20,774 research outputs found
Ancyronyx henningi Freitag & Kodada 2017
Ancyronyx henningi Freitag & Kodada, 2017 (Fig. 2 F) Material examined. 2 LL (NMW, CFM [FR131]): “ INDONESIA: C Sulawesi, Rd. Km 32 Tentena-Gintu; mountain creek, grass-land; submerged wood, c. 1600m asl., c. 01°48′10′′ S 120°28′10′′E 0 4 Jan.1998 leg. Freitag (Sul 13f)”. Differential diagnosis (based on presumably fifth instar, HW: 0.33 mm). In most characters very similar to Ancyronyx tobada except for: dominantly pale brown dorsal side, dark brown areas limited to dorsal portion of head and subapical portion of terminal abdominal segment (Fig. 2 F). Anterior half of pronotum yellowish, remaining thoracic and abdominal segments pale brown with darker posteromedian brown patches (most distinct at pronotum) and darker sublateral areas; abdominal apex yellowish (Fig. 2 F). Entire ventral side more or less pale. Dorsal tubercles moderately elevated, lateral protuberances distinctly protruding. Head slightly tapering anteriad, widest at posterior 0.3. Abdominal segments moderately elevated at posterior portions, with few indistinct scattered short erected setae; posterolateral protuberances with prominent tubercles and two distinct, long setae; terminal segment IX long, ca. 2.3 times as long as wide; basomedian portion almost glabrous, only with few small tubercles; apex distinctly notched.Published as part of Freitag, Hendrik & Kodada, Ján, 2017, Larvae of Ancyronyx Erichson, 1847 (Insecta: Coleoptera: Elmidae) from Sulawesi, using DNA sequences for the assignment of the larval stages, pp. 121-130 in Zootaxa 4299 (1) on page 129, DOI: 10.11646/zootaxa.4299.1.6, http://zenodo.org/record/83515
Ancyronyx skalei Freitag & Kodada 2017
Ancyronyx skalei Freitag & Kodada, 2017 (Fig. 2 B) Material examined. 3LL (NMW, CKB, CFM [FR126]): “ INDONESIA: Sulawesi, T. Toraja, Salu Toriu River, 810m asl., c. 02°56′38′′ S 119°52′24′′E 0 2 Jan.1998 leg. Freitag (Sul 8 Af)”. Differential diagnosis (based on presumably fifth instar, HW: 0.32 mm). Very similar to the previous species, except for the following: anterior quarter of pronotum and posterior quarter of terminal abdominal segment yellowish to pale brown (Fig. 2 B); lateral portions of the head, entire first abdominal segment and posterior half/entire eight abdominal segment pale; small transverse pale spots might also be present at medioposterior margin of pro-, meso-, and metanotum. Dorsal tubercles slightly elevated. Head more tapering anteriad, widest at posterior 0.2; antennae rather small, approximately reaching anterior margin of labrum. Abdominal segment IX moderately long, ca. 1.9 times as long as wide; basomedian portion slightly tuberculate; apex distinctly emarginated.Published as part of Freitag, Hendrik & Kodada, Ján, 2017, Larvae of Ancyronyx Erichson, 1847 (Insecta: Coleoptera: Elmidae) from Sulawesi, using DNA sequences for the assignment of the larval stages, pp. 121-130 in Zootaxa 4299 (1) on page 126, DOI: 10.11646/zootaxa.4299.1.6, http://zenodo.org/record/83515
Ancyronyx tobada Freitag & Kodada 2017
