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Ateuchus earthorum Kohlmann & Solis, sp. nov.
Ateuchus earthorum Kohlmann & Solís, sp. nov. (Figs. 1 –2, 4) Diagnosis: This species is distinguished from other Costa Rican species by the following combination of characters: head disc finely punctured with coarse punctures at anterior margin; pronotum finely punctured with coarse punctures at the base of the pronotal midline; anterior pronotal margin incomplete; eyes viewed from above 3 times longer than wide; head and pronotum with coppery red reflections; body oval; profemur coarsely punctured; proepimeron without punctures; elytra not shagreened; pygidium very convex; last abdominal segment broad; internal sac with three hooks. Holotype description: Male (Figs. 1–2). Total length 6.7 mm. Elytral width 4.4 mm. Body slightly ovoid and convex, dorsum dark brown, head and pronotum with strong cupreous red reflections; venter dark brown. Clypeal margin coarsely punctate and broadly V-shaped, tooth on each side rounded, lateral margin arcuate, dorsal surface of head finely punctate and granulate, frons and vertex feebly tumid, eye dorsally small (ten times the interocular distance). Pronotum finely punctate and granulate, moderately punctate at posterior end of midline, midline impressed only one-third pronotal length, anterior pronotal margin incomplete. Proepisternum finely wrinkled, proepimeron granular. Elytral surface smooth and shiny; striae slightly impressed, more strongly so anteriorly; striae feebly punctate, intervals slightly convex. Pygidium very convex, surface slightly granulate and minutely punctate, completely grooved. Protibia quadridentate, basal tooth small, protibial spur oval; apical one-half of profemur ventrally coarsely punctate, punctures extending along posterior margin to base of femur, punctate area black; mesofemur and metafemur short, thick, with minute punctures near apex. Internal sac of the aedeagus (Fig. 2) with three hooks, one small, two large; three apical lamellae; and a well-developed, spiny fascies. Allotype: Female. Total length: 7.2 mm. Elytral width: 4.9 mm. Differs from the holotype by the following characters: Clypeal margin anteriorly moderately V-shaped, anterior clypeal border moderately punctate, protibia with acute, slender spur slightly bent apically; last abdominal segment broader, pygidium less convex. Variation: Total length: 6.3–7.2 mm. Elytral width: 4.1–4.9 mm. The color intensity varies of the cupreous-red head and pronotum. Material Examined (11 specimens): Holotype, male: COSTA RICA: Est. Cacao, 2 km SW del Cerro Cacao, Prov. Guanacaste, 1100 m, 12–14 SET 1995, C. Scarabaeidae, caca de caballo. L_N _ 323100 _ 375800, # 6292. Allotype, female: ibidem. Paratypes. ibidem, 5 males, 3 females; ibidem, caca de mono, 1 female. Remarks: This species will key to A. ginae in Kohlmann’s (1997) key. These species are cryptic and cannot be separated on the grounds of external morphology; only the internal sac differences will distinguish them. In both species there are three hooks, two are long and similar and the third is long and like a simple bar in A. ginae, whereas it is short and spine-like in A. earthorum (Fig. 2). These sac-hooks morphological differences are not only consistent (external morphology can be very variable and misleading) and geographically circumscribed to Cacao volcano, but also typical for separating Ateuchus species (Kohlmann 1984, 1997, 2000). Moreover, the ecology and geology are also critical; the Guanacaste volcanoes are a wellknown species generating area for small-sized dung beetle taxa like Ateuchus, Canthidium, and Onthophagus (Kohlmann 1997, Kohlmann & Solís 2001, Kohlmann & Wilkinson 2007, Kohlmann et al. 2007, Solís & Kohlmann 2004) and for plants (Araceae, Arecaceae, and Bromeliaceae) as well (Kohlmann et al. accepted). Habitat: The new species lives in mountain tropical forest at 1100 m