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FIGURE 1 in Revision of Heterococcopsis Borchsenius (Hemiptera: Coccoidea: Pseudococcidae), with description of a new genus with two new species from Turkey
No full textFIGURE 1. Heterococcopsis lonicerae Borchsenius.Published as part of Kaydan, Bora, 2011, Revision of Heterococcopsis Borchsenius (Hemiptera: Coccoidea: Pseudococcidae), with description of a new genus with two new species from Turkey, pp. 49-62 in Zootaxa 2970 on page 52, DOI: 10.5281/zenodo.20396
FIGURE 2 in Two new genera and species of Eriococcidae (Hemiptera: Sternorrhyncha: Coccoidea) with new data on the family in Turkey
No full textFIGURE 2. Adult female of Kotejacoccus turcicus Kaydan & Kozár sp. nov.Published as part of Kaydan, Bora & Kozár, Ferenc, 2008, Two new genera and species of Eriococcidae (Hemiptera: Sternorrhyncha: Coccoidea) with new data on the family in Turkey, pp. 16-26 in Zootaxa 1848 on page 22, DOI: 10.5281/zenodo.18334
Two new species of Coccidohystrix Lindinger (Hemiptera: Coccomorpha: Pseudococcidae) with notes on the related genus Artemicoccus Balachowsky
No full textScale insect (Hemiptera: Coccomorpha: Pseudococcidae) samples were collected in Turkey (Hakkari-
Yüksekova) and Bulgaria (Kresna Gorge) between 2007 and 2012. Two new Coccidohystrix Lindinger species, are
described and illustrated. An identification key including the new species is also provided. Additionaly, the type
material of C. zangherii was studied and the species is transferred into the genus Artemicoccus Balachowsky. A new
identification key for species in the genus Artemicoccus is also provided
Heterobrevennia
No full textKey to <i>Heterobrevennia</i> species <p>1. Multilocular pores scattered throughout dorsum; coxa without transculent pores.................................... 2</p> <p> - Multilocular pores present dorsally only on abdominal segments VI and VII: coxa with transculent pores............................................................................................... <i>H. kozari</i> Kaydan <b>spec. nov.</b></p> <p> 2. Cerarii totalling 6 pairs; with two groups of multilocular pores on dorsum of head; many trilocular pores present on dorsum............................................................................. <i>H. gullanae</i> Kaydan <b>spec. nov.</b></p> <p> - Cerarii totalling 7 pairs; without multilocular pores on dorsum of head; trilocular pores few on dorsum, restricted to thorax, on posterior abdominal segments, in cerarii and on ostioles..................................... <i>H. opertus</i> Borchsenius</p>Published as part of <i>Kaydan, Bora, 2011, Revision of Heterococcopsis Borchsenius (Hemiptera: Coccoidea: Pseudococcidae), with description of a new genus with two new species from Turkey, pp. 49-62 in Zootaxa 2970</i> on page 54, DOI: <a href="http://zenodo.org/record/203960">10.5281/zenodo.203960</a>
Heterococcus
No full textKey to adult females of the <i>Heterococcus</i> group <p> 1. Multilocular pores absent..................................................................... <i>Stachycoccus</i></p> <p>- Multilocular pores present at least around vulva............................................................. 2</p> <p> 2. Circuli numbering 4; tubular ducts small, short and stout............................................ <i>Annulococcus</i></p> <p>- Circuli numbering 0-3; tubular ducts elongate............................................................... 3</p> <p>3. Trilocular pores totally absent from body................................................................... 4</p> <p>- Trilocular pores present at least around spiracle opening, in each cerarius and on ostioles............................. 6</p> <p> 4. Tarsal claw without a denticle; anterior ostioles absent............................................ <i>Pseudorhodania</i></p> <p>- Tarsal claw with a denticle; anterior ostiole present........................................................... 