Ancyronyx tobada Freitag & Kodada, 2017 (Figs 2 C; 4A, C, E) Material examined. 3LL (CKB, CFM [FR114]): “ INDONESIA: C Sulawesi, Rd. Km 43 Tentena-Gintu; mountain creek; cloud forest; submerged wood, c. 1680m asl., c. 01°48′39′′ S 120°23′17′′ E 0 4 Jan.1998 leg. Freitag (Sul 14f)”; 6LL (CKB [SEM], NMW, CFM [FR 119]): “ INDONESIA: C Sulawesi, Rd. Tentena-Pendola, 9km E Tentena, secondary forest; submerged wood, c. 860m asl., c. 01°46′31′′ S 120°43′05′′ E 0 3 Jan.1998 leg. Freitag (Sul 10f)”; 4LL (CFM [FR121]): “ INDONESIA: S Sulawesi, Rd. Km 56 Tentena-Gintu; black-water river; submerged wood, c. 1190m asl., c. 01°49′28′′ S 120°21′43′′E 0 4 Jan.1998 leg. Freitag (Sul16f)”. Differential diagnosis (based on presumably sixth instar, HW: 0.32 mm). Generally similar to Ancyronyx schoedli except for the following characters: pronotum yellowish in anterior third, yellowish lateral portions of head extending posteriad along the frontal suture; posterior margins of thoracic and abdominal terga slightly paler, somewhat dark yellow-brown; terminal abdominal segment dark along anterior 0.1, yellow-brown along anterior 0.1–0.6, dark brown along posterior 0.6–0.9 and pale yellow-brown at the tip. Entire ventral side pale yellowish, only abdominal segment IX occasionally slightly darker, but still somewhat pale. Dorsal tubercles moderately elevated (Figs 4 A, C). Head (Figs 2 C, 4A) slightly tapering anteriad, widest at posterior 0.2; antennae overreaching anterior margin of labrum. Abdominal segments gently elevated near entire posterior portion; dorsum with scattered short setae (Figs 4 C, E); lateral protuberances with tubercles almost as large as spiracles; segment IX (Figs 2 C, 4E) very long, ca. 2.4 times as long as wide; basomedian portion glabrous, without tubercles; apex deeply emarginated.Published as part of Freitag, Hendrik & Kodada, Ján, 2017, Larvae of Ancyronyx Erichson, 1847 (Insecta: Coleoptera: Elmidae) from Sulawesi, using DNA sequences for the assignment of the larval stages, pp. 121-130 in Zootaxa 4299 (1) on page 126, DOI: 10.11646/zootaxa.4299.1.6, http://zenodo.org/record/83515
Ancyronyx longiparamerus Freitag & Kodada 2017
Ancyronyx longiparamerus Freitag & Kodada, 2017 (Figs 2 D; 4B, D, F) Material examined. 2LL (NMW, CFM): “ INDONESIA: C Sulawesi, Rd. Km 43 Tentena-Gintu; mountain creek; cloud forest; submerged wood, c. 1680m asl., c. 01°48′39′′ S 120°23′17′′E 0 4 Jan.1998 leg. Freitag (Sul 14f)”; 2LL (CFM [FR132], CKB [SEM]): “ INDONESIA: S Sulawesi, 17.5km NNW Rantepao 1570m asl., small mount. river “Salu Malarak”, pine forest, run/riffle; submerged wood, c. 02°48′51′′ S 119° 51′21′′E 31 Dec.1997 leg. Freitag (Sul7f)”. Differential diagnosis (based on presumably sixth instar, HW: 0.33 mm). Very similar to Ancyronyx tobada except for the following characters: generally the colour is somewhat paler brown, surface of pronotum yellowish on extensive oblique anterolateral portion (Fig. 2 D); yellowish lateral portions of head extending posteriad; entire posterior half of abdominal segments I and VIII indistinctly paler; terminal abdominal segment dark except of paler brownish apex. Entire ventral side pale. Dorsal tubercles prominent and distinctly elevated (Figs 4 D, F); anterolateral portion of all thoracic and abdominal segments with glabrous signa, most conspicuous at pronotum. Head (Fig. 4 B) distinctly tapering anteriad, widest at posterior 0.1. Abdominal segments with scattered short erected setae (Fig. 4 F); posterolateral protuberances with prominent tubercles, which are distinctly smaller than spiracles; terminal segment IX (Fig. 4 F) moderately short, ca. 1.9 times as long as wide; basomedian portion distinctly and densely tuberculate; apex slightly emarginated.Published as part of Freitag, Hendrik & Kodada, Ján, 2017, Larvae of Ancyronyx Erichson, 1847 (Insecta: Coleoptera: Elmidae) from Sulawesi, using DNA sequences for the assignment of the larval stages, pp. 121-130 in Zootaxa 4299 (1) on pages 127-128, DOI: 10.11646/zootaxa.4299.1.6, http://zenodo.org/record/83515
Ancyronyx schoedli Freitag & Kodada 2017
