1,284 research outputs found
Copris matthewsi subsp. pacificus Delgado and Kohlmann
No full textCopris matthewsi pacificus Delgado and Kohlmann (Fig. 5) We record here a new locality for this subspecies in El Salvador in cloud forest. EL SALVADOR. Chalatenango. El Pital, 3.IX.2002, 2640 m, 14 ° 39 ’ 45 ’’N, 89 ° 12 ’ 23 ’’W, E. Echeverría (4).Published as part of Kohlmann, Bert & Delgado, Enio Cano And Leonardo, 2003, New species and records of Copris (Coleoptera: Scarabaeidae; Scarabaeinae) from Central America, pp. 1-16 in Zootaxa 167 on page 15, DOI: 10.5281/zenodo.15678
Ateuchus earthorum Kohlmann & Solis, sp. nov.
No full textAteuchus earthorum Kohlmann & Solís, sp. nov. (Figs. 1 –2, 4) Diagnosis: This species is distinguished from other Costa Rican species by the following combination of characters: head disc finely punctured with coarse punctures at anterior margin; pronotum finely punctured with coarse punctures at the base of the pronotal midline; anterior pronotal margin incomplete; eyes viewed from above 3 times longer than wide; head and pronotum with coppery red reflections; body oval; profemur coarsely punctured; proepimeron without punctures; elytra not shagreened; pygidium very convex; last abdominal segment broad; internal sac with three hooks. Holotype description: Male (Figs. 1–2). Total length 6.7 mm. Elytral width 4.4 mm. Body slightly ovoid and convex, dorsum dark brown, head and pronotum with strong cupreous red reflections; venter dark brown. Clypeal margin coarsely punctate and broadly V-shaped, tooth on each side rounded, lateral margin arcuate, dorsal surface of head finely punctate and granulate, frons and vertex feebly tumid, eye dorsally small (ten times the interocular distance). Pronotum finely punctate and granulate, moderately punctate at posterior end of midline, midline impressed only one-third pronotal length, anterior pronotal margin incomplete. Proepisternum finely wrinkled, proepimeron granular. Elytral surface smooth and shiny; striae slightly impressed, more strongly so anteriorly; striae feebly punctate, intervals slightly convex. Pygidium very convex, surface slightly granulate and minutely punctate, completely grooved. Protibia quadridentate, basal tooth small, protibial spur oval; apical one-half of profemur ventrally coarsely punctate, punctures extending along posterior margin to base of femur, punctate area black; mesofemur and metafemur short, thick, with minute punctures near apex. Internal sac of the aedeagus (Fig. 2) with three hooks, one small, two large; three apical lamellae; and a well-developed, spiny fascies. Allotype: Female. Total length: 7.2 mm. Elytral width: 4.9 mm. Differs from the holotype by the following characters: Clypeal margin anteriorly moderately V-shaped, anterior clypeal border moderately punctate, protibia with acute, slender spur slightly bent apically; last abdominal segment broader, pygidium less convex. Variation: Total length: 6.3–7.2 mm. Elytral width: 4.1–4.9 mm. The color intensity varies of the cupreous-red head and pronotum. Material Examined (11 specimens): Holotype, male: COSTA RICA: Est. Cacao, 2 km SW del Cerro Cacao, Prov. Guanacaste, 1100 m, 12–14 SET 1995, C. Scarabaeidae, caca de caballo. L_N _ 323100 _ 375800, # 6292. Allotype, female: ibidem. Paratypes. ibidem, 5 males, 3 females; ibidem, caca de mono, 1 female. Remarks: This species will key to A. ginae in Kohlmann’s (1997) key. These species are cryptic and cannot be separated on the grounds of external morphology; only the internal sac differences will distinguish them. In both species there are three hooks, two are long and similar and the third is long and like a simple bar in A. ginae, whereas it is short