192 research outputs found

    Protochorebus pervushini Kostromina, Timokhov & Belokobylskij, 2016, sp. nov.

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    Protochorebus pervushini sp. nov. (Figs 10–23) Type material. Holotype: female, Middle Urals, Ekaterinburg City outskirts, Verkh-Isetsk pond, Baran Island, sweeping on sedge, 25.V.2015, T. Kostromina coll. (ZISP). Paratypes. 1 female, 5 males, same label as holotype (ZISP, TK); 1 male, same locality, 14.V.2009 (ZISP); 1 female, 13 males, same locality, 19.V.2009 (ZISP, TK); 1 female, 4 males, same locality, 10–15.V.2011 (ZISP, TK); 1 male, same locality, reared from sedge tussock infested by Selachops flavocinctus (Agromyzidae), 7.V.2013 (ZISP); 2 females (one damaged), same locality, from sedge tussock infested by Selachops flavocinctus (Agromyzidae), collected 30.IV.2014, reared 12.V.2014 (ZISP); 1 male, same locality, from sedge tussock infested by Selachops flavocinctus (Agromyzidae), collected 1.V.2014, reared 11.V.2014 (ZISP); 1 male, same locality, “[9– 8] Reared from pupa of Selachops flavocinctus (Agromyzidae) 12.05.2015, sedge tussock collected 3.V.2015 ” (TK). Description. Female (Fig. 10). Body length 3.3–4.7 mm; fore wing length 3. 0–3.6 mm. Head (Fig. 12) behind eyes (dorsal view) evenly and more or less distinctly widened anteriorly, roundly narrowed posteriorly, 1.7–1.8 × as wide as median length, about 1.5 × as wide as maximum length, 1.40–1.45 × as wide as mesoscutum. Occiput medially distinctly concave. Frons with shallow and rather wide median longitudinal furrow in posterior half. Temple 1.1 × as long as transverse diameter of eye (1.3 × if measured in a straight line). Ocelli arranged in almost equilateral triangle. POL 1.5–1.6 × OD, 0.3–0.4 × OOL. Eyes glabrous, 1.2–1.3 × as high as broad. Face (Fig. 11) width equal to height of eye, 1.5–1.7 × median height of face. Width of clypeus 2.5 × its median height. Mandible (Fig. 14) weakly widened towards subapex, its median length 1.5–1.7 × maximum subapical width; with five teeth, formula of its teeth 1b+2b+3 (three main teeth and two, lower first and lower second, additional teeth). Upper tooth rather short, wide, weakly outstanding, subrectangular, apically with distinct median excavation dividing apex into two obtuse teeth. Median tooth long, rather narrow, almost straight, (sub)pointed apically, slightly direct outward, submedially with small obtuse additional ventral tooth. Excision between upper and median teeth not deep and narrow. Lower tooth short, rather wide, slightly direct outward, subpointed. Antenna (Figs 15, 16) rather thick but narrowed apically, weakly fusiform in basal one-third and here with transverse segments, with subsquare or weakly elongate segments in apical half, 42-segmented, about 0.9 × as long as body. Scape 1.6–1.8 × longer than its maximum width. First flagellar segment 2.0–2.3 × longer than its apical width, 1.3–1.4 × longer than second segment. Transverse flagellar segments situated in basal third (dorsal view) 1.3–1.4 × wider medially that their length. Penultimate segment 1.3–1.4 × longer than wide, 0.4 × as long as first flagellar segment, 0.7 × as long as apical segment. Last (apical) segment more or less pointed apically but without spine. Mesosoma (Figs 17, 18) 1.5–1.6 × longer than height. Pronotum with deep pronope. Mesonotum with distinct median longitudinal and weakly crenulate depression in posterior half, 0.85–0.90 × as long as maximum width. Notauli deep and crenulate in anterior 0.3–0.4, shallow submedially, absent in posterior half. Prescutellar depression deep, coarsely crenulate (with five to six carinae), 0.3–0.4 × as long as the convex scutellum. Metanotum with distinct median longitudinal carina (dorsal view) transformed posteriorly in short, wide and obtuse tooth (lateral view). Precoxal sulcus (sternaulus) (Fig. 18) wide, deep, sinuate, strongly rugose-crenulate, reaching anterior and posterior margins of mesopleuron. Subalar depression rather shallow, wide, coarsely striate. Metapleural flange wide, short and obtuse. Propodeal spiracles very small and subround. Wings (Fig. 21). Fore wing 2.5–2.7 × longer than wide. Pterostigma short and