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Deltochilum acanthus Kohlmann & Solis, new species
No full textDeltochilum acanthus Kohlmann & Solís, new species Figs. 6, 7, 15 Diagnosis. This species is distinguished from other Costa Rican Deltochilum species by the following combination of characters: Pygidium with apex thickened, very acutely angled (90 °) and outwardly produced (Fig. 7 f). Apical half of metatibia bent inward. Description. Holotype. Male (Fig. 6): Length 11.9 mm. Humeral width 8.0 mm. Body color black, head and pronotum distinctly punctate. Head approximately as long as wide; clypeus with two distinct, narrowly separated, upwardly reflexed, pointed teeth, pointed anterior teeth; each clypeal-genal margin with low obtuse teeth; vertex closely punctate, nearly flat; dorsal eyes large. Pronotum with lateral margin angulate at middle, otherwise straight to slightly sinuate before and behind; pronotal surface densely punctate, giving the appearance of a honeycomb. Elytral striae indicated by a double line, line expanded by widely separated, shallow strial punctures; disc intervals opaque, with numerous shallow, very close punctures (Fig. 7 h); humeral umbone and apices of third to seventh intervals carinate; elytral surface shagreened. Epipleural upper carina interrupted (Figs. 7 b, d). Pygidium with apex thickened, very acutely angled and outwardly produced (Fig. 7 f); disc nearly flat, shallowly and densely punctate, punctures umbilical. First abdominal sternite forms a rearward projection at the middle of the posterior border. Profemur with margin unmodified; protibia with three apical teeth (the middle one smaller) and numerous serrations on outer margin. Apical half of metatibia bent inwardly. Allotype. Female. Length 11.2 mm. Humeral width 7.6 mm. Similar to male, but the first abdominal sternite is evenly arched at the middle of the posterior border. Variation. Fifty-eight specimens examined, 38 males and 20 females. Length 10.4–13.0 mm. Humeral width 7.3–8.0 mm. Examined material (58 specimens). Holotype, male: COSTA RICA. Prov. Puntarenas. Res. Biol. Carara, Est., Quebrada Bonita. 50 m, jun 1993. J.C. Saborío. L-N- 194500, 469850, CRI 001185073. Allotype, female: COSTA RICA. Prov. Puntarenas. R.B. Carara, Est. Quebrada Bonita. 50 m, nov. 1993. J.C. Saborío. L S 194500 _ 469850, # 2470, CRI 001969700. Paratypes. COSTA RICA. Prov. Puntarenas. Centro Juvenil Tropical. Alrededor de la Estación. 100m. 5–12 JUL 1997. M. Lobo. Colecta Nocturna. L_S_ 294700 _ 517100 # 47733, 1 female; Estación Esquinas, 0 m, Península de Osa, Abr 1993. M. Segura, L-S 301400 _ 542200, 1 female; Estación Quebrada Bonita, 50 m, Res. Biol. Carara, 1 a 29 jul 1992, R. Guzmán, L- N 194500 _ 469850, 1 male; Oct 1993. J. C. Saborío, L N 194500 _ 469850 # 2396, 1 female; Set 1993. J. Saborío, L N 194500 _ 469850 # 2354, 2 males; Nov 1993, R. Guzmán, L N 194500 _ 469850 # 2447, 2 males; Jun 1993, R. Guzmán, L N 194500 _ 469850 # 2202, 1 male; 2 a 23 set 1992, R. Guzmán, L-N 194500 _ 469850, 3 males; agos 1993, R.M. Guzmán, L N 194500 _ 469850 # 2297, 1 female; Ene 1994, R. M. Guzmán, L N 194500 _ 469850 # 2572, 1 female; Feb 1994, R. M. Guzmán, L N 194500 _ 469850 # 2613, 1 male; May 1994, R. M. Guzmán, L N 194500 _ 469850 # 2914, 3 males, 1 female; Ago 1994, R. M. Guzmán, L N 194500 _ 469850 # 3163, 1 female; Oct 1994. J.C. Saborío, Desconocido L_N_ 469850 _ 194500 # 3288, 1 female; Jun 1996. R. Guzmán. L_N_ 195250 _ 469850 # 7648, 1 male; Set 1994. R. M. Guzmán, L_N_ 194500,469850 # 3214, 2 males, 1 female; May 1994. J. Saborío. L_N_ 470000 _ 195200 # 