and spine-like in A. earthorum (Fig. 2). These sac-hooks morphological differences are not only consistent (external morphology can be very variable and misleading) and geographically circumscribed to Cacao volcano, but also typical for separating Ateuchus species (Kohlmann 1984, 1997, 2000). Moreover, the ecology and geology are also critical; the Guanacaste volcanoes are a wellknown species generating area for small-sized dung beetle taxa like Ateuchus, Canthidium, and Onthophagus (Kohlmann 1997, Kohlmann & Solís 2001, Kohlmann & Wilkinson 2007, Kohlmann et al. 2007, Solís & Kohlmann 2004) and for plants (Araceae, Arecaceae, and Bromeliaceae) as well (Kohlmann et al. accepted). Habitat: The new species lives in mountain tropical forest at 1100 m and has been collected in September in horse manure and from monkey’s dung. Geographical distribution (Fig. 4): The new species is so far only known from the Pacific slope of Cacao volcano, in the province of Guanacaste. Chorological affinities: The new species is found at the same altitude, in the Guanacaste Cordillera, as its sister species, A. ginae Kohlmann, in the Central Cordillera (Kohlmann 1997). Taxonomic relationships: Ateuchus earthorum is hypothesized to be the sister species to A. ginae Kohlmann based on shared morphological characters discussed above. Etymology: The name is a Latinized noun in the genitive case. This species is dedicated to EARTH University in Costa Rica, an institution committed to the sustainable development of the humid tropics, to celebrate its 20 th anniversary.Published as part of Kohlmann, Bert & Solis, Angel, 2009, New species of Ateuchus and Canthidium (Coleoptera: Scarabaeidae: Scarabaeinae) from Costa Rica, pp. 31-37 in Zootaxa 2219 on pages 31-34, DOI: 10.5281/zenodo.19001
Onthophagus turgidus Kohlmann & Solis, new species
No full textOnthophagus turgidus Kohlmann & Solís, new species Figs. 8, 14 Diagnosis. This species is distinguished from other species of the O. dicranius species group by the following combination of characters: body dark reddish brown (Fig. 8); clypeal horn upright, basal portion wide, forked in its apical third, apical portion nearly parallel-sided; pronotum broadly tumescent anteriorly (Fig. 8), apical portion of tumescence with well separated lateral tubercles; anterior pronotal apical bead angularly reflexed medially. Description. Holotype. Male (Fig. 8): Length 10.3 mm. Humeral width 5.9 mm. Body oblong and dark brown. Vertex, pronotum, and elytral intervals closely punctate; vertex and pronotal punctures ocellate, surface between smooth; most elytral punctures with very minute setae. Clypeus at anterior median edge with an upright, slightly arched, and flattened horn (Fig. 8); horn in its apical third bifurcate, Y-shaped, apical portion nearly parallel-sided (Fig. 8). Side of clypeus from rounded edge of gena to base of horn almost straight. Clypeal surface behind the horn concave. Frons surface feebly convex medially and with scattered punctures. Vertex with very small tubercle on each side near anterior inner edge of each eye; surface anterior to and between tubercles punctate, punctures between eyes ocellate. Pronotum with anterior margin raised medially; anterior median half with large transverse tumosity (Fig. 8), delimited on each side by a distinct conical tubercle, tubercles evidently separated; anterior face of tumosity almost vertical with a small, slightly convex, tuberculated mid-line, tubercles very small. Pronotal surfaces near anterior lateral angles concave; marginal bead of posterior margin obsolete medially. Elytron with striae distinctly impressed with ocellate punctures at regular intervals; intervals on disc with two or more