and has been collected in September in horse manure and from monkey’s dung. Geographical distribution (Fig. 4): The new species is so far only known from the Pacific slope of Cacao volcano, in the province of Guanacaste. Chorological affinities: The new species is found at the same altitude, in the Guanacaste Cordillera, as its sister species, A. ginae Kohlmann, in the Central Cordillera (Kohlmann 1997). Taxonomic relationships: Ateuchus earthorum is hypothesized to be the sister species to A. ginae Kohlmann based on shared morphological characters discussed above. Etymology: The name is a Latinized noun in the genitive case. This species is dedicated to EARTH University in Costa Rica, an institution committed to the sustainable development of the humid tropics, to celebrate its 20 th anniversary.Published as part of Kohlmann, Bert & Solis, Angel, 2009, New species of Ateuchus and Canthidium (Coleoptera: Scarabaeidae: Scarabaeinae) from Costa Rica, pp. 31-37 in Zootaxa 2219 on pages 31-34, DOI: 10.5281/zenodo.19001
Copris matthewsi subsp. pacificus Delgado and Kohlmann
Copris matthewsi pacificus Delgado and Kohlmann (Fig. 5) We record here a new locality for this subspecies in El Salvador in cloud forest. EL SALVADOR. Chalatenango. El Pital, 3.IX.2002, 2640 m, 14 ° 39 ’ 45 ’’N, 89 ° 12 ’ 23 ’’W, E. Echeverría (4).Published as part of Kohlmann, Bert & Delgado, Enio Cano And Leonardo, 2003, New species and records of Copris (Coleoptera: Scarabaeidae; Scarabaeinae) from Central America, pp. 1-16 in Zootaxa 167 on page 15, DOI: 10.5281/zenodo.15678
Anthony Kohlmann, S.J. (7 of 11)
Contains ALS: from E. Fenwick to Roger Baxter, SJ 7/7/1820; from Kenney to Marshall on personnel at Georgetown 8/7/1820; from A. Kohlmann to E. Fenwick urging latter to be rector of Georgetown 8/10/1820; from E. Fenwick to A. Kohlmann on former's appointment as rector 8/10/1820; from Dubuisson on events at Conewago 8/11/1820; from Marshall to E. Fenwick on bad state of plantations 8/14/1820; from DeBarth to Marshall on bad state of Conewago farms 8/26/1820, on Goshenhoppen law suit 9/17/1820, on German Fund in Pennsylvania and needs of Conewago 9/26/1820; from Grassi to McElroy on arrival of Americans in Naples 8/27/1820; from Joseph Schneller, SJ to Mulledy 9/2/1820; from E. Fenwick to F. Neale complaining of condition of Georgetown 9/6/1820; from Marshall to Neale 9/17/1820, on overseer at St. Thomas and Goshenhoppen law suit 9/22/1820, asking power to sell bank stock 9/27/1820; from Abp Marechal to A. Kohlmann on Baxter's suspension (undated).**Former finding aid locations: 119_59_6; 205G1-205G12*
Onthophagus turgidus Kohlmann & Solis, new species
Onthophagus turgidus Kohlmann & Solís, new species Figs. 8, 14 Diagnosis. This species is distinguished from other species of the O. dicranius species group by the following combination of characters: body dark reddish brown (Fig. 8); clypeal horn upright, basal portion wide, forked in its apical third, apical portion nearly parallel-sided; pronotum broadly tumescent anteriorly (Fig. 8), apical portion of tumescence with well separated lateral tubercles; anterior pronotal apical bead angularly reflexed medially. Description. Holotype. Male (Fig. 8): Length 10.3 mm. Humeral width 5.9 mm. Body oblong and dark brown. Vertex, pronotum, and elytral intervals closely punctate; vertex and pronotal punctures ocellate, surface between smooth; most elytral punctures with very minute setae. Clypeus at anterior median edge with an upright, slightly arched, and flattened horn (Fig. 8); horn in its apical third bifurcate, Y-shaped, apical portion nearly parallel-sided (Fig. 8). Side of clypeus from rounded edge of gena to base of horn almost straight. Clypeal surface behind the horn concave. Frons surface feebly convex medially and with scattered punctures. Vertex with very small tubercle on each side near anterior inner edge of each eye; surface anterior to and