Ancyronyx schoedli Freitag & Kodada, 2017 (Figs 2 A; 3A–F) Material examined. 4LL (CKB [SEM], CFM [FR128, FR129]): “ INDONESIA: Sulawesi, T. Toraja, Salu Toriu River, 810m asl., c. 02°56′38′′ S 119°52′24′′E 0 2 Jan.1998 leg.Freitag (Sul8Af)”; 1L (NMW): “S - SULAWESI 1992 Malino-Manipi 700m (31) leg. Jäch 1.V. ”. Larval diagnosis (based on presumably fifth instar, HW: 0.30 mm). Body elongate, nearly semicircular in cross section, dorsally vaulted, ventrally almost flat. Posterolateral margins of abdominal segments I–VII moderately produced (Fig. 3 C). Colour (Fig. 2 A) dorsally dominantly dark brown except for pale brown to yellowish lateral portion of head, apex of the terminal abdominal segment (up to posterior 0.15) and the entire first and eight abdominal segments; clypeus and labrum reddish brown; legs, antennae, remaining mouthparts, entire ventral side including operculum and except abdominal segment IX yellowish to pale brown. The larva is very similar in the external characters to that of A. minerva (see Freitag & Balke 2011), except for the following characters: lateral abdominal projections (Figs 3 C, D) with two large robust long setae (referred to as “trichoid teeth” in Freitag & Balke 2011), one robust short seta and one long thin trichoid seta. Dorsal tubercles slightly elevated (Figs 3 A–D). Head (Fig. 3 A) widest at posterior 0.3, sides arcuate in posterior half; frontal suture V-shaped, conspicuous in wet specimens; dorsolateral portion with three pairs of conspicuous setae: first pair consists of short setae situated mediofrontally, second pair of moderately long setae inserted near the frontoclypeal suture and third pair of long setae originate near frontal suture behind the eyes; lateral portion with about 4 pairs of moderately long setae. Antennae moderately long, overreaching anterior margin of labrum; setae of scapus trichoid. Labrum with subapical fringe of ramose setae and few trichoid setae laterally. Maxillary palpus ca. as long as stipes wide. Mentum subrectangular, slightly narrower along posterior 0.1–0.4; pair of trichoid setae moderately long, not reaching anterior margin, inserted sublaterally at anterior 0.35 of mentum. Pro- meso- and metathorax subtrapezoidal, slightly narrowed anteriorly, broader than long; with slightly produced lateral rim; lateral rim with few scattered long trichoid setae; dorsosagittal carinae small, narrow and present at anterior half of mesonotum; surface of meso- and metanotum with microtubercles; pronotum with ca. six pairs of rather inconspicuous, irregularly shaped and partly fused signa (flat, subglabrous areas) in posterior half (Figs 3 A, B); remaining areas of pronotum including anterolateral grooves with setiferous tubercles. Ventral sclerites of thorax granulose. Abdomen (Figs 2 A, 3C–F) without dorsosagittal carina, with distinct mesal suture except for segment IX (Figs 3 C, E); lateral rim with few scattered long, trichoid setae; squamose setae at posterior rim of segments large; ventral sclerites of segment I with rather indistinct sagittal ridge in anterior half; segment IX (Fig. 3 E) moderately short, ca. 1.6 times as long as wide; basomedian portion tuberculate, apical portion rugose; apex slightly emarginated; operculum ca. 1.7 times as long as wide.Published as part of Freitag, Hendrik & Kodada, Ján, 2017, Larvae of Ancyronyx Erichson, 1847 (Insecta: Coleoptera: Elmidae) from Sulawesi, using DNA sequences for the assignment of the larval stages, pp. 121-130 in Zootaxa 4299 (1) on pages 124-126, DOI: 10.11646/zootaxa.4299.1.6, http://zenodo.org/record/83515