wide, 3.5–4.0 × longer than its maximum width. Radial (marginal) cell distinctly shortened, finishing far from apex of wing. Metacarp (1-R1) 0.85–0.90 × as long as pterostigma, 1.9–2.0 × longer than distance from apex of radial (marginal) cell to apex of wing. Radial (r) vein arising somewhat before or nearly middle of pterostigma; inner anterior margin of pterostigma 0.9 × as long as its inner posterior margin. Second radial abscissa (SR1+3-SR) evenly curved, almost straight in posterior third. First radial abscissa (r) 0.2 × as long as second abscissa (SR1+3-SR) (if measured on straight line). Recurrent (m-cu) vein strongly antefurcal. First abscissa of longitudinal anal vein (1-1A) weakly evenly curved submedially. Distance between basal vein (1-M) and nervulus (cu-a) 0.6 × nervulus (cu-a) length. Brachial (first subdiscal) cell weakly widened to apex, widely open apically. Parallel vein (CU1a) arising behind middle of apical margin of brachial (first subdiscal) cell. Hind wing 3.8–4.0 × longer than its maximum width. First costal abscissa (C+SC+R) 0.8–0.9 × as long as second abscissa (1-SC+R). First abscissa of mediocubital vein (M+CU) 3.0–3.7 × longer than second abscissa (1-M). Legs. Hind femur (Fig. 19) 3.8–4.4 × longer than its maximum width. Hind tarsus 0.85–0.90 × as long as hind tibia. Hind basitarsus 0.55–0.60 × as long as second-fifth segments combined. Second segment of hind tarsus 0.6 × as long as hind basitarsus, 1.3–1.4 × longer than hind fifth segment (without pretarsus). Metasoma (Figs 22, 23) more or less elongated, weakly depressed, 2.6–2.8 × longer than its maximum width, 0.9–1.0 × as long as head and mesosoma combined. First tergite distinctly widened from base to spiracles, then almost parallel-sided, with wide dorsope, with distinct dorsal carinae fused basally and medially following till apex of tergite as single carina. Length of first tergite 1.3–1.5 × its apical width; apical width 1.6–1.9 × its basal width. Second tergite medially 0.5 × as long as basal width, 0.8–0.9 × as long as third tergite. Median length of second and third tergites 1.0–1.4 × its basal width, 0.7–1.0 × its apical width. Second suture very weak. Second-sixth tergites without separated laterotergites. Ovipositor sheath (Fig. 20) very short, usually not or sometimes weakly protruding behind tip of metasoma, about as long as third segment of hind tarsus. Sculpture and pubescence. Head mainly smooth, face finely punctate, clypeus almost smooth. Mesoscutum mainly smooth, finely rugulose-punctate anteriorly, sometimes partly with very fine granulation. Scutellum entirely smooth. Mesopleuron mainly smooth. Propodeum coarsely and densely rugose-reticulate with additional dense and fine granulation, with high median longitudinal carina. First tergite entirely densely rugose-reticulate with additional fine granulation. Remaining tergites smooth. Vertex entirely and mesoscutum almost entirely covered by dense pale short setae. Propodeum, metapleuron and first metasomal tergites almost entirely covered by long and rather dense pale setae. Hind coxa dorsally in long and rather dense pale setae not forming bunch. Third to sixth tergites with two-three lines of transverse pale setae in their posterior thirds. Sheath of ovipositor with cluster оf dense setae apically. Colour. Body mainly black, metasoma partly with reddish tint, its sternites reddish brown on wide areas. Mandible reddish brown to dark reddish brown, almost black basally. Palpi reddish yellow, darker basally. Antenna mainly black, two basal segments reddish brown to dark reddish brown. Tegula almost black. Fore leg light reddish brown, fore coxa and trochanter brown to black; middle coxa, trochanter, trochantellus and basal half of femur dark brown to black, apical half of femur, tibia and tarsus (except dark apical segment) light reddish yellow with infuscation; hind coxa, trochanter and femur black, trochantellus and tibia mainly light reddish brown or reddish brown, apex of hind tibia and tarsus dark reddish brown. Fore wing faintly infuscate; veins brown to dark brown, but basally mainly brownish yellow or yellow. Pterostigma entirely dark brown. Male. Body length 3.7–5.1 