1,722,307 research outputs found
Pseudagrion sarepi Kipping & Dijkstra, sp. nov.
No full textPseudagrion sarepi Kipping & Dijkstra sp. nov. – Sarep Sprite (Type Photo 22, Photos 33–34, Fig. 13) Taxonomy Genetically, morphologically, ecologically and geographically close to P.fisheri Pinhey, 1961 and P. greeni Pinhey, 1961; combines characters of both, but genetically distinct from them. Material studied Holotype ♂ . Angola, Cuando-Cubango Province, Longa River, 82 km E of Menongue at road to Cuito Cuanavale, clear and sandy river in wide open valley (Photo 34), 1326 m a.s.l. (14.6063 ° S 18.4673 ° E), 20 -v- 2012, leg. J. Kipping, RMNH. Further material. ANGOLA (Cuando-Cubango Province): 1 ♂ 1 ♀, as holotype, RMNH. 4 ♂ 1 ♀, as holotype, CJKL. 1 ♂ (RMNH.INS.507931), Cuito River, 9 km N of Cuito Cuanavale, large, clear and sandy river, 1187 m a.s.l. (15.0872 ° S 19.1497 ° E), 19 -v- 2012, leg. J. Kipping, RMNH. 4 ♂ 6 ♀, Luassingua River, 55 km E of Menongue at road to Cuito Cuanavale, clear and sandy river in wide open valley, 1326 m a.s.l. (14.5899 ° S 18.1705 ° E), 20 -v- - 2012, leg. J. Kipping, CJKL. 2 ♂ (RMNH.INS.507940, RMNH.INS.507941), same locality and date, leg. J. Kipping, RMNH. Genetics Two unique haplotypes (n= 3) nearest to four of P. greeni (n= 8) and two of P. fisheri (n = 2). Male morphological diagnosis Similar to P. fisheri and P. greeni by (a) the uniformly orange labrum; (b)the inner border of the cerci lacking a basal tooth or toothed flange in dorsal view; (c) the upper branch of cerci being longer than the lower; and (e) the paraprocts that are more than half as long as the cerci. However, (1) is distinctly smaller, Hw 18.0–21.0 mm (mean 19.2 mm; n = 10), compared to both P. fisheri (21.0– 23.5 mm; mean 22.3 mm; n= 14) and P. greeni (21.0–23.0 mm; mean 22.0 mm; n= 22); (2) with maturity the dorsa of the head and thorax are wholly dark, rather than with tiny postocular spots and continuous though narrow ante-humeral stripes as in both other species; (3) S 8–9 are dorsally wholly blue like in P. greeni, not entirely dark like all other segments as in P. fisheri; and (4) the upper branch of the cerci is small and in lateral view reaches about as far as the rather deep lower branch as in P. fisheri, rather than being huge and reaching far beyond the small lower branch as in P. greeni (Fig. 13). Etymology Named after the SAREP (Southern African Regional Environmental Program) field survey during which the species was discovered (neuter singular genitive noun). Range and ecology Like P. greeni and P. fisheri prefers clear, sandy and swift rivers in open landscapes. Perches low on overhanging grass or floating vegetation. Known from three sites between about 1 150 and 1 350 m a.s.l. in grassy valleys of the Bié highlands between Menongue and Cuito Cuanavale in southern Angola, but probably more widespread.Published as part of Dijkstra, Klaas-Douwe B., Kipping, Jens & Mézière, Nicolas, 2015, Sixty new dragonfly and damselfly species from Africa (Odonata), pp. 447-678 in Odonatologica 44 (4) on pages 532-534, DOI: 10.5281/zenodo.3538
Ceriagrion banditum Kipping & Dijkstra, sp. nov.
No full textCeriagrion banditum Kipping & Dijkstra sp. nov. – Band-eyed Citril (Type Photo 15, Photo 21–22, Fig. 10) Taxonomy Dijkstra (2005 a) reviewed the suave -complex of Ceriagrion and suggest- ed that various taxa may be synonymous. However, up to four superficially similar taxa that differ clearly genetically and subtly in ecology, coloration and morphology overlap in southern central Africa (Tree 2). Additional taxa may occur elsewhere: western African specimens appear similar to C.suave Ris, 1921 but are slightly distinct morphologically (Fig. 10) and genetically (Tree 2). These could represent C. moorei Longfield, 1952 of which C. hamoni Fraser, 1955 is possibly a synonym. Marshall & Gambles (1977) separated C. suave and C. moorei from northern Ghana by the penis shape, but while this suggests that similar taxa overlap there too, whether their interpretation agrees with type material must be investigated. Dijkstra & Clausnitzer (2014) also provided insufficient information to distinguish the various taxa in the complex, which they separated from C. ignitum Campion, 1914 and C. kordofanicum Ris, 1924 only by (a) the moderate size, Hw 17.0–23.0 mm rather than 15.0–17.0 mm; although (b) the greenish rather than deep reddish eyes with maturity are also diagnostic. All species further have (c) rather dull orange to brown dorsa of the head and thorax; (d) uniformly orange to red abdomen; (e) apically pointed penis with acute or finger-like lateral lobes; (f) no toothed processes on the apical border of S 10; and (g) paraprocts that do not or barely reach beyond the cerci and that have a fairly distinct angular heel ventrally. True C. suave favours temporary pools such as rain puddles and, as confirmed with the holotype and a paratype in MRAC, is (1) notably uniform in colour, brown to orange on the head and thorax grading to yellow on the face and flanks, and typically without a darker greyish or greenish tinge to the head dorsum, nor with distinctly whitish thoracic flanks and/or blackish dots on the humeral and metapleural sutures; (2) often has largely yellow-stained