2849, 4 males, 1 female; Estación Sirena, 0–100m, P. N. Corcovado, Ago 1991, J. C. Saborío, L S 270500 _ 508300, 1 female; G. Fonseca, Jun 1991, L- S 270500 _ 508300, 1 male; Jun 1991, J. C. Saborío, L- S 270500 _ 508300, 1 male; Oct 1993. G. Fonseca, L S 270500 _ 508300 # 2380, 1 male; Jan 1990, G. Fonseca, L_S_ 270500 _ 508300, 1 female; G. Fonseca, Abr 1991, L- S 270500 _ 508300, 1 male, 1 female; G. Fonseca, Oct 1989, L- S 270500 _ 508300, 1 female; F. Quesada, Jun 1990, L- S 270500 _ 508300, 1 male; Refugio de Vida Silvestre Golfito, Estación Naranjales, 0 – 100m, 25 ABR 2004, W. Porras, Libre, L_S_ 289900 _ 553450 # 76842, 1 male; 22 – 27 ABR 2004, W. Porras, B. Gamboa, D. Briceño, M. Moraga, Amarilla, L_S_ 289900 _ 553450 # 76946, 2 males, 3 females; Rancho Quemado, 200 m, Península de Osa. 12 a 24 may 1993. A. Gutiérrez, L S 292500 _ 511000, 1 male; Oct 1990. F. Quesada. L-S 292500, 511000, 1 male; 11–28 Oct 1993, A. H. Gutiérrez, L S 292500 _ 511000 # 2409, 2 males, 1 female; Río Agujas. Estación Agujas. Send. Ajo. 300m. 6–12 ENE 1998. M. Lobo. L_S_ 276750 _ 526550 # 49736, 1 male. PANAMA. Canal Zone, Barro Colorado, Poscher’s Peninsula, 6 jun 1986. H. Wolda, 1 male; 11 jun 1986. H. Wolda, 1 female; 18 jun 1986. H. Wolda, 1 male; 25 jun 1986, H. Wolda, 1 male; 11 sep 1987. H. Wolda, 1 male. Habitat. The species has been collected with flight interception traps in tropical rain forest, ranging from 0– 100 m altitude, during the months of April to November. Geographical distribution (Fig. 15). This species is known so far from the Pacific rain forest of Costa Rica and the Canal Zone of Panama. Chorological affinities. Deltochilum acanthus seems to show a geographic vicariant pattern in relation to the similar species D. valgum acropyge, which inhabits the Caribbean slope of Costa Rica. Taxonomic relationships. It would appear that the new species originated from a vicariant event, when the Talamanca range rose up approximately 3 million years ago, isolating the rain forest on the Pacific coast from the rainforests on the Caribbean coast. This mechanism seems to account for the origin of a great number of other animal vicariant species, examined in Kohlmann & Wilkinson (2007). We believe that D. valgum needs to be studied, and that its different subspecies represent a species complex in need of hierarchical revaluation. We therefore describe D. acanthus as a species and not as a subspecies, in anticipation of this process. Deltochilum acanthus can be easily separated from D. valgum by its pygidium, which has the thickened apex, very acutely angled and outwardly produced (Fig. 7 f), whereas D. valgum has a much less thickened and projected pygidium (Fig. 7 e). There are also differences in dorsal punctation: the pronotum in D. acanthus is very densely punctured, producing the effect of a honeycomb, whereas in D. valgum the punctures are more spaced, by at least the length of one puncture. The elytral punctures are also different, in D. acanthus punctation is dense and the elytral striae are broad (Fig. 7 h), whereas D. valgum is less densely punctured and the striae are thin (Fig. 7 g). Etymology. Acanthus (ĸανθοζ = acanthos), a Latinized Greek noun in apposition, meaning thorn, making reference to the spiny pygidial apex.Published as part of Kohlmann, Bert & Solís, Ángel, 2012, New species and revalidations of scarab beetles (Coleoptera: Geotrupidae: Athyreini and Coleoptera: Scarabaeidae: Scarabaeinae) from Costa Rica and Panama, pp. 28-52 in Zootaxa 3193 on pages 36-38, DOI: 10.5281/zenodo.21112
Canthidium margaritae Kohlmann & Solis, sp. nov.