between tubercles punctate, punctures between eyes ocellate. Pronotum with anterior margin raised medially; anterior median half with large transverse tumosity (Fig. 8), delimited on each side by a distinct conical tubercle, tubercles evidently separated; anterior face of tumosity almost vertical with a small, slightly convex, tuberculated mid-line, tubercles very small. Pronotal surfaces near anterior lateral angles concave; marginal bead of posterior margin obsolete medially. Elytron with striae distinctly impressed with ocellate punctures at regular intervals; intervals on disc with two or more irregular rows of punctures, surface between shiny. Pygidium closely ocellate-punctate, each puncture with a short stiff seta. Metasternum with ocellate punctures, except along mid-line. Protibia elongate, with terminal tuft of setae; apical and subapical teeth distinctly closer to each other than second to third or third to fourth teeth. Ventral surface of all femora with regular, shallow punctures. Examined material (1 specimen). Holotype, male: PANAMA. Panamá, Bocas del Toro. Fila a 1.5 km este de río Tskui, 800 m. 9.4453º N - 82.8471º W. Col: A. Solís y M. Moraga. Trampa 11. Proyecto Darwin. Habitat. The specimen was collected with a trap baited with pig manure at an altitude of 800 m inside a primary tropical rain forest, during the month of October. Geographical distribution (Fig. 14). This species is known so far from the Caribbean slope on the Panamanian Central Cordillera. Chorological affinities. Onthophagus turgidus is found at similar altitudes in the Chiriquí Cordillera as O. solisi (500–1250 m; Kohlmann & Solís 2001; Fig. 9), its ecological equivalent (a small dung tunneller), in the Guanacaste and Tilarán Cordilleras. Taxonomic relationships. More material is needed, especially females, in order to establish taxonomic relationships. Presently, and using the similar clypeal horn, the anterior pronotal marginal bead, and the pronotal tumescence, the new species would seem to be closely related to O. solisi Howden & Gill, and it might actually be its sister species. This species will key to O. solisi in Kohlmann & Solís’ (2001) key. The male of the new species (Fig. 8) can be easily separated from the males of O. solisi (Fig. 9) by the form of the clypeal horn, which is thicker and more robust in the new species; as well as having a developed pronotal tumescence with more divergent lateral tubercles. With the description of these new species of Onthophagus in this paper, there are now 24 known from Panama. Etymology. Turgidus, Latin adjective in the genitive case meaning swollen, in reference to the transverse pronotal tumosity.Published as part of Kohlmann, Bert & Solís, Ángel, 2012, New species and revalidations of scarab beetles (Coleoptera: Geotrupidae: Athyreini and Coleoptera: Scarabaeidae: Scarabaeinae) from Costa Rica and Panama, pp. 28-52 in Zootaxa 3193 on pages 38-40, DOI: 10.5281/zenodo.21112
Anthony Kohlmann, S.J. (9 of 11)
Contains ALS: from A. Kohlmann to Cary on Newport and Cobb Neck properties Jan 1821; from Beschter to Cary on procuring a painting for St. John's, Baltimore 1/3/1821; from Marshall to F. Neale on income for Georgetown, debt of Conewago, destructive influence of A. Kohlmann on Georgetown 1/5/1821, on conditions for sale of New York property and New York debts 1/20/1821; from Beschter to F. Neale on clergy problems in Philadelphia, arrival of Bp Dubourg in New Orleans 1/11/1821; from F. Neale to Marshall on sale of New York and Philadelphia property 1/29/1821, on willingness to sell St. Thomas to lessen debt, cost of board at Georgetown 2/14/1821; from McElroy to Grassi on enrollment at Georgetown 2/6/1821; from DeBarth to Marshall on Conewago debt 2/7/1821, on Goshenhoppen farming 3/6/1821, on Conewago debt 3/21/1821; from Joseph Snyder to Marshall on problems with Rev. Wm. Hogan at St. Mary's, Phila 3/1/1821, 3/12/1821, 3/28/1821; from Beschter to Grassi on difficulties of reforming Georgetown, suspension of Hogan 3/31/1821.**Former finding aid locations: 119_59_4; 205D1-205D13*