wings; (3) the penis does not have drawn-out finger-like lateral lobes; (4) the dorsal excision of the apical border of S 10 is up to half as deep as the segment is dorsally long, and its borders are pale; (5) the cerci have the apical black teeth turned downward and thus usually not visible in dorsal view; and (6) the tips of the paraprocts reach at most as far as those (or even the apical teeth) of the cerci (Fig. 10). The suave -complex species’ similarity stems from the shared lack of conspicuous characters rather than from common ancestry (Tree 2): ecology, genetics, coloration and appendages and penis shape indicate that C. suave is very close to the ubiquitous C. glabrum (Burmeister, 1839), while two new species described here are nearer C. bakeri Fraser, 1941 but lack its bright blue head and thorax. Known only from the coastal plain of Mozambique and Tanzania, the similarly dull C. mourae Pinhey, 1969 differs by having 10 rather than 12–14 Fw postnodal cross-veins, the rectangular rath- er than trapezoidal Pt, and in the shape of the male appendages (Fig. 10). Material studied Holotype ♂ . RMNH.INS.508068, Zambia, Northern Province, Lake Chila near Mbala, boggy eastern lake shore, at shallow grassy pools, 1619 m a.s.l. (8.8341 ° S 31.3948 ° E), 08-xii- 2014, leg. J. Kipping, RMNH (Photo 21). Further material. MALAWI (Central Region): 2 ♂, 45 km SW of Lilongwe, Lilongwe District, Dzalanyama Forest Reserve, SE of Dzalanyama Forest Lodge, miombo woodland with rocky areas and marshy grassland, 1250 m a.s.l. (14.26 ° S 33.455 ° E), 27–29 -xii- 2001, leg. K.- D.B. Dijkstra, RMNH. 1 ♂ (identification suspected), MALAWI (Southern Region): Blantyre District, Blantyre, Sunnyside, Smythe Road, rough grass in suburban garden, 1000m a.s.l. (15.7922 ° S 34.9943 ° E), 06-xi- 2001, leg. K.- D.B. Dijkstra, RMNH. MO- ZAMBIQUE (Zambezia Province): 1 ♂, 15 km NE of Gurue, Namuli Massif, Muretha Plateau, grassy plateau with streams, boggy pits and forest patches, 1860 –1890 m a.s.l. (15.3883 ° S 37.0467 ° E), 05-xii- 2001, leg. K.- D.B. Dijkstra, RMNH. ZAMBIA (Copperbelt Province): 1 ♂ (RMNH.INS.508067), Chimfunshi Wildlife Reserve near Chingola, shallow pool in grassy dambo near gallery forest, 1296 m a.s.l. (12.3676 ° S 27.4831 ° E), 20 -xi- 2014, leg. J. Kipping, RMNH. 3 ♂ 3 ♀, same locality, same dates, CJKL. ZAMBIA (Northern Province): 1 ♂ (RMNH.INS.508066), Shiwa N’gandu, Kapishya Hot Springs, shallow grassy pools near Mansha River (Photo 22), 1434 m a.s.l. (11.1703 ° S 31.6057 ° E), 04-xii- 2014, leg. J. Kipping, RMNH. 1 ♂ 1 ♀, same locality, 06-xii- 2014, leg. J. Kipping, RMNH. 1 ♂ 1 ♀ same locality, 04-xii- - 2014, leg. J. Kipping, BMNH. 1 ♂ 1 ♀ same locality, 04-xii- 2014, leg. J. Kipping, CJKL. ZAMBIA (Northwestern Province): 1 ♂ (RMNH.INS.507960), Ikelenge, Sakeji School on Hillwood Farm, grassy ditches at forest edge 1 405 m a.s.l. (11.2332 ° S 24.3125 ° E), 25 -ii- 2010, leg. J. Kipping, CJKL. 1 ♀ (RMNH.INS.508069), as holotype, RMNH. 2 ♂ (RMNH.INS.508070), 1 ♀, locality as holotype, 11 -xii- 2014, leg. J. Kipping, CJKL. Genetics Three unique haplotypes (n= 6) nearest to C. bakeri and C. junceum sp. nov. (Tree 2). Male morphological diagnosis Distinct within the suave -complex (see above) by its (1) greater size, Hw 21.5–22.5 mm (n = 13) rather than 17.0–21.0 mm; (2) sleeker build with the wing tips reaching at most a third down the length of S 6, rather than halfway or beyond, and Hw 50–51 % of full length (n= 8), versus 51–57 % in C.suave (n= 9); (3) two narrow horizontal dark bands on the greenish eyes, which are typically lost with maturity in other species; (4) rather intermediate colour of head and thorax, i.e., neither quite uniformly orange like C.suave, nor more two-toned brown and cream with dark dots on the sutures as C.sakejii Pinhey, 1963 and C. junceum; (5) often distinctly reddish rather than pale brown Pt; (6) penis with finger-like lateral lobes, rather like C. bakeri (Fig. 10); (7) apical excision on S 10 being about a third as deep as the segment and bordered with tiny black denticles that appear as dark ridges; (8) cerci that in dorsal view appear narrowed and twisted distally, with the apical black tooth turned in- and base-wards, from above thus being well visible and lying almost halfway the cerci; and (9) tips of the paraprocts that typically fall clearly short of those of the cerci (Fig. 10). Etymology Medieval Latin “banished”, the origin of the word bandit, which refers to the banded eyes recalling the mask of a bandit (neuter adjective). Range and ecology Known from northern Zambia to northern Mozambique between 1 000 and 1 900 m a.s.l. While poorly preserved, males in MRAC from Lubumbashi in southern Congo-Kinshasa also appear to belong to this species (Map 4). Exact habitat difference with other species unclear, but occurs at shallow grassy pools in dambos, open depressions that flood seasonally (see C. junceum). Also found away from water in half-open miombo woodland.Published as part of Dijkstra, Klaas-Douwe B., Kipping, Jens & Mézière, Nicolas, 2015, Sixty new dragonfly and damselfly species from Africa (Odonata), pp. 447-678 in Odonatologica 44 (4) on pages 504-510, DOI: 10.5281/zenodo.3538
Zygonyx denticulatus Dijkstra & Kipping, sp. nov.