No full text<i>Canthidium margaritae</i> Kohlmann & Solís sp. nov. (Figs. 1–3) <p> <b>Diagnosis.</b> This species is distinguished by the following combination of characters: body globose; head and pronotum uniformly and strongly punctate; frons swollen; frontoclypeal region with a small, smooth swelling medial to each eye; eye dorsally at posterior end of gena narrow, eight to ten facets wide, interocular distance separated by approximately 8–10 times maximal eye width; basal pronotal border not margined; elytra with eight striae, surface lightly shagreened, microsculpture coarser towards apex; hind wing brachypterous.</p> <p> <b>Description of Holotype.</b> Male. Length: 4.5 mm, humeral width: 2.9 mm. Body form globose (Fig. 1). Head and pronotum coppery­red, elytra dark brown (Fig.1). Head and pronotum strongly punctate and devoid of setae.</p> <p>Clypeus rugosely punctate, apex strongly bidentate, median emargination broadly Vshaped (Fig. 2 a); eye dorsally at posterior end of gena narrow, eight to ten facets wide, eyes separated by approximately 8–10 times maximum eye width (Fig. 2 a). Frons and vertex of head strongly punctate, frons swollen; two small, black, smooth swellings between eyes (Fig. 2 a).</p> <p>Pronotum on disc and posterior angles slightly shagreened; surface strongly and uniformly punctate; lateral fovea oval; posterior margin lacking elongated punctures or groove.</p> <p>Elytral striae consisting of clear indented lines interrupted by fairly evenly spaced oval punctures (approximately the maximum length of one puncture); intervals shiny, finely punctate; surface shiny and slightly shagreened especially towards apex. Elytra globose; hindwing brachypterous (Fig. 2 h).</p> <p>Pygidium distinctly punctate, more coarsely so in basal half, surface shiny and slightly shagreened at base. Genitalia as in Fig. 2 g.</p> <p>Protibia with three teeth on external border (Fig. 2c), inner apical margin of protibia produced into a rounded anterior projection and slightly bent downwards (Fig. 2c), apical spur with incurved apex (Fig. 2c). Pro­, meso­ and metafemora with ventral surface finely punctate and finely shagreened.</p> <p> <b>Allotype.</b> Female. Length: 4.6 mm, humeral width: 3.1 mm. Differing from male in the following major characters: clypeus slightly more transversely rugose (Fig. 2 b), apical spur not forming incurved apex and claw­bearing protarsus not as thick (Fig. 2 d), pygidium broader and less heavily punctate (Fig. 2 f), last abdominal segment broader, inner apex of protibia not forming a rounded lobe (Fig. 2 d).</p> <p> <b>Variation.</b> Elytral surface can range from completely shagreened to shagreened only on the apical third.</p> <p> <b>Material examined</b> (8 specimens). <b>Holotype</b> male: MÉXICO. Estado de México. Sierra de Nanchititla, Palos Prietos, 28–29­julio­2005, Alt. 1750 m, coprotrampa, M. Castillo, A. y L. Delgado cols. <b>Allotype</b> female: <i>ibidem</i>. Paratypes: MÉXICO. Estado de México. Sierra de Nanchititla, 18–19­VIII­96, 1800 msnm, G. Nogueira col., (1 female); same data as holotype (4 males, 1 female).</p> <p> <b>Habitat.</b> This species has been found in cloud forest mixed with oaks in elevations ranging from 1,750 to 1,800 m above sea level.</p> <p> <b>Geographical distribution.</b> It is only known in the State of México in the Sierra de Nanchititla on the upper reaches of the Balsas river depression (Fig. 3).</p> <p> <b>Chorological affinities.