mm; fore wing length 3.4–4.1 mm. Temple (Fig. 13) 1.2–1.3 × longer than transverse diameter of eye (1.4–1.7 × if measured on straight line). Antennae thickened, weakly setiform, 45–52- segmented, 1.0–1.1 × as long as body. Submedian antennal segments about as long as their maximum width. Mesoscutum sometimes with longitudinal striation in submedian area. Pterostigma 4.2–5.3 × longer than its maximum width. First radial abscissa 0.8–1.0 × as long as first radiomedial vein. Hind femur 4.0–4.3 × longer than its maximum width. Metasoma narrow, sometimes second and third tergites brownish. First tergite often weakly narrowed from spiracular tubercles towards apex or subparallel, its apical width 0.85–1.2 × width at level of spiracles, 1.4–1.8 × minimum width; length 1.5–1.6 × its apical width. Otherwise similar to female. Etymology. This species is named in honour of Artem A. Pervushin (Yekaterinburg), who prematurely passed away, and who helped very much during Braconidae study of the first author. Host. Selachops flavocinctus Wahlberg, 1844 (Diptera: Agromyzidae). Distribution. Russia (Middle Urals). Comparative diagnosis. The differences between new species P. pervushini sp. nov. and type species of this genus, P. kasparyani Perepechaenko (Figs 24–34), are shown in the key below: 1. Upper (first) tooth of mandible compound, with additional distinct tubercle below (Fig. 14). Mesoscutum of female partly finely punctate, its median posterior furrow rather long (Fig. 17). Dorsal tooth of metanotum (lateral view) short (Fig. 18). Recurrent vein (m-cu) of fore wing less strongly antefurcal, about 2.0 × longer than second abscissa of medial vein (2-SR+M) (Fig. 21). Second radial abscissa (SR1+3-SR) of fore wing not sinuate in apical half (Fig. 21). Hind femur black (Fig. 19). Body length 3.3–5.1 mm.................................................... P. pervushini sp. nov. (Figs 10–23) Upper (first) tooth of mandible simple, without additional tubercles (Fig. 24). Mesoscutum of female mainly distinctly punctate, its median posterior furrow rather short (Fig. 32). Dorsal tooth of metanotum (lateral view) long (Fig. 31). Recurrent vein (m-cu) of fore wing strongly antefurcal, 1.3–1.5 × longer than second abscissa of medial vein (2-SR+M) (Fig. 30). Second radial abscissa (SR1+3-SR) of fore wing more or less distinctly sinuate in apical half (Fig. 30). Hind femur light reddish brown (Fig. 25). Body length 4.2–4.4 mm................................. P. kasparyani Perepechaenko, 1997 (Figs 24–34) Based on the shape and structure of mandibles, P. pervushini sp. nov. resembles Chorebus (Pentalexis) mysteriosus Perepechaenko, 2004, described from two males from Zabaykal'skiy Territory of Russia and separated in the new subgenus mainly on the basis of mandible structure (Perepechaenko, 2004). The differences between these species are shown in the following key: 1. Eyes less strongly convergent below and without setae (Fig. 11). Face distinctly transverse (Fig. 11). Antennal segments in basal third distinctly widened, transverse (Fig. 15). Mesosoma 1.6 × longer than high (Fig. 18). Dorsal tooth of metanotum (lateral view) short (Fig. 18). Metapleuron without oval sculptured area. Hind coxa dorsally without tuft of setae. Radial vein (r) of fore wing arising almost from middle of pterostigma (Fig. 21)............................ P. pervushini sp. nov. - Eyes strongly convergent below, with short and sparse setae. Face subsquare. Antennal segments in basal third distinctly narrow, elongate. Mesosoma about twice longer than high. Dorsal tooth of metanotum (lateral view) long. Metapleuron with large oval sculptured area. Hind coxa dorsally with more or less distinct tuft of setae. Radial vein (r) of fore wing arising distinctly before middle of pterostigma............................... Chorebus (Pentalexis) mysteriosus Perepechaenko, 2004Published as part of Kostromina, Tatiana S., Timokhov, Alexander V. & Belokobylskij, Sergey A., 2016, Braconid wasps of subfamily Alysiinae (Hymenoptera: Braconidae) as endoparasitoids of Selachops flavocinctus Wahlberg, 1844 (Diptera: Agromyzidae) in the Central Urals, Russia, pp. 305-319 in Zootaxa 4200 (2) on pages 309-314, DOI: 10.11646/zootaxa.4200.2.3, http://zenodo.org/record/18217