No full textZygonyx denticulatus Dijkstra & Kipping sp. nov. – Pale Cascader (Type Photo 59, Photos 47, 76– 77, Fig. 35) Taxonomy Morphologically distinct from Z. flavicosta (see Z. annika sp. nov.) and genetically closer to western African specimens of that species than those found within 40 km of it around Upemba National Park, Katanga (Tree 10). Treated as a distinct species by Dijkstra & Clausnitzer (2014), but the photograph on page 218 shows this species, and not Z. flavicosta (Map 12). Material studied Holotype ♂ . RMNH.INS.505335, Congo-Kinshasa, Katanga, Kiubo, Lufira (large, Kiubo Falls) and Luvilombo (small, rocky) rivers, and adjacent flooded areas, gallery forest and degraded miombo woodland, 840–880 m a.s.l. (9.52 ° S 27.05 ° E), 05-xi- 2011, leg. K.-D.B. Dijkstra, RMNH. Further material. ANGOLA (Huambo Province): 2 ♂ (RMNH.INS.507908), Keve (Cuvo) River near Alto Hama, 1415 m a.s.l. (12.2031 ° S 15.5952 ° E), 14 - v- 2012, leg. J. Kipping, CJKL. 1 ♂ (RMNH.INS.507907), same locality and date, RMNH. CONGO-KINSHASA (Katanga): 6 ♂ (RMNH.INS.505334, RMNH.INS.505338, RMNH.INS.505357), 1 ♀ (RMNH.INS.505336), as holotype, RMNH. 2 ♂ 1 ♀ (RMNH.INS.505657), Kundelungu National Park, Masansa, small river with some gallery forest and falls in miombo woodland, 1440 – 1480 m a.s.l. (10.532 ° S 27.978 ° E), 22 -xi- 2011, leg. K.-D.B. Dijkstra, RMNH. 1 ♂, Kundelungu National Park, Lutshipuka, small river with some gallery forest and falls in miombo woodland (Photo 47), 1500 – 1530 m a.s.l. (10.557 ° S 27.958 ° E), 26 -xi- 2011, leg. K.-D.B. Dijkstra, RMNH. ZAMBIA (Luapula Province): 1 ♂ (RMNH.INS.508074), 1 ♂ ♀, Mumbuluma Falls, 35 km NW of Mansa, large waterfall, 1195 m a.s.l. (10.9301 ° S 28.7354 ° E), 05-iv- 2011, leg. J. Kipping, CJKL. 1 ♂, Ntumbachushi Falls, 21 km SE of Mbereshi, large waterfall of Ngona River (Photo 77), 1154 m a.s.l. (9.8537 ° S 28.9448 ° E), 07-iv- 2011, leg. J. Kipping, RMNH. 2 ♂, same locality, same dates, BMNH. 5 ♂ 1 ♀, same locality, same dates, CJKL. ZAMBIA (Northern Province): 1 ♂ (RMNH.INS.508071), Shiwa N’gandu, Kapishya Hot Springs, Mansha River, fast flowing rapids, 1434 m a.s.l. (11.1702 ° S 31.6053 ° E), 06-xii- 2014, leg. J. Kipping, RMNH. 1 ♂, same locality, same dates, CJKL. 1 ♂ ♀ (♂ RMNH.INS.508072, ♀ RMNH.INS.508073), Chishimba Falls W of Kasama, Kaela Fall, large waterfall, 1315 m a.s.l. (10.1089 ° S 30.9167 ° E), 07- xii- 2014, leg. J. Kipping, RMNH. Genetics Nine unique haplotypes (n = 12) nearest to Z. flavicosta from western Africa, but more distant to the sympatric clade of Z. flavicosta (Tree 10). Male morphological diagnosis Belongs to flavicosta -complex by (a) the modest size, Hw 35.0–39.0 mm (n = 24); (b) the moderately dense venation, with 10 ½– 12 ½ Ax in Fw; (c)the largely black hind legs; (d) the pointed and forward-curved apex of the anterior lamina (Fig. 35); and (e) the pale subapical band on the dor- sum of S 7 that stands out against largely dark S 5–6. Differs from sympatric Z. flavicosta by (1) the dorsal dark metallic area that does not extend to the anterior side of the frons; (2) the pale brown and not blackish Pt (Photo 76); and (3)the presence of denticles on the apical border of the genital lobe (Fig. 35). Also, (4) the labrum is often pale brown at least at its base, although it can be wholly black as in Z. flavicosta; (5) the background colour of the thorax is browner and the pale stripes are often more fragmented and reduced, e.g., the stripe on the metepisternum is typically broken between the metastigma and wing base, or may be reduced to a single spot in that position, while the metepimeron may bear only one to three spots rather than a T-shaped stripe; and (6) the hook of the hamule often seems longer and more strongly curved, about as long as the width of its base, rather than about half (Fig. 35). Etymology Latin “with small teeth” for the denticles on the genital lobe (masculine adjective). Range and ecology Fast-flowing sections and waterfalls of large streams and small rivers in gallery forest and open miombo woodland in Katanga, eastern Angola and north-eastern Zambia, recorded between 840 and 1 530 m a.s.l.Published as part of Dijkstra, Klaas-Douwe B., Kipping, Jens & Mézière, Nicolas, 2015, Sixty new dragonfly and damselfly species from Africa (Odonata), pp. 447-678 in Odonatologica 44 (4) on pages 665-669, DOI: 10.5281/zenodo.3538
Pseudagrion tanganyicum Dijkstra & Kipping, sp. nov.