</b> The known range of <i>Canthidium margaritae</i> is widely separated from that of a very similar species, <i>C. riverai</i>, which is distributed in the Manantlán (Jalisco State) and the Coacolmán (Michoacán state) mountain ranges at similar altitudes (960–2,000 m above sea level) and in cloud forests.</p> <p> <b>Taxonomic relationships</b>. <i>Canthidium margaritae</i> is postulated to be the sister species of <i>C. riverai</i>. They are both globose in body shape and brachypterous, they also have a swollen frons, clearly punctate pronotum, shagreened elytra, evenly impressed striae, and they both inhabit cloud forests. This species pair apparently represents a vicariant speciation event between the Sierra Madre del Sur (<i>C</i>. <i>riverai</i>) and the Neovolcanic Axis (<i>C</i>. <i>margaritae</i>).</p> <p> <i>C. margaritae</i> will key to <i>C. riverai</i> Kohlmann and Solís in couplet 12 in the Kohlmann and Solís (2006) key for the <i>Canthidium</i> of Mexico. The two species are very similiar, but <i>C. margaritae</i> can be easily separated from <i>C. riverai</i> by having bigger eyes (8–10 eye facets versus 2–3 eye facets), head and pronotal punctures less coarse, elytra less shagreened, a different metatibial form (internal apical angle projected like a tapering rectangle versus apically obliquely truncated), and differences in the form of the parameres (parameres taper evenly towards apex versus parameres with a small hump at apical two­thirds).</p> <p> <b>Etymology.</b> We dedicate this species to Margarita Castillo, who has always supported Luis Leonardo Delgado in his entomological studies and also helped collect this new species. The name is derived from the latinized (<i>margarita</i>) Greek word μαργαρίτη, meaning a pearl.</p>Published as part of <i>Kohlmann, Bert & Solís, Ángel, 2006, New species of dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae) from Mexico and Costa Rica, pp. 61-68 in Zootaxa 1302</i> on pages 62-65, DOI: <a href="http://zenodo.org/record/173658">10.5281/zenodo.173658</a>
Onthophagus xiphias Solis & Kohlmann, sp. nov.
No full text<i>Onthophagus xiphias</i> Solís & Kohlmann, sp. nov. (Figs. 3­4) <p> <i>Onthophagus quetzalis</i> Howden & Gill, 1993 (<i>in part)</i>: 1101.</p> <p> <b>Diagnosis.</b> This species is distinguished by the following combination of characters: pronotal surface without setae; body colour uniformly reddish­brown; meso­and metafemora yellow in their central part; profemur brown; elytral surface with setae; functional wings; clypeal horn long with a thin base and forked distally; pronotal horn long, grooved dorsally and ventrally, ending in a thin rounded tip.</p> <p> <b>Description. Holotype. Male:</b> Length 6.1 mm. Humeral width: 3.7 mm. Body coloration is reddish­brown, more so in the pronotum, the meso­ and metafemora are yellowish.</p> <p>The head presents a clypeal horn, slender at its base but broadening and forking toward the apex. The frontal area of the head is smooth, with punctures toward the base; eyes small.</p> <p>The pronotum has a long and slender horn grooved dorsally and ventrally at its base and ends in a rounded tip. Pronotal surface slightly shagreened and covered with umbilicate punctures.</p> <p>Elytra with eight distinct striae, each with umbilicate punctation, intervals broad and slightly convex, with regular and coarse punctation. Elytral apices and epipleural stria with short and erect setae.</p> <p>Pygidium slightly shagreened, with umbilical punctures bearing short, stiff and erect setae.