S–T. Nidularium rutilans (Leme 1072). U–V. Wittrockia superba (Leme 4751). W– X. Wittrockia gigantea (Leme 1289). Y–a. Aechmea aculeatosepala (Leme 3235). b–c. Aechmea castelnavii (Leme 4168). d–e. Aechmea hoppii (Leme 2339). Bars = 5 mm.Published as part of Leme, Elton M. C., Zizka, Georg, Paule, Juraj, Santoro, Julián Aguirre-, Heller, Sascha, Ramírez-Morillo, Ivón M., Halbritter, Heidemarie, Mariath, Jorge E. A., De, Jordano D. T. & Forzza, Rafaela C., 2021, Re-evaluation of the Amazonian Hylaeaicum (Bromeliaceae: Bromelioideae) based on neglected morphological traits and molecular evidence, pp. 1-60 in Phytotaxa 499 (1) on page 22, DOI: 10.11646/phytotaxa.499.1.1, http://zenodo.org/record/542441

    FIGURE. Seeds of genera and subgenera of Bromelioideae. A–C. Bromelia aff. reversacantha (Leme 4609). A. Longitudinal section of the fruits with seeds connected to placenta by the funicular appendage. B. Seeds with the funicular appendage manually detached. C. Frontal view of the distal funicular pole of the seeds with the funicular appendage backwardly inrolled to the seeds and attached under an aril-like coat. D. Disteganthus lateralis (Leme 9390). E. Aechmea of the "Streptocalycoid complex": A. nidularioides (Leme 2150). F. Aechmea subg. Chevaliera s.l.: A. digitata (Leme 4019). G. Araeococcus subg. Araeococcus: A. flagellifolius (Leme 9501). H. Pseudaraeococcus lageniformis (Leme 9603). I. Cryptanthus pickelii (Leme 3873). J. Fernseea bocainensis (Leme 1422). K. Greigia stenolepis (Leme 9738). L. Neoglaziovia variegata (Leme 9631). M. Aechmea subg. Chevaliera s.l.: A. ornata (Leme 6760). N. Billbergia subg. Billbergia: B. amoena var. stolonifera (Leme 215-B). O. Billbergia subg. Helicodea: B. zebrina (Leme 128). P. Quesnelia subg. Quesnelia: Q. quesneliana (Leme 107). Q. Portea petropolitana var. noetiigii (Leme 5277). R. Billbergia subg. Billbergia: B. tweedieana (Leme 278). S. Quesnelia subg. Billbergiopsis: Q. liboniana (Leme 2361). T. Wittmackia lingulatoides (Leme 9585). Bars = 1 mm. in Re-evaluation of the Amazonian Hylaeaicum (Bromeliaceae: Bromelioideae) based on neglected morphological traits and molecular evidence

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    FIGURE. Seeds of genera and subgenera of Bromelioideae. A–C. Bromelia aff. reversacantha (Leme 4609). A. Longitudinal section of the fruits with seeds connected to placenta by the funicular appendage. B. Seeds with the funicular appendage manually detached. C. Frontal view of the distal funicular pole of the seeds with the funicular appendage backwardly inrolled to the seeds and attached under an aril-like coat. D. Disteganthus lateralis (Leme 9390). E. Aechmea of the "Streptocalycoid complex": A. nidularioides (Leme 2150). F. Aechmea subg. Chevaliera s.l.: A. digitata (Leme 4019). G. Araeococcus subg. Araeococcus: A. flagellifolius (Leme 9501). H. Pseudaraeococcus lageniformis (Leme 9603). I. Cryptanthus pickelii (Leme 3873). J. Fernseea bocainensis (Leme 1422). K. Greigia stenolepis (Leme 9738). L. Neoglaziovia variegata (Leme 9631). M. Aechmea subg. Chevaliera s.l.: A. ornata (Leme 6760). N. Billbergia subg. Billbergia: B. amoena var. stolonifera (Leme 215-B). O. Billbergia subg. Helicodea: B. zebrina (Leme 128). P. Quesnelia subg. Quesnelia: Q. quesneliana (Leme 107). Q. Portea petropolitana var. noetiigii (Leme 5277). R. Billbergia subg. Billbergia: B. tweedieana (Leme 278). S. Quesnelia subg. Billbergiopsis: Q. liboniana (Leme 2361). T. Wittmackia lingulatoides (Leme 9585). Bars = 1 mm.Published as part of Leme, Elton M. C., Zizka, Georg, Paule, Juraj, Santoro, Julián Aguirre-, Heller, Sascha, Ramírez-Morillo, Ivón M., Halbritter, Heidemarie, Mariath, Jorge E. A., De, Jordano D. T. & Forzza, Rafaela C., 2021, Re-evaluation of the Amazonian Hylaeaicum (Bromeliaceae: Bromelioideae) based on neglected morphological traits and molecular evidence, pp. 1-60 in Phytotaxa 499 (1) on page 41, DOI: 10.11646/phytotaxa.499.1.1, http://zenodo.org/record/542441