No full textPseudagrion tanganyicum Dijkstra & Kipping sp. nov. – Tanganyika Sprite (Type Photo 23, Photos 35–36, Fig. 12) Taxonomy Structurally probably indistinguishable from P. massaicum Sjöstedt, 1909, but coloration and ecology of this potentially sympatric species (Map 5) differ strongly, and the two were treated as distinct by Dijkstra & Clausnitzer (2014). The two are also inseparable by COI sequence, but this is notably frequent in the Pseudagrion B-group, including the well-differentiated pairs P. acaciae and P. vaalense, P. camerunense (Karsch, 1899) and P. glaucum (Sjöstedt, 1900), P. coeleste Longfield, 1947 and P. nubicum Selys, 1876, and P. commoniae (Förster, 1902) and P. hamoni Fraser, 1955. Material studied Holotype ♂ . Tanzania, Kigoma Region, Kibirizi, Lake Tanganyika at Nondwa point, sandy and reedy lakeshore, 770 m a.s.l. (4.86 ° S 29.61 ° E), 20 -viii- - 2009, leg. K.-D.B. Dijkstra, RMNH. Further material. TANZANIA (Kigoma Region): 11 ♂ (RMNH.INS.502706, RMNH.INS.502707, RMNH.INS.502715), as holotype, RMNH. 8 ♂ (RMNH.INS.502811, RMNH.INS.502816, RMNH.INS.502820), 6 km SW of Ilagala, Lower Malagarasi Basin, Lake Tanganyika at mouth of Malagarasi, reedy and sandy lakeshore, 770 m a.s.l. (5.3256 ° S 29.8021 ° E), 15 -viii- 2009, leg. K.-D.B. Dijkstra, RMNH. ZAMBIA (Northern Province): 2 ♂ (RMNH.INS.508095), Lake Tanganyika, Isanga Bay Lodge, 15 km NE of Mpulungu, rocky and sandy lake shore (Photo 36), 777 m a.s.l. (8.6549 ° S 31.1932 ° E), 09-xii- 2014, leg. J. Kipping, RMNH. 5 ♂, same locality, same dates, CJKL. 13 ♂ 1 ♀, same locality, 10 -xii- 2014, leg. J. Kipping, CJKL. Genetics Six haplotypes (n= 7), of which one shared with P. massaicum (n = 8; three additional haplotypes); all nine haplotypes are very similar and mixed. Male morphological diagnosis Structurally like P. massaicum with (a) the fairly small size, Hw 17.0– 19.0mm (n = 24); (b) the pointed lateral and apical lobes of the penis; (c)the similarly long cerci and paraprocts; and (c) the paraprocts with a short blunt apex and a prominent dorsal knob in lateral view (Fig. 12). However, with maturity (1) the face, eyes and postocular spots are yellow rather than red; (2) the ante-humeral stripes are green rather than red; and (3) the thorax is quite uniformly pale blue below the humeral suture rather than red grading to bluish pruinose below the metapleural suture. Etymology The name is an amalgam of “Tanganyika” and the specific epithet of P. massaicum (neuter adjective). Range and ecology Inhabits sandy and rocky beaches and wave-battered reeds in shallows of Lake Tanganyika (elevation around 770 m a.s.l.), i.e., notably different from the marshy fringes of pools and streams favoured by P. massaicum. Known sites are over 400 km apart, suggesting the species occurs all around the lake, just like the similarly endemic Platycypha pinheyi Fraser, 1950 that prefers rocky shores.Published as part of Dijkstra, Klaas-Douwe B., Kipping, Jens & Mézière, Nicolas, 2015, Sixty new dragonfly and damselfly species from Africa (Odonata), pp. 447-678 in Odonatologica 44 (4) on pages 535-537, DOI: 10.5281/zenodo.3538
Chlorocypha aurora Dijkstra, Kipping & Schutte, sp. nov.
No full textChlorocypha aurora Dijkstra, Kipping & Schütte sp. nov. – Dawn Jewel (Type Photo 3, Photo 7, Fig. 2) Taxonomy Genetically falls within a poorly resolved group of Chlorocypha Fraser, 1928 species that predominates on more open waters, such as large rivers and woodland streams, and includes widespread species like C. curta (Hagen in Selys, 1853), C. fabamacula Pinhey, 1961, C. pyriformosa Fraser, 1947 and C.victoriae (Förster, 1914). The present taxon is distinctly marked and therefore, within a genus where structural differences are almost non-existent, worthy of specific recognition. Material studied Holotype ♂ . Cameroon, South Province, Kienké River at Kribi, large river in degraded rainforest, 10–15 m a.s.l. (2.9347 ° N 9.9134 ° E), 10 -vi- 2008, leg. K.- D.B. Dijkstra, J. Kipping & K. Schütte, RMNH. Further material. CAMEROON (South Province): 3 ♂ (RMNH.INS.229137, RMNH.INS.500154), 2 ♀, as holotype, RMNH. 3 ♂, as holotype, CJKL. Genetics One unique haplotype (n = 2) that is close to those of the complex of species mentioned above. Male morphological diagnosis Near the Upper Guinean C. luminosa (Karsch, 1893) and potentially sympatric C. neptunus (Sjöstedt, 1900) and C. pyriformosa Fraser, 1947 by (a)small size, Hw 18.5–19.5 mm (n = 7); and (b) fairly extensive black abdominal markings (Fig. 2). However, is the only Chlorocypha species to combine this with (1) a contrastingly yellow-marked head and thorax; (2)yellow-stained wings, especially at their bases; (3) entirely black tibiae with some light pruinosity, rather than with yellow or white anterior streaks; and (4) a reddish orange abdomen with the less extensive black markings on S 2 (almost) continuous between the basal and apical border, S 3–5 and sometimes S 6 each with an apical black bar, and yellow lateral carinae of S 4–8 (Fig. 2). The colour of C. luminosa has a deeper yellow tone and is more uniform across the body and wings, C. neptunus has an especially extensively black abdomen, and C. pyriformosa becomes almost red on the abdomen and black on the thorax. Etymology Latin “dawn” refers to the orange coloration (noun in apposition) and the first author’s colleagues of DAWN; the Damselfly Workers at Naturalis (see Dijkstra et al. 2014). Range and ecology Near sea level on the Kienké, a large forested river at Kribi in southern Cameroon.Published as part of Dijkstra, Klaas-Douwe B., Kipping, Jens & Mézière, Nicolas, 2015, Sixty new dragonfly and damselfly species from Africa (Odonata), pp. 447-678 in Odonatologica 44 (4) on pages 467-469, DOI: 10.5281/zenodo.3538
Anax gladiator Dijkstra & Kipping, sp. nov.