irregular rows of punctures, surface between shiny. Pygidium closely ocellate-punctate, each puncture with a short stiff seta. Metasternum with ocellate punctures, except along mid-line. Protibia elongate, with terminal tuft of setae; apical and subapical teeth distinctly closer to each other than second to third or third to fourth teeth. Ventral surface of all femora with regular, shallow punctures. Examined material (1 specimen). Holotype, male: PANAMA. Panamá, Bocas del Toro. Fila a 1.5 km este de río Tskui, 800 m. 9.4453º N - 82.8471º W. Col: A. Solís y M. Moraga. Trampa 11. Proyecto Darwin. Habitat. The specimen was collected with a trap baited with pig manure at an altitude of 800 m inside a primary tropical rain forest, during the month of October. Geographical distribution (Fig. 14). This species is known so far from the Caribbean slope on the Panamanian Central Cordillera. Chorological affinities. Onthophagus turgidus is found at similar altitudes in the Chiriquí Cordillera as O. solisi (500–1250 m; Kohlmann & Solís 2001; Fig. 9), its ecological equivalent (a small dung tunneller), in the Guanacaste and Tilarán Cordilleras. Taxonomic relationships. More material is needed, especially females, in order to establish taxonomic relationships. Presently, and using the similar clypeal horn, the anterior pronotal marginal bead, and the pronotal tumescence, the new species would seem to be closely related to O. solisi Howden & Gill, and it might actually be its sister species. This species will key to O. solisi in Kohlmann & Solís’ (2001) key. The male of the new species (Fig. 8) can be easily separated from the males of O. solisi (Fig. 9) by the form of the clypeal horn, which is thicker and more robust in the new species; as well as having a developed pronotal tumescence with more divergent lateral tubercles. With the description of these new species of Onthophagus in this paper, there are now 24 known from Panama. Etymology. Turgidus, Latin adjective in the genitive case meaning swollen, in reference to the transverse pronotal tumosity.Published as part of Kohlmann, Bert & Solís, Ángel, 2012, New species and revalidations of scarab beetles (Coleoptera: Geotrupidae: Athyreini and Coleoptera: Scarabaeidae: Scarabaeinae) from Costa Rica and Panama, pp. 28-52 in Zootaxa 3193 on pages 38-40, DOI: 10.5281/zenodo.21112
A new Ateuchus guatemalensis (Bates, 1887) (Coleoptera: Scarabaeidae) synonym from Mexico
No full textKohlmann, Bert (2019): A new Ateuchus guatemalensis (Bates, 1887) (Coleoptera: Scarabaeidae) synonym from Mexico. Zootaxa 4624 (4): 593-593, DOI: 10.11646/zootaxa.4624.4.1
History Of Scarabaeoid Classification
No full textKohlmann, Bert (2006): History Of Scarabaeoid Classification. The Coleopterists Bulletin (mo5) 60: 19-34, DOI: 10.1649/0010-065x(2006)60[19:hosc]2.0.co;
Canthon inusitatus Kohlmann & Solis, sp. nov.
No full textCanthon inusitatus Kohlmann & Solís, sp. nov. (Figs. 4, 5) Diagnosis. This species is distinguished by the following combination of characters: body oval, convex, black; dorsal surface shagreened; legs dark brown with a reddish hue. Posterior border of head clearly margined; second labial palpomere smaller than first; mentum entire; thorax strongly transverse; elytra not carinate; meso and metatibiae lacking transverse carinae; meso and metatarsomeres narrow and elongate, distal four tarsomeres as a group parallelsided or nearly so, first tarsomere very small, its length about half that of second; metatibial spur spinose and sharp. Description of holotype. Male. Length: 7.9 mm, humeral width: 5.9 mm. Body oval and convex, completely black; dorsal surface shagreened. Head with surface smooth, with fine punctures; posterior