5</p> <p> 5. Anal lobe cerarii each with more than 2 enlarged setae; quinquelocular pores only in cerarii and on ostioles..... <i>Boreococcus</i></p> <p> - Anal lobe cerarii each with only 2 slender cerarian setae; quinquelocular pores scattered all over body........ <i>Heterococcus</i></p> <p>6. Ostioles completely absent.............................................................................. 7</p> <p>- At least posterior ostiole present.......................................................................... 8</p> <p> 7. Circuli oval, cerari absent..................................................................... <i>Laingicoccus</i></p> <p> - Circuli absent; cerarii present at least last two abdominal segments................................... <i>Paramococcus</i></p> <p> 8. Circuli present, numbering 3; legs stout........................................................ <i>Heterococcopsis</i></p> <p>- Circuli numbering 0 or 1; legs slender..................................................................... 9</p> <p> 9. Tarsal claw without a denticle; anterior ostiole absent; antennae 6 or 7 segmented........................... <i>Brevennia</i></p> <p> - Tarsal claw with a denticle; anterior ostiole present; antennae 8 or 9 segmented............... <i>Heterobrevennia</i> <b>gen. nov.</b></p>Published as part of <i>Kaydan, Bora, 2011, Revision of Heterococcopsis Borchsenius (Hemiptera: Coccoidea: Pseudococcidae), with description of a new genus with two new species from Turkey, pp. 49-62 in Zootaxa 2970</i> on page 50, DOI: <a href="http://zenodo.org/record/203960">10.5281/zenodo.203960</a>
Heterococcopsis Borchsenius
No full text<i>Heterococcopsis</i> Borchsenius <p> Type species: <i>Heterococcopsis lonicerae</i> Borchsenius, 1948</p> <p> <i>Heterococcopsis lonicerae</i> Borchsenius 1948: 955, by original designation.</p> <p> <b>Generic diagnosis. Adult female.</b> Body elongate to oval, 1.8–2.0 mm long, 0.82–1.28 mm wide. Antennae 9 segmented. Eyes present on margin. Labium 3 segmented. Posterior pair of spiracles always larger than anterior spiracles. Legs well developed, with translucent pores on hind tibia; tibia broad, tarsus shorter than tibia, claw with a denticle; tarsal digitules setose, claw digitules capitate, thicker than tarsal digitules. Posterior ostioles well developed, anterior ostioles very weakly developed or not detected. Heptalocular pores smaller than other multilocular pores, scattered on venter; absent from dorsum. Oral-collar tubular ducts elongate, generally situated on body margin. Anal ring with 6 setae.</p> <p> <b>Dorsum.</b> Cerarii numbering 2 pairs, situated on posterior 2 abdominal segments. Dorsal body setae spine-like. Multilocular pores present on abdominal segments and near body margin. Quinquelocular pores scattered over body. Trilocular pores present on last abdominal segment (VIII) in cerarii and around ostioles. Oral-collar tubular ducts few, generally present on body margin.</p> <p> <b>Venter.</b> Body setae slender, generally short, longest setae present medially on head; apical setae long. Three circuli present. Multilocular disc pores present on posterior abdominal segments (IV–VIII+IX) plus a few on thorax and head. Heptalocular pores and quinquelocular pores about same size, scattered over venter. Minute discoidal pores scattered on body surface. Oral-collar tubular ducts few.</p> <p> <b>Comment.</b> Hitherto, <i>Heterococcopsis</i> included three species: <i>H. desertus</i> Bazarow & Nurmamatov, <i>H. lonicera</i> Borchsenius and <i>H. opertus</i>. Herein, <i>H. desertus</i> is transferred to <i>Phenacoccus</i>, and <i>H. opertus</i> is transferred to <i>Heterobrevennia</i> <b>gen. nov.</b>, leaving <i>Heterococcopsis</i> as a monotypic genus.</p>Published as part of <i>Kaydan, Bora, 2011, Revision of Heterococcopsis Borchsenius (Hemiptera: Coccoidea: Pseudococcidae), with description of a new genus with two new species from Turkey, pp. 49-62 in Zootaxa 2970</i> on pages 50-51, DOI: <a href="http://zenodo.org/record/203960">10.5281/zenodo.203960</a>
Phenacoccus desertus Bazarov & Nurmamatov, comb. nov.