Anthony Kohlmann, S.J. (4 of 11)
Contains ALS: from Grassi to E. Fenwick on former's staying at Rome 7/14/1819; from Grassi to A. Kohlmann on various artworks sent to America 7/14/1819; from Grassi to McElroy on shipment of paintings for Georgetown 7/14/1819; from E. Fenwick to G. Fenwick 7/20/1819; from Grassi to F. Neale 7/22/1819; to Ryder on Ryder and Mulledy's trip to Conewago 8/15/1819; to G. Fenwick 8/22/1819; from E. Fenwick to his mother Margaret on yellow fever epidemic in Baltimore 9/1/1819; from E. Fenwick to F. Neale on yellow fever epidemic 9/29/1819, on Baltimore Cathedral 12/29/1819; from Kenney on disagreements among Georgetown clergy 10/5/1819; from Kenney to F. Neale asking advice for running of Georgetown 10/8/1819; from A. Kohlmann to Beschter on overseer for St. Thomas 10/8/1819; from Abp Marechal to F. Neale on misc debt 11/9/1819, on conveyance of land to ladies of Georgetown 12/8/1819; from E. Fenwick to Scholastics on paintings for Georgetown mistakenly sent to Baltimore Cathedral 12/20/1819.Former finding aid locations: 119_59_9; 205M1-202M1
Canthon inusitatus Kohlmann & Solis, sp. nov.
Canthon inusitatus Kohlmann & Solís, sp. nov. (Figs. 4, 5) Diagnosis. This species is distinguished by the following combination of characters: body oval, convex, black; dorsal surface shagreened; legs dark brown with a reddish hue. Posterior border of head clearly margined; second labial palpomere smaller than first; mentum entire; thorax strongly transverse; elytra not carinate; meso and metatibiae lacking transverse carinae; meso and metatarsomeres narrow and elongate, distal four tarsomeres as a group parallelsided or nearly so, first tarsomere very small, its length about half that of second; metatibial spur spinose and sharp. Description of holotype. Male. Length: 7.9 mm, humeral width: 5.9 mm. Body oval and convex, completely black; dorsal surface shagreened. Head with surface smooth, with fine punctures; posterior border of head clearly margined, margin with punctation at regular intervals; antenna brown, club grayishbrown. Clypeus anteriorly bidentate (Fig. 4), with a Vshaped emargination between teeth. Eyes small, dorsally only 7 facets wide and approximately twice as long as wide, separated by approximately 36 times their dorsal width (Fig. 4). Thorax much wider than long (Fig. 4); anterior angles well developed and acute; lateral borders forming angled arch; posterior angles poorly defined; anterior and lateral borders margined; disc very convex, finely punctured; without evident prescutellar impression. Prosternum with proepimeral carina absent. Elytra with striae nearly obsolete and finely punctate; interstriae shagreened, finely punctate, convex. Pygidium large (3.5 mm width versus 3.6 mm head width) and triangular; base not margined; disc convex, finely punctate. Protibia with three teeth on external edge, the apical tooth broadened towards apex with slender, acute apical spur (Fig. 4); ventral surface of metafemur lacking lines adjacent to posterior margin; metatibia slender and curved; meso and metatarsi long and slender, first tarsal article clearly shorter than second (Fig. 4); basal onethird of metafemur slender. Female. Unknown. Material examined (1 specimen). Holotype male: COSTA RICA. Heredia. 