</p> <p>Protibiae long and slender, with tarsi present and four external teeth. Meso­ and metatibiae dilated toward the apex. First metatarsus long and almost rectangular in shape.</p> <p> <b>Allotype. Female:</b> Length 5.8 mm. Humeral width: 3.6 mm. Similar to male but unarmed, with a transversal carina at the middle of the head and the anterior border of the clypeus shows two well developed teeth separated by a “v” notch. Protibiae broader and shorter than the male.</p> <p> <b>Variation.</b> Length 5.2­6.6 mm. Humeral width: 2.9­3.8 mm. Sometimes the meso­ and metafemora are not yellow, but brown in their central parts.</p> <p> <b>Examined material</b> (101 specimens). <b>Holotype</b>, male: PANAMA. Chiriquí, Prov. Hornito, Finca la Suiza, 1220 m, 31.V.2000. H. & A. Howden, FIT. <b>Allotype</b>, female: <i>ibid</i>, 2.VI.2000. <b>Paratypes.</b> PANAMA. Chiriquí, Hornito, Finca La Suiza, 1220 m, H. & A. Howden, FIT, 1 specimen, 28.V.2000, 3 specimens, 29.V.2000, 7 specimens, 8 specimens, 30.V.2000, 31.V.2000, 10 specimens, 1.VI.2000, 4 specimens, 2.VI.2000, 9 specimens, 3.VI.2000, 7 specimens, 4.VI.2000, 11 specimens, 5.VI.2000, 8 specimens, 6.VI.2000, 18 specimens; Finca La Suiza, 20 km N Gualaca, 1350 m, 10­13.VI.1995, J. Ashe & R. Brooks, FIT, 1 specimen, 22­24.V.1995, J.S. & A.K. Ashe, 1 specimen, 1350 m, 10­ 13.VI.1995. J. Ashe & R. Brooks, ex: f.i.t.(196). 1 specimen; 2 km NW Hornitos, N 8°39.5’ W 82°12.3’, 24.V­6.VI.2000, 1250 m, B. & J. Gill, 7 specimens; Finca La Suiza; 5.3 km N Los Planes, N 8º39’, W 82º12’, V­26­30­1995, elev. 4500’. B. Ratcliffe & M. Jameson, 5 specimens, N 8º39’, W 82º12’, VII­10­13­1994, elev. 4500’. B. & I. Ratcliffe, M. Jameson, 1 specimen; Hartmann Finca, 15 km NW H[ato el] Volcan, 1500 m, S.Peck, 25.V.1975, dung trap, 1 specimen.</p> <p> <b>Habitat.</b> The species was collected with flight interception and dung traps inside cloud forest at altitudes varying from 1220 m to 1500 m.</p> <p> <b>Geographical distribution.</b> The species is known so far from the Pacific slope of the Chiriquí Volcano and probably occurs in the Talamanca Range in Costa Rica.</p> <p> <b>Chorological affinities.</b> <i>Onthophagus xiphias</i>, is widely separated from the known range of its closest relative, <i>O. quetzalis</i>, which is distributed in the Central, Tilarán and Guanacaste Cordilleras at similar altitudes (1000­1750 m) in relation to this new species. The other related species, <i>O. inediapterus</i>, is known from the Talamanca Range, but has been collected at higher altitudes (1650­1800 m) than this new species.</p> <p> <b>Taxonomic relationships.</b> <i>Onthophagus xiphias</i> is postulated to be the sister species of <i>O. quetzalis.</i> This species­pair represents another geographic and phylogenetic dichotomy between the Talamanca Range and the Central, Tilarán and Guanacaste Ranges as has been already noted in Kohlmann and Solís (2001).</p> <p> <i>O. xiphias</i> will key to <i>O. quetzalis</i> in Kohlmann and Solís (2001). It is easily distinguished by its much longer clypeal horn, thinner at its base and with a strongly forked distal end, whereas <i>O. quetzalis</i> has a short clypeal horn, thicker at its base and weakly forked distally. Moreover, <i>O. xiphias</i> has a very long pronotal horn, grooved dorsally and ventrally and finishing in a thin rounded tip. On the contrary <i>O. quetzalis</i> has a shorter horn, grooved only dorsally and finishing in a