border of head clearly margined, margin with punctation at regular intervals; antenna brown, club grayishbrown. Clypeus anteriorly bidentate (Fig. 4), with a Vshaped emargination between teeth. Eyes small, dorsally only 7 facets wide and approximately twice as long as wide, separated by approximately 36 times their dorsal width (Fig. 4). Thorax much wider than long (Fig. 4); anterior angles well developed and acute; lateral borders forming angled arch; posterior angles poorly defined; anterior and lateral borders margined; disc very convex, finely punctured; without evident prescutellar impression. Prosternum with proepimeral carina absent. Elytra with striae nearly obsolete and finely punctate; interstriae shagreened, finely punctate, convex. Pygidium large (3.5 mm width versus 3.6 mm head width) and triangular; base not margined; disc convex, finely punctate. Protibia with three teeth on external edge, the apical tooth broadened towards apex with slender, acute apical spur (Fig. 4); ventral surface of metafemur lacking lines adjacent to posterior margin; metatibia slender and curved; meso and metatarsi long and slender, first tarsal article clearly shorter than second (Fig. 4); basal onethird of metafemur slender. Female. Unknown. Material examined (1 specimen). Holotype male: COSTA RICA. Heredia. 6 km ENE Vara Blanca, 2000 m, 13 abril 2002, 20/TF/05, D. Brenes, M. Paniagua y R. Vargas. Habitat. The species was collected in cloud forest (lower montane rain forest according to the Holdridge [1967] life zone system). Geographical distribution. This species is known only from the Caribbean slope of the Central Cordillera of Costa Rica (Fig. 5). Chorological affinities. The distribution of this new species represents, together with C. moniliatus, the northernmost distribution of the members of the “ Scybalocanthon ” group. Taxonomic relationships. Canthon inusitatus seems to have a number of different characters from the other species of the “ Scybalocanthon ” group, which Medina et al. (2003) have concluded is an artificial group. Canthon inusitatus will key to C. moniliatus Bates in couplet 8 in the Solís and Kohlmann (2002) key to the Canthon of Costa Rica. However, these two species differ in several characters and C. inusitatus can be easily separated from C. moniliatus by color (body all black versus head and elytra black and pronotum yellowbrown), male protibial spur (broad versus slender), and femoral color (unicolored versus black and yellow). Etymology. The word inusitatus is a Latin adjective in the nominative singular case, meaning unusual or extraordinary, in reference to such a big species having been found in such an unusual place for a Canthon, a cloud forest, after more than fifteen years of systematic collections in the area.Published as part of Kohlmann, Bert & Solís, Ángel, 2006, New species of dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae) from Mexico and Costa Rica, pp. 61-68 in Zootaxa 1302 on pages 65-68, DOI: 10.5281/zenodo.17365
FIGURE 2 in New species and revalidations of scarab beetles (Coleoptera: Geotrupidae: Athyreini and Coleoptera: Scarabaeidae: Scarabaeinae) from Costa Rica and Panama
No full textFIGURE 2. Dorsal habitus of the female allotype of Athyreus gulesseriani Kohlmann & Solís, new species.Published as part of Kohlmann, Bert & Solís, Ángel, 2012, New species and revalidations of scarab beetles (Coleoptera: Geotrupidae: Athyreini and Coleoptera: Scarabaeidae: Scarabaeinae) from Costa Rica and Panama, pp. 28-52 in Zootaxa 3193 on page 30, DOI: 10.5281/zenodo.21112
Checklist and distribution atlas of the Scarabaeinae (Coleoptera: Scarabaeidae) of Costa Rica