    Hylaeaicum levianum Leme & Forzza 2021, comb. nov.

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    <i>Hylaeaicum levianum</i> (L.B. Sm.) Leme & Forzza, <i>comb. nov.</i> <p> Basionym:— <i>Neoregelia leviana</i> L.B. Sm., <i>Phytologia</i> 16: 460, pl. 1, fig. 5. 1968. Type:— BRAZIL. Amazonas: Rio Caubouris, 1967, <i>M. Mee s.n.</i> (holotype US!).</p>Published as part of <i>Leme, Elton M. C., Zizka, Georg, Paule, Juraj, Santoro, Julián Aguirre-, Heller, Sascha, Ramírez-Morillo, Ivón M., Halbritter, Heidemarie, Mariath, Jorge E. A., De, Jordano D. T. & Forzza, Rafaela C., 2021, Re-evaluation of the Amazonian Hylaeaicum (Bromeliaceae: Bromelioideae) based on neglected morphological traits and molecular evidence, pp. 1-60 in Phytotaxa 499 (1)</i> on page 48, DOI: 10.11646/phytotaxa.499.1.1, <a href="http://zenodo.org/record/5424411">http://zenodo.org/record/5424411</a&gt

    Hylaeaicum margaretae Leme & Forzza 2021, comb. nov.

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    <i>Hylaeaicum margaretae</i> (L.B. Sm.) Leme & Forzza, <i>comb. nov.</i> <p> Basionym:— <i>Neoregelia margaretae</i> L.B. Sm., in Mee, <i>Flowers of the Brazilian Forests Pl. 25</i>. 1968. Type:— BRAZIL. Amazonas: Rio Içana, tributary of upper Rio Negro, January 1965, <i>M. Mee s.n.</i> (holotype US!).</p>Published as part of <i>Leme, Elton M. C., Zizka, Georg, Paule, Juraj, Santoro, Julián Aguirre-, Heller, Sascha, Ramírez-Morillo, Ivón M., Halbritter, Heidemarie, Mariath, Jorge E. A., De, Jordano D. T. & Forzza, Rafaela C., 2021, Re-evaluation of the Amazonian Hylaeaicum (Bromeliaceae: Bromelioideae) based on neglected morphological traits and molecular evidence, pp. 1-60 in Phytotaxa 499 (1)</i> on page 48, DOI: 10.11646/phytotaxa.499.1.1, <a href="http://zenodo.org/record/5424411">http://zenodo.org/record/5424411</a&gt

    Hylaeaicum tarapotoense Leme, Zizka & Aguirre-Santoro 2021, comb. nov.