No full textAnax gladiator Dijkstra & Kipping sp. nov. – Swordbearer Emperor (Type Photo 24, Photos 31, 37– 38, Fig. 14) Taxonomy Belongs to the genetically, morphologically and ecologically distinct speratus -group of Anax Leach, 1815 (see below). Overlaps geographically with A. speratus Hagen, 1867 that is most similar in coloration, but probably also with A. congoliath Fraser, 1953 that is closer structurally and genetically (Tree 3) (Map 6). However, is highly distinct from both, and treated as such by Dijkstra & Clausnitzer (2014). A fourth species in this group has long remained in synonymy with A. speratus but replaces it from the Cape Verde Islands to Nigeria and possibly further east: A. rutherfordi McLachlan, 1883 is distinct genetically (Tree 3) and has (1) smaller size on average, Hw 49.0– 54.0 mm (n= 5) rather than 52.0–57.0 mm (n = 10); (2) the apices of the mid and hind femora and anterio-exterior side of the front femora black, rather than all femora all red; (3) the apical borders of S 2–6 without lateral black dots; (4) the basal half of the dorsal ridge on S 10 distinctly raised, rather than low and sloping to apex; and (5) the epiproct 60 % as long as the cerci, rather than 50 % (Fig. 14) [rejected synonymy]. Material studied Holotype ♂ : RMNH.INS.505566, Congo-Kinshasa, Katanga, Upemba National Park, Kabwekanono, source area of Kafwi, stream through gallery forest and bogs in open grassy plains, 1770 – 1820 m a.s.l. (8.937 ° S27.166 ° E), 16 -xi- 2011, leg. K.-D.B. Dijkstra, RMNH. Further material: CONGO-KINSHASA (Katanga): 1 ♂ (RMNH.INS.505567), as holotype, RMNH. 1 ♂ (RMNH.INS.505596), Upemba National Park, Munte, river with grassy verges, gallery patches and rapids in miombo woodland, 1450 m a.s.l. (8.77 ° S26.84 ° E), 18 -xi- 2011, leg. K.-D.B. Dijkstra, RMNH. 3 ♂, Lubumbashi, xi- 1952, leg. C. Seydel, MRAC. 1 ♂, Karavia (Elisabethville), ix- 1951, leg. C. Seydel, MRAC. ZAMBIA (Northern Province): 1 ♂ (RMNH.INS.508082), Shiwa N’gandu, Kapishya Hot Springs, Mansha River, calm section of clear blackwater river, 1434 m a.s.l. (11.1702 ° S31.6053 ° E), 13 -xii- 2014, leg. J. Kipping, RMNH. 1 ♂ (Photo 37), same locality, 04-xii- 2014, CJKL. 1 ♂, Shiwa N’gandu, Kapishya Hot Springs, small clear stream and pools W of Kapishya, 1456 m a.s.l. (11.1939 ° S31.5617 ° E), 06-xii- 2014, leg. J. Kipping, CJKL. 1 ♀, Shiwa N’gandu, Kapishya Hot Springs, ovipositing at small sandy and warm (36 ° C) stream flowing off hot springs to the nearby Mansha River, 1434 m a.s.l. (11.1706 ° S31.6013 ° E), 04-xii- 2014, leg. J. Kipping, CJKL. ZAMBIA (Northwestern Province): 1 ♂ (RMNH.INS.508081), Sakeji River S of Sakeji Mission School, E of Ikelenge, calm section of clear blackwater river, 1385 m a.s.l. (11.2355 ° S24.3100° E), 25 -xi- 2014, leg. J. Kipping, RMNH. 1 ♂, same locality, same dates, CJKL. Genetics Five unique haplotypes (n= 5) nearest to A. congoliath (Tree 3). Male morphological diagnosis Belongs to the speratus -group on account of (a) the unmarked frons; (b)the reddish femora contrasting with the black tibiae; (c) the presence of an accessory lamina on S 10, a small process visible between the cerci in dorsal view; (d) the cerci with broadened and rounded apices; and (e) the epiproct that is at least half as long as the cerci (Fig. 14). Most recalls the sympatric A.speratus by (f) the entirely orange to red abdomen. However, is (1)small- er on average, Hw 49.0–54.0 mm (n= 12) rather than 52.0–57.0 mm (n = 11); (2) with maturity the frons and thorax remain green, rather than becoming orange to reddish; (3) the apical borders of S 2–6 lack lateral black dots; (4)S 10 has its dorsal ridge raised into a high triangular spike, rather than low and sloping down to apex; and (5) the epiproct is well over half as long as the cerci, rather than about half (Fig. 14). Etymology Latin “sword-bearer” refers to the blade-like spike at the abdomen tip (noun in apposition). Range and ecology As with all speratus -group species inhabits streams and small rivers, rather than standing waters as other African Anax. These are nutrient-poor and slightly acidic with peat moss and/or blackwater, and typically open with some gallery or miombo forest nearby (Photo 38), on sandy plateaus between 1 385 and 1 820 m a.s.l. in Katanga and northern Zambia. Flow can be fast but there are always calm sections, e.g., it was seen in Zambia along a lake below its inflow. Ovipositing females without guarding males were observed on a stream flowing from hot springs and in small pools beside a river. There is an unconfirmed record near the Nyika Plateau of northern Malawi (Allen Barlow pers. comm.) and a possible sighting in suitable habitat near Negage in northern Angola (obs. K.-D.B. Dijkstra) suggesting the species is widespread across the sandy plateaus of southern central Africa.Published as part of Dijkstra, Klaas-Douwe B., Kipping, Jens & Mézière, Nicolas, 2015, Sixty new dragonfly and damselfly species from Africa (Odonata), pp. 447-678 in Odonatologica 44 (4) on pages 539-544, DOI: 10.5281/zenodo.3538
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Data files for “Evidence for a Large Exomoon Orbiting Kepler-1625b” by Teachey & Kipping (2018)
No full textHubble WFC3 observations of Kepler-1625 taken in October 2017 were reported in Teachey & Kipping (2018). In that work, we the authors find evidence for an exomoon associated with the planetary transit of Kepler-1625b. We here provide the processed frame images, light curves, centroid time series, stellar isochrones and model posteriors/results data used in our study. Each folder contains a short readme file aimed at professional astronomers familiar with these types of analyses
Eleuthemis libera Dijkstra & Kipping, sp. nov.