Tepuidessus grulai sp. nov. from Acopán Tepui in Venezuela (Coleoptera: Dytiscidae: Hydroporinae: Bidessini)
Kodada, Jan, Hendrich, Lars, Balke, Michael (2018): Tepuidessus grulai sp. nov. from Acopán Tepui in Venezuela (Coleoptera: Dytiscidae: Hydroporinae: Bidessini). Zootaxa 4434 (3): 561-572, DOI: 10.11646/zootaxa.4434.3.1
FIGURE 9 in Tepuidessus grulai sp. nov. from Acopán Tepui in Venezuela (Coleoptera: Dytiscidae: Hydroporinae: Bidessini)
FIGURE 9. Habitat of Tepuidessus breweri. Shallow puddles on rocks of summit plateau of Roraima Tepui (Fotos: M. Hornburg).Published as part of Kodada, Jan, Hendrich, Lars & Balke, Michael, 2018, Tepuidessus grulai sp. nov. from Acopán Tepui in Venezuela (Coleoptera: Dytiscidae: Hydroporinae: Bidessini), pp. 561-572 in Zootaxa 4434 (3) on page 570, DOI: 10.11646/zootaxa.4434.3.10, http://zenodo.org/record/129224
Tepuidessus grulai Kodada & Hendrich & Balke 2018, sp. nov.
Tepuidessus grulai sp. nov. Type locality. Acopán Tepui, ca. 2,000 m, 5.194N 62.045W, Venezuela. Holotype, male (ZSM): " Venezuela: Estado Boliver, puddles on rocks in Acopantepui base camp" / " 2022m, 14.xi.2015, 5.19413°[N] -62.04478°[W], (VEN 6 /Nov2015)" / " Holotype Tepuidessus grulai sp. nov., Kodada, Hendrich & Balke des. 2018" [red printed label]. Paratypes: 43 exs with the same label data as holotype (CHF, CKB, MIZA, NMPC, ZSM). Each paratype is provided with a red printed paratype label. Description of holotype. Habitus slightly broad oval, outline with particularly distinct discontinuity between pronotum and elytra (Figs 3–4). Pronotum broadest before its midlength. Elytra widest at about midlength. Total length: 1.8 mm; maximum width: 0.9 mm. Colouration. Black dorsally, dark brown to black ventrally (Figs 3, 4, 5). Surface sculpture. Head with distinct microreticulation and few setiferous punctures (Fig. 4A). Pronotum and elytron shiny (Fig. 3A), mostly polished and distinct microreticulation only visible along pronotal margins; whole surface with dense and coarse setiferous punctation, exceptionally coarse and deep on disc (Fig. 6C). Ventral surface microreticulate, abdominal ventrites 3–6 shinier; venter with distinct setiferous punctation; posterior margins of ventrites 3–5 without denticles (present in Neotropical genus Bidessodes Régimbart, 1895; see Miller & Bergsten 2016). Structures. Antenna stout, moniliform. Head without occipital line and with rounded clypeus (as in Fig. 4). Pronotum without lateral bead; with distinct and very deep basal striae (as in Figs 6A, C). Elytron with short and deep basal striae, but without sutural line, with distinct longitudinal depression on elytral disc roughly in middle of each elytron (as in Fig. 3B, arrow). Basal epipleural transverse carina absent. Metathoracic wings vestigial, reduced to short membranous vestiges without any sign of venation (Fig. 6). Pro- and mesotarsi appearing stout because tarsomeres 1–3 distinctly dilated laterally; metatrochanter moderately offset. Male genitalia. Median lobe of aedeagus simply curved, long in lateral view; in ventral view, slender and gently narrowed towards tip (Figs 7A, B); lateral lobes (parameres) bisegmented and of a general Bidessini type (see Biström 1988), tip of distal joint rounded (Fig. 7A) and with no obvious "nose" or hook (as in e.g. Balke et al. 2015: fig. 31; Biström 1988: fig. 15). Female. Dorsal and ventral surface dull due to well impressed microreticulation between surface punctation (Figs 3B, 4B, 5). Apical ventrite flattened before posterior margin, tip in lateral view beak-shaped (Fig. 6B). Size variation. Total length: 1.7–1.8 