6 km ENE Vara Blanca, 2000 m, 13 abril 2002, 20/TF/05, D. Brenes, M. Paniagua y R. Vargas. Habitat. The species was collected in cloud forest (lower montane rain forest according to the Holdridge [1967] life zone system). Geographical distribution. This species is known only from the Caribbean slope of the Central Cordillera of Costa Rica (Fig. 5). Chorological affinities. The distribution of this new species represents, together with C. moniliatus, the northernmost distribution of the members of the “ Scybalocanthon ” group. Taxonomic relationships. Canthon inusitatus seems to have a number of different characters from the other species of the “ Scybalocanthon ” group, which Medina et al. (2003) have concluded is an artificial group. Canthon inusitatus will key to C. moniliatus Bates in couplet 8 in the Solís and Kohlmann (2002) key to the Canthon of Costa Rica. However, these two species differ in several characters and C. inusitatus can be easily separated from C. moniliatus by color (body all black versus head and elytra black and pronotum yellowbrown), male protibial spur (broad versus slender), and femoral color (unicolored versus black and yellow). Etymology. The word inusitatus is a Latin adjective in the nominative singular case, meaning unusual or extraordinary, in reference to such a big species having been found in such an unusual place for a Canthon, a cloud forest, after more than fifteen years of systematic collections in the area.Published as part of Kohlmann, Bert & Solís, Ángel, 2006, New species of dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae) from Mexico and Costa Rica, pp. 61-68 in Zootaxa 1302 on pages 65-68, DOI: 10.5281/zenodo.17365
Anthony Kohlmann, S.J. (1 of 11)
Contains ALS: from Grassi to E. Fenwick on procurement of art for Baltimore Cathedral 3/13/1818; from Abp Marechal to F. Neale 7/6/1817; from Peter Menard about a dictionary of Indian language 7/21/1817; from Grassi to A. Kohlmann on arrival in France 8/7/1817; from Grassi to E. Fenwick on appointment of Marechal as Abp, efforts to obtain bishop for Virginia 9/21/1817; from Grassi to Cary on trip to Rome 9/21/1817; to Cary on unspecified land purchase 10/26/1817.Former finding aid locations: 119_59_12; 205R1-205S
Canthidium margaritae Kohlmann & Solis, sp. nov.
<i>Canthidium margaritae</i> Kohlmann & Solís sp. nov. (Figs. 1–3) <p> <b>Diagnosis.</b> This species is distinguished by the following combination of characters: body globose; head and pronotum uniformly and strongly punctate; frons swollen; frontoclypeal region with a small, smooth swelling medial to each eye; eye dorsally at posterior end of gena narrow, eight to ten facets wide, interocular distance separated by approximately 8–10 times maximal eye width; basal pronotal border not margined; elytra with eight striae, surface lightly shagreened, microsculpture coarser towards apex; hind wing brachypterous.</p> <p> <b>Description of Holotype.</b> Male. Length: 4.5 mm, humeral width: 2.9 mm. Body form globose (Fig. 1). Head and pronotum coppery­red, elytra dark brown (Fig.1). Head and pronotum strongly punctate and devoid of setae.</p> <p>Clypeus rugosely punctate, apex strongly bidentate, median emargination broadly Vshaped (Fig. 2 a); eye dorsally at posterior end of gena narrow, eight to ten facets wide, eyes separated by approximately 8–10 times maximum eye width (Fig. 2 a). Frons and vertex of head strongly punctate, frons swollen; two small, black, smooth swellings between eyes (Fig. 2 a).</p> <p>Pronotum on disc and posterior angles slightly shagreened; surface strongly and uniformly punctate; lateral fovea oval; posterior margin lacking elongated punctures or groove.</p> <p>Elytral striae consisting of clear indented lines interrupted by fairly evenly spaced oval punctures (approximately the maximum length of one puncture); intervals shiny, finely punctate; surface shiny and slightly shagreened especially towards apex. Elytra globose; hindwing brachypterous (Fig. 2 h).</p> <p>Pygidium distinctly punctate, more coarsely so in basal half, surface shiny and slightly shagreened at base. Genitalia as in Fig. 2 g.</p> <p>Protibia with three teeth on external border (Fig. 2c), inner apical margin of protibia produced into a rounded anterior projection and slightly bent downwards (Fig. 2c), apical spur with incurved apex (Fig. 2c). Pro­, meso­ and metafemora with ventral surface finely punctate and finely shagreened.