No full textEleuthemis libera Dijkstra & Kipping sp. nov. – Free Firebelly (Type Photo 45, Fig. 28) Taxonomy The only Eleuthemis species that is well-separated from all others by its secondary genitalia, which together with its highly distinct COI haplotypes suggests it is the sister-species of all remaining species. It is potentially sympatric with congeners, being found in Angola 31 km from E. eogaster sp. nov. and in north-western Zambia 18 km from a species that morphologically is nearest to E. buettikoferi but genetically distinct; see E. eogaster (Tree 7). Material studied Holotype ♂ . RMNH.INS.508918, Angola, Uíge Province, 13 km N of Negage, Lucunga river near Hinda, gravel river in open swamp and forest gallery, 1200 m a.s.l. (7.6462 ° S 15.2598 ° E), 24 -xi- 2012, leg. K.-D.B. Dijkstra, RMNH. Further material. ANGOLA (Uíge Province): 3 ♂ (RMNH.INS.508376), as holotype, RMNH. 4 larvae (RMNH.INS.559575, RMNH.INS.559576, RMNH.INS.559577, RMNH.INS.559578), locality as holotype, 04-x- 2013, leg. K.-D.B. Dijkstra, RMNH. ZAMBIA (Northwestern Province): 1 ♂ (RMNH.INS.507955), Luakera River, 20 km N of Mwinilunga, fast flowing rocky and sandy river with several cascades in gallery forest, 1380 m a.s.l. (11.5271 ° S 24.4110 ° E), 25 -ii- 2010, leg. J. Kipping, CJKL. Genetics Four unique haplotypes (n= 7) are the most distinct within the genus (Tree 7). Male morphological diagnosis Superficially similar to E. buettikoferi by the abdomen being (a) entirely whitish pruinose on the upperside with maturity; and (b) largely orange on the underside. Differs from other Eleuthemis species by (1) the pale labrum with at most a hint of a black border; (2) the pale frons darkened basally at least anteriorly to the lateral ocelli; (3) the uniformly brown occipital triangle; (4) the dark metapleural stripe expanded markedly onto the metepisternum between the metastigma and wing bases, rather than being of fairly even width throughout; (5) the Fw discoidal field with 2 to 5 single cells, rather than entirely of two or more rows of cells, although rarely up to 4 are single in others; (6) the largely pale brown rather than blackish secondary genitalia; (7) the relatively large and less divergent posterior branch of the genital lobe; and (8) the hook of the hamule with a right-angled rather than acute tip. Also notable are (9) the modest size, Hw 23.0–24.0 mm (n = 4) rather than 23.5–26.5 mm (n= 10); and (10) the dark tips of Fw that extend almost halfway along Pt, rather than just touching its distal end. Etymology Latin “free” refers to the separate position in the genus and its Greek equivalent eleutho used in the genus name (feminine adjective). The name Eleuthemis was presumably derived from Liberia, the nation of freed slaves, from which the type species E. buettikoferi was described. Range and ecology Like all Eleuthemis favours rather swift streams and rivers with overhanging trees, recorded between 1 200 and 1 400 m a.s.l. Near Negage in Angola larvae were found in an especially gravelly and fast-flowing section, while the site in north-western Zambia was also notably rapid, including waterfalls.Published as part of Dijkstra, Klaas-Douwe B., Kipping, Jens & Mézière, Nicolas, 2015, Sixty new dragonfly and damselfly species from Africa (Odonata), pp. 447-678 in Odonatologica 44 (4) on pages 615-617, DOI: 10.5281/zenodo.3538
Umma gumma Dijkstra, Meziere & Kipping, sp. nov.
No full textUmma gumma Dijkstra, Mézière & Kipping sp. nov. – Robust Sparklewing (Type Photo 1, Photos 1–2, Fig. 1) Taxonomy Pinhey (1969) remarked that the variability of U. longistigma (Selys, 1869) might conceal unnamed taxa. Vick (1999) noted that two forms with different paraprocts occur together in Cameroon, the typical one having more slender paraprocts, as confirmed by the holotype of U. longistigma from that nation in ISNB (Fig. 1). Both forms overlap widely across central Africa, are easily separated by other characters as well, and differ genetically where they co-occur in Gabon, thus representing good species. Material studied Holotype ♂ . RMNH.INS.554425, Gabon, Haut-Ogooué Province, Alanga- Aboumi road, Moyol, gravelly and sandy forest river (2–3 m wide, 10–50 cm deep), 424 m a.s.l. (0.8403 ° S 13.9385 ° E), 28 -ix- 2012, leg. N. Mézière, RMNH. Further material. CAMEROON (South Province): 1 ♂, 26 km E of Campo, Campo-Ma’an National Park (buffer zone), Bitandé River 1 km E of Nkoélon, rocky rainforest river and adjacent roadsides, 70–100 m a.s.l. (2.3940 ° N 10.0540 ° E), 16 -vi- 2008, leg. K.