    Hydrochemical Atlas of the Arctic Ocean

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    Introduction: Chemical composition of water determines its physical properties and character of processes proceeding in it: freezing temperature, volume of evaporation, density, color, transparency, filtration capacity, etc. Presence of chemical elements in water solution confers waters special physical properties exerting significant influence on their circulation, creates necessary conditions for development and inhabitance of flora and fauna, and imparts to the ocean waters some chemical features that radically differ them from the land waters (Alekin & Liakhin, 1984). Hydrochemical information helps to determine elements of water circulation, convection depth, makes it easier to distinguish water masses and gives additional knowledge of climatic variability of ocean conditions. Hydrochemical information is a necessary part of biological research. Water chemical composition can be the governing characteristics determining possibility and limits of use of marine objects, both stationary and moving in sea water. Subject of investigation of hydrochemistry is study of dynamics of chemical composition, i.e. processes of its formation and hydrochemical conditions of water bodies (Alekin & Liakhin 1984). The hydrochemical processes in the Arctic Ocean are the least known. Some information on these processes can be obtained in odd publications. A generalizing study of hydrochemical conditions in the Arctic Ocean based on expeditions conducted in the years 1948-1975 has been carried out by Rusanov et al. (1979). The “Atlas of the World Ocean: the Arctic Ocean” contains a special section “Hydrochemistry” (Gorshkov, 1980). Typical vertical profiles, transects and maps for different depths – 0, 100, 300, 500, 1000, 2000, 3000 m are given in this section for the following parameters: dissolved oxygen, phosphate, silicate, pH and alkaline-chlorine coefficient. The maps were constructed using the data of expeditions conducted in the years 1948-1975. The illustrations reflect main features of distribution of the hydrochemical elements for multi-year period and represent a static image of hydrochemical conditions. Distribution of the hydrochemical elements on the ocean surface is given for two seasons – winter and summer, for the other depths are given mean annual fields. Aim of the present Atlas is description of hydrochemical conditions in the Arctic Ocean on the basis of a greater body of hydrochemical information for the years 1948-2000 and using the up-to-date methods of analysis and electronic forms of presentation of hydrochemical information. The most wide-spread characteristics determined in water samples were used as hydrochemical indices. They are: dissolved oxygen, phosphate, silicate, pH, total alkalinity, nitrite and nitrate. An important characteristics of water salt composition – “salinity” has been considered in the Oceanographic Atlas of the Arctic Ocean (1997, 1998). Presentation of the hydrochemical characteristics in this Hydrochemical Atlas is wider if compared with that of the former Atlas (Gorshkov, 1980). Maps of climatic distribution of the hydrochemical elements were constructed for all the standard depths, and seasonal variability of the hydrochemical parameters is given not only for the surface, but also for the underlying standard depths up to 400 m and including. Statistical characteristics of the hydrochemical elements are given for the first time. Detailed accuracy estimates of initial data and map construction are also given in the Atlas. Calculated values of mean-root deviations, maximum and minimum values of the parameters demonstrate limits of their variability for the analyzed period of observations. Therefore, not only investigations of chemical statics are summarized in the Atlas, but also some elements of chemical dynamics are demonstrated. Digital arrays of the hydrochemical elements obtained in nodes of a regular grid are the new form of characteristics presentation in the Atlas. It should be mentioned that the same grid and the