No full textPhenacoccus desertus (Bazarov & Nurmamatov), comb. nov. Heterococcopsis desertus Bazarov & Nurmamatov, 1975: 63. Material examined. Holotype adult female: Tadzhikistan, Gbao, Vosim, Gasur Lurgap, Org. Artemisia demissa (Asteraceae), L. Kuarzivsheva, 18.vii. 1924 (ZIAS). Adult female (Fig. 5) Live appearance. Body oval. Mounted material. Body oval, 2.20 mm long, 1.34 mm wide. Eye marginal, 35.0 µm wide. Antenna 9 segmented, 130–140 µm long; apical segment 47.5 µm long, 20 µm wide, with apical setae 22.5 µm long, plus 3 fleshy setae, each 20–30 µm long. Clypeolabral shield 145 µm long, 120 µm wide. Labium 110 µm long, 75 µm wide. Anterior spiracles 45 µm long, 15 µm wide across atrium; posterior spiracles 45.0– 47.5 µm long, 25.0– 27.5 µm wide across atrium. Legs well developed; coxa 40 µm long, hind trochanter + femur 150 µm long, hind tibia + tarsus 165 µm long, hind claw 25.0 µm long. Ratio of lengths of hind tibia + tarsus to hind trochanter + femur 1.10, ratio of lengths of hind tibia to tarsus 1.3: 1, ratio of length of hind trochanter + femur to greatest width of femur 3.3: 1. Tarsal digitules setose, each 10 µm long. Claw digitules subequal, each 17.5 µm long, knobbed and broader than tarsal digitules. Translucent pores present on femur and tibia of hind legs, totalling 16–24. Both pairs of ostioles present, posterior ostioles each with 9 trilocular pores and 1 setae, anterior ostioles each with 4 trilocular pores and 2 or 3 setae. Anal ring 55 µm wide, with 6 anal-ring setae, each 60–65 µm long. Dorsum. Cerarii numbering 2 pairs, with pairs on posterior 2 abdominal segments; anal lobe cerarii each with 2 conical setae (broken, as shown on figure), plus 5–7 trilocular pores; C 17 each with 2 conical setae, each 10 µm long, plus 2 trilocular pores. Dorsal body setae spine-like, each 5–10 µm long. Multilocular pores each 7.5 –9.0 µm wide with 11 loculi, present in laterally in abdominal segment I and in rows or bands on posterior abdominal segments (II–VII): 12 pores on segment on II, 19 on III, 37 on IV, 37 on V, 24 on VI, 30 on VII and no pores on segment VIII. Quinquelocular pores very few (only three pores seen on head and thorax), each 5–6 µm. Trilocular pores each 4–5 µm in diameter, scattered throughout. Oral-collar tubular ducts each 7.5 –10.0 µm long, 2–3 µm wide, present in rows on all abdominal segments plus 8 on head and thorax laterally; distributed on abdomen as follows: 3 on I; 5 on II; 6 on III; 6 on IV; 7 on V; 2 on VI; 3 on VII; 4 on VIII in row. Venter. Body setae slender, each 12.5 –40.0 µm long, longest setae medially on head; apical setae of anal lobe broken. Multilocular disc pores present, each 7.5 –8.0 µm in diameter, present laterally on abdominal segments I–II and in rows or bands on posterior abdominal segments (III–VIII + IX): 22 pores on segment III, 34 on IV, 36 on V, 48 on VI, 37 on VII, and 27 on VIII + IX. Quinquelocular pores each 5–6 µm wide, scattered throughout. Oral-collar tubular ducts each 7.5 –9.0 µm long, 2.5 –3.0 µm wide, present in rows on all abdominal segments plus a few on head and thorax; distributed as follows: 1–2 on each side of abdominal segments I–III; 13 on IV; 14 on V; 8 on VI in row or band and 3 on VII; 3 on VIII laterally. Comment. Species in the Heterococcus -group differ from those in Phenacoccus Cockerell in having: (i) quinquelocular pores on the dorsum and either no trilocular pores or trilocular pores restricted to around each spiracle. Because Heterococcopsis desertus lacks both of these character states, it is not congeneric with Heterococcopsis lonicerae Borchsenius, the type species of the genus Heterococcopsis but does appear to be congeneric with Phenacoccus aceris Signoret, the type species of the genus Phenacoccus. H. desertus shares the following important features with P. aceris: (i) presence of many trilocular pores on dorsum and venter, (ii) very few quinquelocular pores (only 3 were found) on dorsum, and (iii) a long labium. Based on the morphological affinities of H. desertus with P. aceris, I here transfer H. desertus to Phenacoccus, as Phenacoccus desertus (Bazarov & Nurmamatov) comb. nov. and illustrate the species based on the type material. This species is very close to Phenacoccus tergrigorianae Borchsenius & Ter-Grigorian, and P. p u m i l u s Kiritshenko in having multilocular pores on dorsum, but differs from both in having many quinquelocular pores on the venter. The genus Phenacoccus has many species and needs further study to clarify the status of the new combination of P. desertus.Published as part of Kaydan, Bora, 2011, Revision of Heterococcopsis Borchsenius (Hemiptera: Coccoidea: Pseudococcidae), with description of a new genus with two new species from Turkey, pp. 49-62 in Zootaxa 2970 on pages 60-62, DOI: 10.5281/zenodo.20396
Heterobrevennia opertus Borchsenius, comb.nov.