- D.B. Dijkstra, J. Kipping & K. Schütte, RMNH. CONGO-BRAZZAVILLE (Region de Koulilou): 1 ♂, Conkouati National Park, forest, 50 m a.s.l. (3.7951 ° S 11.3859 ° E), 25–26 -xi- - 2010, leg. P.H. Lambret, RMNH. CONGO-KINSHASA (Province Orientale): 1 ♂, Lilanda and Baombo streams at and upstream of Lilanda and Yafake villages, sandy clear and blackwater streams and river, 390–450 m a.s.l. (0.83 ° N 24.34 ° E), 04-v- 2010, leg. K.- D.B. Dijkstra, RMNH. 1 ♂, 20 km NW of Lokutu, between Lukomete and Lunua River, forest streams and clearings, 375–400 m a.s.l. (1.275 ° N 23.425 ° E), 01–06-xi- 2004, leg. K.- D.B. Dijkstra, RMNH. 1 ♂, 20 km NW of Lokutu, large shallow, sandy, clear stream (3–5 m wide) in disturbed forest, 410 m a.s.l. (1.0866 ° N 23.5368 ° E), 01- xi- 2004, leg. K.- D.B. Dijkstra, RMNH. 3 ♂, Ubundu Road, Yoko Forest, Mokonoka Stream, rock-bottomed (and some sand, gravel, leaflitter) stream and small tributaries in rainforest, 413 m a.s.l. (0.2940 ° N 25.2888 ° E), 08- vi- 2010, leg. K.- D.B. Dijkstra, RMNH. GABON (Estuaire Province): 1 ♂ (RMNH.INS.508732), Kougouleu-Medouneu road, Monts de Cristal, Assegone-Essala, mountain stream with rapids and small falls (2–4 m wide, 50 cm deep), 316 m a.s.l. (0.5596 ° S 10.2362 ° E), 28 -xii- 2012, leg N. Mézière, RMNH. GABON (Haut-Ogooué Province): 1 ♂, Ndjima, 618 m a.s.l. (1.6643 ° S 13.0958 ° E), 21 -ii- 2009, leg. N. Mézière, RMNH. 1 ♂, Plateau d’Okouma, road to Vuku, Mounana, gravel-bottomed stream (1–2 m wide) in grassland, 507 m a.s.l. (1.3998 ° S 13.1920 ° E), 18 -ii- 2009, leg. N. Mézière, RMNH. 1 ♂ (RMNH.INS.506164), righthand road before “Africa No 1 ” radio station in the direction of Franceville, Moyabi, sandy forest river (4 m wide,> 50 cm deep) bordered by ponds, 483 m a.s.l. (0.6476 ° S 13.6800° E), 27 -iii- 2012, leg. N. Mézière, RMNH. 1 ♂ (RMNH.INS.554468), same locality, small gravelly and sandy forest stream bordered by swamp and springs, 474 m a.s.l. (1.6877 °S 13.3113 °E), 04-xi- 2012, leg. N. Mézière, RMNH. 8 ♂, Moyabi, concession Rougier, 17 km SSE of Moanda, large sandy stream in dense forest (Photo 2), 398 m a.s.l. (1.7065 °S 13.2558 °E), 24 -ix- 2013, leg. J. Kipping, CJKL. 1 ♂, same locality, same date, leg. A. Günther, CAGF. 1 ♂, same locality, 17 -ix-2013, 1♂ same locality, 01-x- 2013, leg. J. Kipping, CJKL. Genetics Four unique haplotypes (n= 4) nearest to but distinct from eleven of true U.longistigma (n = 14). Male morphological diagnosis Similar to sympatric sister-species U. longistigma by the combination of (a) a uniformly green to blue metallic head, thorax and abdomen, but dark brown to black legs and poststernum; (b) the largely pale rather than dark mandibular bases; (c) the sparse whitish hairs on the poststernum, rather than a dense patch of long dark hairs; (d) 6–10 cross-veins in the Fw quadrilateral cell; (e)the large Pt with an acute proximal corner, 2.7 –3.2 mm; (f) the absence of a clump of thick bristles near the tips of the cerci; and (g)paraprocts that are almost as long as the cerci (Fig. 1). However, is (1)larger with Hw 31.5 – 35.0 mm (mean 33.6; n = 15) rather than 30.0–33.0 mm (31.9; n = 20), with notably more robust build; (2) has a largely pale anterior face to each basal antennal segment, which is entirely dark in most U. longistigma specimens, although about a third have indistinct or partial markings; (3)no expansion on the bend of the penis’s lateral lobes, but their tips are more widened instead; (4)wide-based cerci that do not expand notably distally and end in unmodified rounded or squarish tips, while in U. longistigma the internal flanges widen notably towards the tips, which bear a transverse ridge and hollow, with often a subapical notch or tooth where the flange and ridge meet; and (5) paraprocts that are broad throughout with square-cut tips bearing an inward-directed tooth, rather than slender and tapering to rounded hooked tips (Fig. 1). Etymology Name refers to the classic 1969 Pink Floyd album »Ummagumma« (noun in apposition). Range and ecology Three Umma Kirby, 1890 species overlap widely in central Africa’s rainforests: U. mesostigma (Selys, 1879) – including the probably synonymous U.saphirina Förster, 1916 – is found closest to the source on often gravelly sections, while U. longistigma and U. gumma appear to co-occur on larger and often sandy streams (Photo 2). The new species has been recorded up to 620 m a.s.l. and favours less shady sections, occurs at lower densities, and comes down to the water only with brighter sunshine.Published as part of Dijkstra, Klaas-Douwe B., Kipping, Jens & Mézière, Nicolas, 2015, Sixty new dragonfly and damselfly species from Africa (Odonata), pp. 447-678 in Odonatologica 44 (4) on pages 457-460, DOI: 10.5281/zenodo.3538
Trithemis legrandi Dijkstra, Kipping & Meziere, sp. nov.