same boxes were used in the Atlas, as those that had been used by creation of the US-Russian climatic Oceanographic Atlas. It allows to combine hydrochemical and oceanographic information of these Atlases. The first block of the digital arrays contains climatic characteristics calculated using direct observational data. These climatic characteristics were not calculated in the regions without observations, and the information arrays for these regions have gaps. The other block of climatic information in a gridded form was obtained with the help of objective analysis of observational data. Procedure of the objective analysis allowed us to obtain climatic estimates of the hydrochemical characteristics for the whole water area of the Arctic Ocean including the regions not covered by observations. Data of the objective analysis can be widely used, in particular, in hydrobiological investigations and in modeling of hydrochemical conditions of the Arctic Ocean. Array of initial measurements is a separate block. It includes all the available materials of hydrochemical observations in the form, as they were presented in different sources. While keeping in mind that this array contains some amount of perverted information, the authors of the Atlas assumed it necessary to store this information in its primary form. Methods of data quality control can be developed in future in the process of hydrochemical information accumulation. It can be supposed that attitude can vary in future to the data that were rejected according to the procedure accepted in the Atlas. The hydrochemical Atlas of the Arctic Ocean is the first specialized and electronic generalization of hydrochemical observations in the Arctic Ocean and finishes the program of joint efforts of Russian and US specialists in preparation of a number of atlases for the Arctic. The published Oceanographic Atlas (1997, 1998), Atlas of Arctic Meteorology and Climate (2000), Ice Atlas of the Arctic Ocean prepared for publication and Hydrochemical Atlas of the Arctic Ocean represent a united series of fundamental generalizations of empirical knowledge of Arctic Ocean nature at climatic level. The Hydrochemical Atlas of the Arctic Ocean was elaborated in the result of joint efforts of the SRC of the RF AARI and IARC. Dr. Ye. Nikiforov was scientific supervisor of the Atlas, Dr. R. Colony was manager on behalf of the USA and Dr. L. Timokhov – on behalf of Russia

    Russian-German Cooperation: Laptev Sea System : [2. Workshop Russian-German Cooperation: Laptev Sea System ; St. Petersburg, November 1994]

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    PREFACE : The Laptev Sea System The Arctic Ocean, in particular the wide Eurasian shelf seas comprise some of the most sensitive elements of the global environment which are believed to respond at a very early time to Global Change. The renewed interest in the Arctic, the large scale international research efforts devoted to the Arctic, as well as the presently available new technology to carry out research in ice-infested areas, have opened many new avenues to conduct investigations On the variability of the depositional environments of the Eurasian shelf seas. The Laptev Sea is of particular importance in the string of the Eurasian shelf seas because feeding the Transpolar Drift of the Arctic sea-ice Cover it exports relatively the largest amounts of sea ice into the Open Arctic Ocean, because it is farthest away from the influence of the Atlantic and Pacific waters, and because it is under the influence of rapidly changing fresh water fluxes from the Siberian hinterland (Fig. 1, Sea ice drift paths in the Arctic Ocean). The morphology of the seafloor, the rapidly changing coast lines of the fragil Lena Delta Island frame work as well as the presence of submarine permafrost are examples for the dynamics of the entire Laptev Sea System. - Fig. 1 - In order to address the natural properties of the Laptev Sea System a joint research project is carried out between a number of Russian and German research institutions under the framework of the "Laptev Sea System Project" (Fig. 2, Research institutions under the framework of the "Laptev Sea