cannula. <i>O. xiphias</i> has sparse pilosity at the elytral apex; whereas <i>O</i>. <i>quetzalis</i> is densely pilose in this area. <i>O. xiphias</i> has a slightly shagreened pygidium with short setae; <i>O. quetzalis</i> has on the contrary a shagreened pygidium with much shorter setae. <i>O xiphias</i> has slightly shagreened abdominal sternites and the metasternal surface is covered by simple punctation; whereas <i>O. quetzalis</i> has shagreened abdominal sternites and the metasternal surface is covered by umbilicate punctures.</p> <p> Females of <i>O. xiphias</i> are separated from <i>O. quetzalis</i> using head characters. The first species has a more rounded head and a distinct cephalic carina, whereas the second species has a more pointed head and a weak cephalic carina.</p> <p> One specimen of <i>Onthophagus quetzalis</i> had been reported from Hartmann Finca, Chiriquí Prov., Panama (Howden and Gill, 1993). This specimen belongs to the new species here described.</p> <p> <b>Etymology.</b> Latinized word in apposition, taken from the Greek, <i>xiphias</i> (), meaning swordfish; making reference to the long and slender pronotal horn of the developed males of this species.</p>Published as part of <i>Solís, Ángel & Kohlmann, Bert, 2003, New species of dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae) from Costa Rica and Panama, pp. 1-14 in Zootaxa 139</i> on pages 4-9, DOI: <a href="http://zenodo.org/record/156789">10.5281/zenodo.156789</a>
Quality and quantity in item generation and reduction in developing a specific measure for women in mature life
No full textFragebogen zur Erhebung von Stress und Stressbewältigung im Kindes- und Jugendalter (SSKJ 3-8)
No full textLohaus A, Eschenbeck H, Kohlmann C-W, Klein-Heßling J. Fragebogen zur Erhebung von Stress und Stressbewältigung im Kindes- und Jugendalter (SSKJ 3-8). Göttingen: Hogrefe; 2006
Developing of a comprehensive questionnaire in post-menopausal women: results from multinational qualitative studies
No full textMeasuring health-related quality of life in mature women: a systematic overview od standardized questionnaires
No full textInstrumente zur Erfassung von Stress und Coping im Kindesalter
No full textEschenbeck H, Lohaus A, Kohlmann C-W. Instrumente zur Erfassung von Stress und Coping im Kindesalter. In: Seiffge-Krenke I, Lohaus A, eds. Stress und Stressbewältigung im Kindes- und Jugendalter. Göttingen: Hogrefe; 2007: 31-47
Fragebogen zur Erhebung von Stress und Stressbewältigung im Kindes- und Jugendalter (SSKJ 3-8)
No full textLohaus A, Eschenbeck H, Kohlmann C-W, Klein-Heßling J. Fragebogen zur Erhebung von Stress und Stressbewältigung im Kindes- und Jugendalter (SSKJ 3-8). Vollständig überarbeitete, erweiterte und neu normierte Auflage. Göttingen: Hogrefe; 2018
FIGURE 2. C. margaritae. a in New species of dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae) from Mexico and Costa Rica
No full textFIGURE 2. C. margaritae. a) Dorsal view of male head; b) dorsal view of female head; c) dorsal view of right male protibia; d) dorsal view of right female protibia; e) male pygidium; f) female pygidium; g) lateral view of aedeagus; h) dorsal view of brachypterous right wing of male.Published as part of Kohlmann, Bert & Solís, Ángel, 2006, New species of dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae) from Mexico and Costa Rica, pp. 61-68 in Zootaxa 1302 on page 64, DOI: 10.5281/zenodo.17365
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