No full textTrithemis legrandi Dijkstra, Kipping & Mézière sp. nov. – Robust Dropwing (Type Photo 56, Photos 2, 72, Fig. 33) Taxonomy Dijkstra (2007) diagnosed the basitincta -group of Trithemis (see below) and its similar species T. basitincta Ris, 1912 and T. bifida Pinhey, 1970. However, males from Makokou in north-eastern Gabon identified as T. bifida by Jean Legrand in MNHN pertain to neither species, and similar individuals occur with both in south-eastern Gabon. Although available COI evidence is limited, the morphological distinction is sufficient to describe this species, based also on material from extreme south-western Cameroon. Damm et al. (2010) found a small difference with T. bifida in ND1, 16S and ITS; the two species together forming the sister-group of T. basitincta. Material studied Holotype ♂ . Cameroon, South Province, 26 km E of Campo, Campo-Ma’an National Park (buffer zone), Bitandé River 1 km E of Nkoélon, rocky rainforest river (Photo 72) and adjacent roadsides, 70–100 m a.s.l. (2.3940 ° N 10.0540 ° E), 15 -vi- 2008, leg. K.-D.B. Dijkstra, J. Kipping & K.Schütte, RMNH. Further material. CAMEROON (South Province): 2 ♀, as holotype, CJKL. 1 ♀, locality as holotype, 13 -vi- 2008, leg. K.-D.B. Dijkstra, J. Kipping & K. Schütte, RMNH. GABON (Haut-Ogooué Province): 1 ♂ (RMNH.INS.508171), 1 ♀ (RMNH.INS.508175), 38 km SE of Franceville, Franceville- Kessala Road, 3 rd stream E of Onkoua, gravelly stream in rainforest, 411 m a.s.l. (1.8551 ° S 13.8620 ° E), 09-i- 2010, leg. K.-D.B. Dijkstra, N. Mézière & C. Vanappelghem, RMNH. 1 ♂ 4 ♀, Concession Rougier, plot L 21 A on road to Franceville, 10 km after Moanda, Moyabi, sandy stream (1 m wide, 20– 50 cm deep), 377–398 m a.s.l. (1.7390 ° S 13.2977 ° E), 14–28 -x- 2010, leg. N. Mézière, RMNH. 2 ♂ 1 ♀, Concession Rougier, Moanda, sandy stream (Photo 2) (3–4 m, 50 cm deep), 411 m a.s.l. (1.7051 ° S 13.3560 ° E), 26 -iv- 2009, leg. C. Vanappelghem & N. Mézière, RMNH. 1 ♂ 1 ♀, Moanda-Mounana road, Mbéréssé bridge, Nguiassono, sandy stream (2 m wide, 30 cm deep) with rapids and rocks, 420 m a.s.l. (1.4953 ° S 13.1800° E), 08-i- 2011, leg. N. Mézière, RMNH. 1 ♀, Franceville-Okondja road, 2 km before Okila, muddy and gravelly stream in forest (4 m wide, <50 cm deep), 364 m as.l. (1.0790 ° S 13.5669 ° E), 23 -ix- 2010, leg. N. Mézière, RMNH. 1 ♀ (RMNH.INS.508897), same locality, 14 -x- 2013, leg. N. Mézière, RMNH. 1 ♂, Franceville-Okondja road, 1 km after Ondzei, muddy river (silted from laterite of road) bordered by dammed ponds, 351 m a.s.l. (0.7317 ° S 13.5970 ° E), 13 -iii- 2011, leg N. Mézière, RMNH. 1 ♂, Bongoville-Léconi road, swamp close to Boubou Riv- er, 421 m a.s.l. (1.6116 ° S 13.9116 ° E), 30 -x- 2010, leg. N. Mézière, RMNH. GABON (Ogooué-Ivindo Province): 1 ♂, Okondja-Makokou road, 5 km SE of Bakouaka, sandy forest stream, 531 m a.s.l. (0.1298 ° N 13.6772 ° E), 23 - ix- 2014, leg. J. Kipping, CJKL. GABON (Estuaire Province): 1 ♂ (RMNH.INS.508734), Kougouleu-Medouneu road, Monts de Cristal, Assegone-Essala, mountain stream with rapids and small falls, 2–4 m wide, 50 cm deep (0.5596 ° N 10.2362 ° E), 28 -xii- 2012, leg. N. Mézière & J. Lekogo, RMNH. Genetics Two unique haplotypes (n = 4) are very similar to six of T. bifida (n = 11) and the two species are potentially inseparable. Male morphological diagnosis Belongs to the basitincta -group by the combination of (a) Fw discoidal field of 3 rows at base; (b) 4–6 cell-doublings in radial planate and thus two rows of cells there; (c) hamule with long sickle-shaped hook; and (d) the genital lobe directed away from the hamule (Fig. 33). Nearest to the sympatric T.basitincta and T. bifida by (e) the fairly large size, Hw 31.8–33.5 mm (n = 4); (f) 13 ½– 16 ½ Ax in Fw; (g) the large and dark patch at Hw base that fills the cubital space to beyond Cux and extends into the anal field, thus appearing triangular and nearest to T. basitincta as Fig. 6 s in Dijkstra (2007), although sometimes also present in the subcostal space proximal to Ax 1; (h)the parallel-sided anterior lamina with a broad and deeply-notched apex; and (i) the notably slender and sabre-like hamules (Fig. 33). Only separated reliably by (1) the apical peaks of the anterior lamina that in lateral view are not dark and pointedly rounded, but have pale flat-topped swellings, which in caudal or rostal view may appear to close the top of the cleft between them. The swelling is greater and the cleft completely closed in the sympatric T. aconita Lieftinck, 1969 and T. congolica Pinhey, 1970, while T.tropicana Fraser, 1953 has a fairly flat but neither swollen nor cleft anterior lamina (Fig. 33). Also is (2) shorter-bodied, Hw being 79–82 % of full length rather than 73–78 %; and (3) as males darken with age, the pale markings remain visible last ventrally on S 4–7, while any visible dorsal markings lie on the lateral carina and are shorter than half the segments’ length. The dorsal markings that are visible last in T. bifida lie closer to the dorsal than lat- eral carinae and extend over almost the full segment length. The markings are intermediate in T. basitincta, with the rather short final markings lying about halfway the dorsal and lateral carinae. That species, however, can be separated by the presence of short bristles rather than long hairs at the base of the anterior lamina (Fig. 33). Trithemis legrandi also often has large and distinct lateral yellow spots on the labium, postclypeus and frons even in very dark specimens, e.g., those virtually lacking pale abdominal markings. Etymology Named in honour of Dr Jean Legrand, collector of the first material of this species, and successor to Elliot Pinhey as the leading specialist of African Odonata of his time (masculine genitive noun). Range and ecology Sandy, gravelly and rocky streams and small rivers in rainforest between 50 and 500 m a.s.l. in Gabon and southern Cameroon. Males fly rapidly over water but rarely perch by the streams.Published as part of Dijkstra, Klaas-Douwe B., Kipping, Jens & Mézière, Nicolas, 2015, Sixty new dragonfly and damselfly species from Africa (Odonata), pp. 447-678 in Odonatologica 44 (4) on pages 652-656, DOI: 10.5281/zenodo.3538
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