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    <i>Hylaeaicum tarapotoense</i> (Rauh) Leme, Zizka & Aguirre-Santoro, <i>comb. nov.</i> <p> Basionym:— <i>Neoregelia tarapotoensis</i> Rauh, <i>Trop. Subtrop.Pflanzenw.</i> 52: 68. 1985. Type:— PERU. San Martin:via Tarapoto-Yurimaguas, 1200 m elev., 17 August 1983, <i>W. Rauh</i> & <i>Bismarck 63976</i> (holotype HEID!).</p>Published as part of <i>Leme, Elton M. C., Zizka, Georg, Paule, Juraj, Santoro, Julián Aguirre-, Heller, Sascha, Ramírez-Morillo, Ivón M., Halbritter, Heidemarie, Mariath, Jorge E. A., De, Jordano D. T. & Forzza, Rafaela C., 2021, Re-evaluation of the Amazonian Hylaeaicum (Bromeliaceae: Bromelioideae) based on neglected morphological traits and molecular evidence, pp. 1-60 in Phytotaxa 499 (1)</i> on page 49, DOI: 10.11646/phytotaxa.499.1.1, <a href="http://zenodo.org/record/5424411">http://zenodo.org/record/5424411</a&gt

    Hylaeaicum peruvianum Leme, Zizka & Aguirre-Santoro 2021, comb. nov.

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    <i>Hylaeaicum peruvianum</i> (L.B. Sm.) Leme, Zizka & Aguirre-Santoro, <i>comb. nov.</i> <p> Basionym:— <i>Neoregelia peruviana</i> L.B. Sm., <i>Phytologia.</i> 9: 245, pl. 1, figs. 10-12. 1963. Type:— PERU. Loreto: Alto Amazonas, rainforest on lower northwest slopes of Cerros Campanquiz, Rio Marañón, just above Pongo de Manseriche, 250-350 m elevation, 17 October 1962, <i>J.J. Wurdack 2320</i> (holotype US!, isotype USM!).</p>Published as part of <i>Leme, Elton M. C., Zizka, Georg, Paule, Juraj, Santoro, Julián Aguirre-, Heller, Sascha, Ramírez-Morillo, Ivón M., Halbritter, Heidemarie, Mariath, Jorge E. A., De, Jordano D. T. & Forzza, Rafaela C., 2021, Re-evaluation of the Amazonian Hylaeaicum (Bromeliaceae: Bromelioideae) based on neglected morphological traits and molecular evidence, pp. 1-60 in Phytotaxa 499 (1)</i> on page 49, DOI: 10.11646/phytotaxa.499.1.1, <a href="http://zenodo.org/record/5424411">http://zenodo.org/record/5424411</a&gt

    Two New Species of Tillandsia (Bromeliaceae: Tillandsioideae) from Brazil

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    Souza, Everton Hilo De, Cavalcante, Brayan P., França, Rivã Ribeiro Do N., Aona, Lidyanne Y. S., Souza, Fernanda V. D., Leme, Elton M. C. (2022): Two New Species of Tillandsia (Bromeliaceae: Tillandsioideae) from Brazil. Phytotaxa 560 (1): 93-103, DOI: 10.11646/phytotaxa.560.1.7, URL: http://dx.doi.org/10.11646/phytotaxa.560.1.

    Hylaeaicum eleutheropetalum var. bicolor Leme & Forzza 2021, comb. nov.

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    <i>Hylaeaicum eleutheropetalum</i> var. <i>bicolor</i> (L.B. Sm.) Leme & Forzza, <i>comb. nov.</i> <p> Basionym:— <i>Neoregelia eleutheropetala</i> var. <i>bicolor</i> L.B. Sm., <i>Phytologia</i> 8: 227. 1962. Type:— PERU. Loreto: near Loreto, 1961, <i>L. Moore s.n.</i> (holotype US!, isotype NY!).</p>Published as part of <i>Leme, Elton M. C., Zizka, Georg, Paule, Juraj, Santoro, Julián Aguirre-, Heller, Sascha, Ramírez-Morillo, Ivón M., Halbritter, Heidemarie, Mariath, Jorge E. A., De, Jordano D. T. & Forzza, Rafaela C., 2021, Re-evaluation of the Amazonian Hylaeaicum (Bromeliaceae: Bromelioideae) based on neglected morphological traits and molecular evidence, pp. 1-60 in Phytotaxa 499 (1)</i> on page 48, DOI: 10.11646/phytotaxa.499.1.1, <a href="http://zenodo.org/record/5424411">http://zenodo.org/record/5424411</a&gt
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