System Project"). Every year expeditions are carried out in the area on Russian or German research vessels where multi-disciplinary and binational working groups are addressing some of the identified scientific themes. Results from these joint investigations are then discussed in a series of RussianIGerman workshops which are held alternatively in Russia or Germany. The second workshop 'Russian-German Cooperation: Laptev Sea System' was held in November 1994 in St. Petersburg in order to assess (1) the state of knowledge of the Laptev Sea and the adjacent continental margin of the deep Arctic, and (2) to develop a research strategy for the marine geosciences in the Laptev Sea and terrestrial werk in East Siberia. The workshop brought together more than 100 scientists, among them meteorologists, sea ice physicists, oceanographers, biologists, chemists, geologists and geophysicists from various Russian and German research institutions. The main goal of the workshop was to promote and coordinate scientific collaboration among scientists from Russia and Germany. Main emphasis have laid on first scientific results of the expeditions within the scope of the interdisciplinary Russian-German research project 'Laptev Sea System', that is present and past oceanography, ecology, and climatology of the Laptev Sea. The workshop was organized into serveral sessions which followed various themes of the environment of the Laptev Sea from their present situation to their geological record: (I) Ciimate and Ice (11) Modern Environment of the Laptev Sea (111) Environmental History of the Laptev Sea (IV) From Siberia to the Arctic Ocean: Land-Sea Connection (V) Strategy and Plans for Future Work (VI) Mid-long Term Perspectives The scientific content of this workshop is documented in this report containing most of the results and discussions. The publication of this volume serves various purposes. It is primarily a forum for scientists working in the Siberian shelf seas, in which the results of many years of research and preliminary shipboard results can be presented. In order to provide all the participants in the workshop with the opportunity for reporting their results, a speedy way of publication was chosen. Thus, each individual author has presented his opinions and views as he or she sees them, reflecting the diversity and complexity of the Laptev Sea system. On the other hand, this volume offers many researchers the possibility of acquainting themselves with methods and results of research into the East Siberian seas as carried out in other parts of the world. Finally, it is hoped that this collection of papers will function as another step toward joint research projects and are base for the expeditions to be carried out in 1995 and the following years. Many of the papers published identify major scientific problems, thus offering new perspectives for future scientific research in polar regions. The nature of the papers, the discussions and the disciplines of the attendees clearly demonstrate that the study of the Laptev Sea System is a multidisciplinary one in an interesting key area involving all branches of the natural sciences, such as ice physics, oceanography, biology and geology, in particular. It thus remains an important example for GLOBAL CHANGE and CLIMATE IMPACT research within international research efforts, e.g. International Arctic Science Committee (IASC), Arctic Ocean Sciences Board (AOSB) or the Nansen Arctic Drilling Programme (NAD). - Fig. 2 - The editors also made an effort, probably not wholly successful, to edit manuscripts by non-English-speaking authors to make them easier to understand. In this process, we hope we have not changed the meanings of the original papers. Above all we thank Bettina Rohr and Daniel Krüger who kindly assisted in editing the papers. The workshop has been sponsored by the German and Russian Ministries for Research and Technology and the meeting was held from the 21st to the 14th of November in 1994 in the Arctic and Antarctic Research Institute in St. Petersburg. We wish to thank these organizations for their financial and logistic support
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