No full textHeterobrevennia opertus (Borchsenius) comb.nov. Material examined: Syntypic adult females: Tadzikistan, Gissarskii Ridge, Dactylis glomerata (Poaceae), H. Borchsenius, 17.vii. 1944 (2 specimens, ZIAS: 34-45). Adult female (Fig. 4) Live appearance: Body elongate oval. Mounted material. Body elongate oval, 2.82–3.60 mm long, 1.56–2.20 mm wide. Eye marginal, 32.5 –35.0 µm wide. Antenna 9 segmented, 270–290 µm long; apical segment 45–55 µm long, 20–25 µm wide, with apical setae 30–35 µm long, plus 3 fleshy setae, each 30–35 µm long. Clypeolabral shield 145–150 µm long, 110–115 µm wide. Labium 80–85 µm long, 65–70 µm wide. Anterior spiracles 42.5–52.5 µm long, 25.0– 27.5 µm wide across atrium; posterior spiracles 50–60 µm long, 25–35 µm wide across atrium. Legs well developed; coxa 90–105 µm long, hind trochanter + femur 215–220 µm long, hind tibia + tarsus 255–260 µm long, hind claw 25.0– 27.5 µm long. Ratio of lengths of hind tibia + tarsus to hind trochanter + femur 1.16–1.18: 1, ratio of lengths of hind tibia to tarsus 1.8–2.03: 1, ratio of length of hind trochanter + femur to greatest width of femur 4.4–4.8: 1. Tarsal digitules setose, each 32.5 –35.0 µm long. Claw digitules subequal, each 27.5 –35.0 µm long, knobbed and broader than tarsal digitules. Translucent pores present on femur and tibia of hind legs, totalling 16–24. Both pairs of ostioles present, posterior ostioles each with 5–7 trilocular pores and 2 or 3 setae, anterior ostioles each with 5 trilocular pores and 2 or 3 setae. Anal ring 60–62.5 µm wide, with 6 anal-ring setae, each 85–95 µm long. Dorsum. Cerarii numbering 7 pairs, with pairs on posterior 4 abdominal segments, and on 3 pairs on head; anal lobe cerarii each with 2 conical setae, each 15.0– 17.5 µm long, plus 5–7 trilocular pores; C 17 each with 2 conical setae, each 15.0– 17.5 µm long, plus 4 trilocular pores; C 16 each with 2 conical setae, each 15 µm long, plus 2 trilocular pores; C 15 each with 2 conical setae, each 11.0– 12.5 µm long, plus 2 trilocular pores; C 1 each with 2 conical setae, 10 µm long, plus 2 trilocular pores; C 2 each with 2 conical setae, each 10 µm long, plus 3 trilocular pores; and C 3 each with 3 conical setae, each 7.5–10 µm long, plus 2 trilocular pores. Dorsal body setae spine-like, each 10.0– 12.5 µm long. Multilocular pores each 7.5 –8.0 µm wide with 11 loculi, scattered throughout. Quinquelocular pores each 5–6 µm, scattered throughout. Trilocular pores each 5–6 µm in diameter, restricted to posterior abdominal segments VII, VIII and IX, and in cerarii, on ostioles and the middle area of thorax. Oral-collar tubular ducts present along body margin, each 9–11 µm long, 2–3 µm wide. Venter. Body setae slender, each 15–65 µm long, longest setae medially on head; apical setae of anal lobe 145–165 µm long. Multilocular disc pores each 7.5 –8.0 µm in diameter, present in rows or bands on posterior abdominal segments (IV–VIII + IX): 12 pores on segment IV, 12–24 on V, 32–36 on VI, 53–62 on VII, and 27–33 on VIII + IX. Quinquelocular pores each 5–6 µm wide, scattered throughout. Oral-collar tubular ducts each 7.5 – 10.0 µm long, 2.5 –3.0 µm wide, present in rows on all abdominal segments plus a few on head and thorax; distributed as follows: 2–4 on each side of abdominal segments I–III and a few medially; 4 on IV; 15–25 on V; 14–24 on VI; 19 on VII; 18–23 on VIII in row or band. Comment. For a comparison of H. opertus with other species in the genus, see under H. gullae and H. kozari above.Published as part of Kaydan, Bora, 2011, Revision of Heterococcopsis Borchsenius (Hemiptera: Coccoidea: Pseudococcidae), with description of a new genus with two new species from Turkey, pp. 49-62 in Zootaxa 2970 on pages 58-60, DOI: 10.5281/zenodo.20396
Heterococcopsis lonicerae Borchsenius