mm; maximum width: 0.7–0.9 mm. Differential diagnosis. Using the key of Miller & Bergsten (2016: 220), the new species from Acopán Tepui does not key out with Tepuidessus, but with Papuadessus Balke, 2001, a genus endemic to the Papuan region. The two Papuadessus species are morphologically rather divergent and considered members of one lineage only based on DNA sequence data (Balke 2001; Balke et al. 2013; Miller & Bergsten 2016). In the absence of any obvious morphological apomorphy that would group the species treated here with Papuadessus, or indeed any other genus, we opt for the pragmatic approach to assign it to Tepuidessus. Tepuidessus grulai sp. nov. differs from T. breweri by (1) much broader habitus with distinct outline discontinuity between pronotum and elytra, (2) well developed, long and deep basal striae on pronotum and elytra and (3) by unbordered apical abdominal ventrite. The median lobe of aedeagus is more narrowed than in T. breweri in lateral view. Etymology. Named after Daniel Gruľa, one of the collectors of the new species. It is a noun in the genitive singular. Distribution. Only known from the type locality (Fig. 8). This locality is situated around 140 km west of Mount Roraima, the locality of the other Bidessini species described from the plateau of a tepui. Habitat. Puddles on rocks of summit plateau of Acopán Tepui (Fig. 10).Published as part of Kodada, Jan, Hendrich, Lars & Balke, Michael, 2018, Tepuidessus grulai sp. nov. from Acopán Tepui in Venezuela (Coleoptera: Dytiscidae: Hydroporinae: Bidessini), pp. 561-572 in Zootaxa 4434 (3) on pages 563-566, DOI: 10.11646/zootaxa.4434.3.10, http://zenodo.org/record/129224
Ancyronyx lianlabangorum sp. nov., a new spider riffle beetle from Sarawak, and new distribution records for A. pulcherrimus Kodada, Jäch & Čiampor based on DNA barcodes (Coleoptera, Elmidae)
Ancyronyx lianlabangorum sp. nov. (Coleoptera, Elmidae), a new spider riffle beetle from the Kelabit Highlands (Sarawak, northern Borneo), is described. Illustrations of the habitus and diagnostic characters of the new species and the similar, polymorphic A. pulcherrimus Kodada et al. are presented. Differences to closely related species, based on COI nucleotide sequences and morphological characters, are discussed. Ancyronyx pulcherrimus is here recorded from Sarawak for the first time, based on DNA barcoding
Tepuidessus Spangler 1981
Tepuidessus Spangler, 1981 Type species. Tepuidessus breweri Spangler, 1981, by original designation. Revised diagnosis for Tepuidessus . Black Bidessini with dorsal surface densely setose; clypeus not modified; occipital line absent; basal striae on pronotum and elytra absent or present; sutural line on elytron absent; longitudinal elytral carina on disc absent; basal epipleural transverse carina absent; metathoracic wings reduced to short membranous vestiges without any sign of venation (Fig. 6); median lobe simple, with narrowed apical part in ventral view; parameres two-segmented. Species inhabiting tepui plateaus in Venezuela. Note that the presence/absence of pronotal and/or elytral striae has also been shown to possibly vary even within closely related species of other genera, e.g. Limbodessus Guignot, 1939 (see Balke et al. 2015).Published as part of Kodada, Jan, Hendrich, Lars & Balke, Michael, 2018, Tepuidessus grulai sp. nov. from Acopán Tepui in Venezuela (Coleoptera: Dytiscidae: Hydroporinae: Bidessini), pp. 561-572 in Zootaxa 4434 (3) on page 562, DOI: 10.11646/zootaxa.4434.3.10, http://zenodo.org/record/129224
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