</p> <p> <b>Allotype.</b> Female. Length: 4.6 mm, humeral width: 3.1 mm. Differing from male in the following major characters: clypeus slightly more transversely rugose (Fig. 2 b), apical spur not forming incurved apex and claw­bearing protarsus not as thick (Fig. 2 d), pygidium broader and less heavily punctate (Fig. 2 f), last abdominal segment broader, inner apex of protibia not forming a rounded lobe (Fig. 2 d).</p> <p> <b>Variation.</b> Elytral surface can range from completely shagreened to shagreened only on the apical third.</p> <p> <b>Material examined</b> (8 specimens). <b>Holotype</b> male: MÉXICO. Estado de México. Sierra de Nanchititla, Palos Prietos, 28–29­julio­2005, Alt. 1750 m, coprotrampa, M. Castillo, A. y L. Delgado cols. <b>Allotype</b> female: <i>ibidem</i>. Paratypes: MÉXICO. Estado de México. Sierra de Nanchititla, 18–19­VIII­96, 1800 msnm, G. Nogueira col., (1 female); same data as holotype (4 males, 1 female).</p> <p> <b>Habitat.</b> This species has been found in cloud forest mixed with oaks in elevations ranging from 1,750 to 1,800 m above sea level.</p> <p> <b>Geographical distribution.</b> It is only known in the State of México in the Sierra de Nanchititla on the upper reaches of the Balsas river depression (Fig. 3).</p> <p> <b>Chorological affinities.</b> The known range of <i>Canthidium margaritae</i> is widely separated from that of a very similar species, <i>C. riverai</i>, which is distributed in the Manantlán (Jalisco State) and the Coacolmán (Michoacán state) mountain ranges at similar altitudes (960–2,000 m above sea level) and in cloud forests.</p> <p> <b>Taxonomic relationships</b>. <i>Canthidium margaritae</i> is postulated to be the sister species of <i>C. riverai</i>. They are both globose in body shape and brachypterous, they also have a swollen frons, clearly punctate pronotum, shagreened elytra, evenly impressed striae, and they both inhabit cloud forests. This species pair apparently represents a vicariant speciation event between the Sierra Madre del Sur (<i>C</i>. <i>riverai</i>) and the Neovolcanic Axis (<i>C</i>. <i>margaritae</i>).</p> <p> <i>C. margaritae</i> will key to <i>C. riverai</i> Kohlmann and Solís in couplet 12 in the Kohlmann and Solís (2006) key for the <i>Canthidium</i> of Mexico. The two species are very similiar, but <i>C. margaritae</i> can be easily separated from <i>C. riverai</i> by having bigger eyes (8–10 eye facets versus 2–3 eye facets), head and pronotal punctures less coarse, elytra less shagreened, a different metatibial form (internal apical angle projected like a tapering rectangle versus apically obliquely truncated), and differences in the form of the parameres (parameres taper evenly towards apex versus parameres with a small hump at apical two­thirds).</p> <p> <b>Etymology.</b> We dedicate this species to Margarita Castillo, who has always supported Luis Leonardo Delgado in his entomological studies and also helped collect this new species. The name is derived from the latinized (<i>margarita</i>) Greek word μαργαρίτη, meaning a pearl.</p>Published as part of <i>Kohlmann, Bert & Solís, Ángel, 2006, New species of dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae) from Mexico and Costa Rica, pp. 61-68 in Zootaxa 1302</i> on pages 62-65, DOI: <a href="http://zenodo.org/record/173658">10.5281/zenodo.173658</a>
Transcription, "Exhortatio ad Nigros Servos"
Contains a transcription of the “Exhortatio ad Nigros Servos” (“An exhortation to black slaves”). This undated, typed transcription also contains several notes identifying the place, date, and community addressed in the “Exhortatio,” and suggests that its author was Anthony Kohlmann, S.J. There are also photocopies of the collection’s previous card catalog record, which suggests that the author was Paul Kohlmann, S.J.This folder contains materials related to Jesuit slaveholding
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