No full textHeterococcopsis lonicerae Borchsenius Material examined: Lectotype adult female (here designated) + 2 paralectotype adult females (on 1 slide): Kazakhstan, Alma-Ata, Lonicera tatarica (Caprifoliaceae), Saygurov, 02.vi. 36 (ZIAS: 80 / 37). Lectotype specimen in circle on left side, other 2 specimens indicated as paralectotypes. Adult female (Fig. 1) Live appearance: Body oval. Mounted material. Body elongate oval, 1.80 –2.00 mm long, 0.82–1.28 mm wide. Eyes located on margin, each 30 µm wide. Antenna 9 segmented, 270 – 235 µm long; apical segment 35–45 µm long, 17.5 –20.0 µm wide, with apical setae 27.5 –30.0 µm long and with 3 fleshy setae, each 27.5 –30.0 µm long. Clypeolabral shield 150–165 µm long, 110 µm wide. Labium 80–90 µm long, 62.5 –70.0 µm wide. Anterior spiracles 37.5 –45.0 µm long, 20–25 µm wide across atrium; posterior spiracles 50 µm long, 22.5 –25.0 µm wide across atrium. Circuli numbering 3, each oval in shape, each 35 µm, 32.5 µm and 20 µm wide respectively. Legs well developed; coxa 70–80 µm long, hind trochanter + femur 130–145 µm long, hind tibia + tarsus 155–160 µm long, hind claw 20.0– 22.5 µm long. Ratio of lengths of hind tibia + tarsus to hind trochanter + femur 1.10–1.19: 1, ratio of lengths of hind tibia to tarsus 1.71–1.83: 1, ratio of length of hind trochanter + femur to greatest width of femur 3.71–4.14: 1. Tarsal digitules setose, each 15–20 µm long. Claw digitules simalar and capitate, each 17.5 –20.0 μm long, knobbed and thicker than tarsal digitules. Translucent pores present on tibia of hind legs, totalling 32–43. Posterior pairs of ostioles present, each with 4 trilocular pores and 3 setae; anterior ostiole, not easy to detect or poorly developed. Anal ring 55 µm wide, with 6 anal ring setae, each seta 50–60 µm long. Dorsum. Cerarii numbering 2 pairs, situated on posterior 2 abdominal segments: anal lobe cerarii each with 2 conical setae (each 14–16 µm long) plus 5 trilocular pores and C 17 each with 2 conical setae (each 15 µm long) plus 2 trilocular pores. Dorsal body setae spine-like, each 5.0– 7.5 µm long. Multilocular pores each 6.25–7.5 µm wide with more than 10 loculi, present across posterior 2 abdominal segments and along body margin. Quinquelocular pores each 5 µm in diameter, scattered over dorsum but absent from last abdominal segment. Trilocular pores each 4–5 µm in diameter, restricted to last abdominal segment, cerarii and posterior ostioles. Oral-collar tubular ducts, each 9–11 µm long, 2–3 µm wide, present in a transverse row across last abdominal IV-VIII segments and along body margin. Venter. Setae slender, each 20–55 µm long, longest setae present medially on head; apical setae of anal lobe 75–105 µm long. Multilocular disc pores each 7.5 –8.0 µm in diameter, present across abdominal segments IV −VIII + IX as follows: 0–6 pores on posterior margins of segment IV, 1–12 pores on posterior margins of V, 18–22 pores on posterior margins of VI, 30–32 across full width of VII, 19–34 across full width of VIII + IX. Quinquelocular and heptalocular pores each 4–5 µm wide, scattered over venter. Oral-collar tubular ducts each 8–11 µm long, 2.5 – 3.0 µm wide, present in a row on last abdominal segment and on margins and submargins of each abdominal segment; ducts distributed as follows: 2–4 on segment I; 2–4 on II; 1–4 on III; 3–4 on IV; 4 on V; 4 on VI; 4–6 on VII; and 4–6 on VIII and a few on the metathorax and the mesothorax. Comment: For discussion, see under Heterobrevinia gen. nov. below.Published as part of Kaydan, Bora, 2011, Revision of Heterococcopsis Borchsenius (Hemiptera: Coccoidea: Pseudococcidae), with description of a new genus with two new species from Turkey, pp. 49-62 in Zootaxa 2970 on pages 51-53, DOI: 10.5281/zenodo.20396
Heterobrevennia Kaydan, gen. nov.