No full textSolís, Ángel, Kohlmann, Bert (2012): Checklist and distribution atlas of the Scarabaeinae (Coleoptera: Scarabaeidae) of Costa Rica. Zootaxa 3482: 1-32, DOI: 10.5281/zenodo.28237
Onthophagini
No full textONTHOPHAGINI ONTHOPHAGUS Latreille, 1802 Chalcoderus Erichson, 1848 Monapus Erichson, 1848 Psilax Erichson, 1848 Gonocyphus Lansberge, 1885 Tauronthophagus Shipp, 1895 Macropocopris Arrow, 1920 O. acuminatus Harold, 1880 O. andersoni Howden & Gill, 1987 O. anthracinus Harold, 1873 O. atriglabrus Howden & Gill, 1987 O. atrosericeus Boucomont, 1932 O. batesi Howden & Cartwright, 1963 O. championi Bates, 1887 O. chryses Bates, 1887 O. coriaceoumbrosus Kohlmann & Solís, 2001 O. coscineus Bates, 1887 O. digitifer Boucomont, 1932 O. crinitus Bates, 1887 O. panamensis Bates, 1887 O. cryptodicranius Kohlmann & Solís, 2001 O. cyanellus Bates, 1887 O. mesoamericanus Zunino & Halffter, 1988 O. dicranius Bates, 1887 O. dorsipilulus Howden & Gill, 1987 O. gazellinus Bates, 1887 O. genuinus Kohlmann & Solís, 2001 O. grataehelenae Kohlmann & Solís, 2001 O. hoepfneri Harold, 1869 O. incensus Say, 1835 O. inediapterus Kohlmann & Solís, 2001 O. landolti Harold, 1880 O. limonensis Kohlmann & Solís, 2001 O. marginicollis Harold, 1880 O. micropterus Zunino & Halffter, 1981 O. nemorivagus Kohlmann & Solís, 2001 O. notiodes Solís & Kohlmann, 2003 O. nubilus Kohlmann & Solís, 2001 O. nyctopus Bates, 1887 O. orphnoides Bates, 1887 O. praecellens Bates, 1887 O. propraecellens Howden & Gill, 1987 O. quetzalis Howden & Gill, 1993 O. sharpi Harold, 1875 O. singulariformis Kohlmann & Solís, 2001 O. solisi Howden & Gill, 1993 O. stockwelli Howden & Young, 1981 O. tapirus Sharp, 1887 O. viridivinosus Kohlmann & Solís, 2001Published as part of Solís, Ángel & Kohlmann, Bert, 2012, Checklist and distribution atlas of the Scarabaeinae (Coleoptera: Scarabaeidae) of Costa Rica, pp. 1-32 in Zootaxa 3482 on pages 8-9, DOI: 10.5281/zenodo.28237
Ateuchini
No full textATEUCHINI AGAMOPUS Bates, 1887 A. lampros Bates, 1887 ATEUCHUS Weber, 1801 Choeridium LePeletier & Serville, 1828 A. aeneomicans (Harold, 1868) A. alutacius Kohlmann & Solís, 2012 A. candezei (Harold, 1868) Choeridium poropyge Bates, 1887 A. earthorum Kohlmann & Solís, 2009 A. fetteri Kohlmann, 1997 A. ginae Kohlmann, 1997 A. hendrichsi Kohlmann, 1997 A. howdeni Kohlmann, 1997 A. rodriguezi (Preudhomme de Borre, 1886) Choeridium ampliatum Bates, 1887 A. solisi Kohlmann, 1997 A. zoebischi Kohlmann, 1997 BDELYRUS Harold, 1869 B. seminudus Bates, 1887 BRADYPODIDIUM Vaz-de-Mello, 2008 B. bradyporum (Boucomont, 1928) EUTRICHILLUM Martínez, 1969 E. arcus (Solís & Kohlmann, 2003) ONOREIDIUM Vaz-de-Mello, 2008 O. bottimeri (Howden & Young, 1981) SCATIMUS Erichson, 1847 S. erinnyos Kohlmann & Solís, 1996 S. ovatus Harold, 1862 S. patruelis Preudhomme de Borre, 1886 S. quadridentatus Balthasar, 1939 TRICHILLIDIUM Vaz-de-Mello, 2008 T. pilosum (Robinson, 1948) UROXYS Westwood, 1842 Pseuduroxys Balthasar, 1938 U. boneti Pereira & Halffter, 1961 U. bidentis Howden & Young, 1981 U. macrocularis Howden & Young, 1981 new synonym U. deavilai Delgado & Kohlmann, 2006 U. dybasi Howden & Young, 1981 U. gatunensis Howden & Young, 1981 U. gorgon Arrow, 1933 U. metagorgon Howden & Young, 1981 U. microcularis Howden & Young, 1981 U. micros Bates, 1887 U. nebulinus Howden & Gill, 1987 U. pauliani Balthasar, 1940 U. depressifrons Howden & Young, 1981 new synonym U. platypyga Howden & Young, 1981 U. transversifrons Howden & Gill, 1987Published as part of Solís, Ángel & Kohlmann, Bert, 2012, Checklist and distribution atlas of the Scarabaeinae (Coleoptera: Scarabaeidae) of Costa Rica, pp. 1-32 in Zootaxa 3482 on pages 4-5, DOI: 10.5281/zenodo.28237
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