No full textHeterobrevennia Kaydan, gen. nov. Type species: Heterobrevennia opertus ( Borchsenius, 1949) Generic diagnosis. Adult female: Body elongate, 2.82–3.20 mm long, 1.56–2.20 mm wide. Antennae generally 8 or 9 segmented (but occasionally 7 when 2 nd and 3 rd segments fused). Eyes present on margin. Labium 3 segmented. Posterior spiracles slightly larger than anterior spiracles. Legs well developed, with translucent pores on hind femur and tibia; claw with a denticle; tarsal digitules setose, claw digitules capitate, broader than tarsal digitules. Both anterior and posterior ostioles well developed. Multilocular disc pores limited to margins of both body surfaces and to posterior abdominal segments. Quinquelocular pores smaller than multilocular pores, scattered over both body surfaces. Trilocular pores present on dorsum. Oral-collar tubular ducts elongate, generally situated on body margin. Anal ring with 6 setae. Dorsum. Cerarii numbering always 5 or more pairs, situated on posterior abdominal segments and on head. Dorsal body setae spine-like. Multilocular pores situated on body margin. Quinquelocular pores scattered throughout. Trilocular pores scattered throughout, sometimes few, but always present at least on cerarii, ostioles and near spiracle opening. Minute discoidal pores scattered throughout. Oral-collar tubular ducts elongate scattered throughout. Venter. Body setae slender, longest setae present medially on head; apical setae long; spine-like setae present submarginally on body. Multilocular disc pores present on posterior abdominal segments and on margin of thorax and head. Quinquelocular pores scattered over body. Trilocular pores present around atrium of spiracle, absent elsewhere. Minute discoidal pores scattered over body. Oral-collar tubular ducts in rows on posterior abdominal segment and scattered on thorax and head. Comment. Heterobrevennia gen. nov. is most similar to Heterococcopsis in having trilocular pores on the dorsum and 8 - or 9 -segmented antennae. The two genera differ as follows (features of Heterococcopsis in parentheses): i) no circuli (with 3 circuli), ii) trilocular pores scattered on dorsum (trilocular pores on dorsum restricted to cerarii, ostioles and posterior abdominal segments), iii) at least five cerarii, of which 3 are present on head (only 2 cerarii, restricted to posterior abdominal segments), and iv) legs slender (legs stout). Heterobrevennia is also similar to all the other genera in the Heterococcus -group. The affinities of Heterobrevennia with other related genera are discussed below. Asphodelococcus can be readily separated from Heterobrevennia by the following combination of features (features of Heterobrevennia in parenthesis): (i) antennae 6 segmented (antennae 9 segmented), (ii) multilocular pores of unusual shape and often quadrate, and with heavily sclerotised hexagonal pores (pores normal), and (iii) cerarii numbering 4 pairs and restricted to abdomen (cerarii numbering 5 or more pairs, present on both abdomen and head). Annulococcus differs from Heterobrevennia in the following combination of features (features of Heterobrevennia in parenthesis): (i) cerarii poorly developed, present along the body margin (cerarii numbering 5 or more pairs, present on both abdomen and head), (ii) quinquelocular pores larger than other multilocular pores (quinquelocular pores smaller than other multilocular pores), (iii) oral-collar tubular ducts very short and stout (oral-collar tubular ducts elongate), and (iv) trilocular pores absent (trilocular pores present) (Miller, 1975; Foldi and Cox, 1989). Brevennia Goux shares many morphological features with Heterobrevennia, namely, the presence of trilocular pores on the dorsum and venter and the abundance of quinquelocular pores on both dorsum and venter, but the two genera differ in the absence of claw denticles and Brevennia also lacks anterior ostioles. Boreococcus Danzig, Heterococcus Borchsenius and Pseudorhodania can be readily distinguished from Heterobrevennia due to the complete absence of trilocular pores in these three genera. Laingococcus Williams differs from Heterobrevennia in having the following combination of features (features of Heterobrevennia in parenthesis): (i) ostioles absent (ostioles present), (ii) cerarii absent (5 or more cerarii present on both abdomen and head); (iii) a circulus present (absent); (iv) dorsal body setae similar in size to those on venter (dorsal body setae spinose, ventral setose); and (v) in having a rotund body (body elongate oval). In having trilocular pores on the dorsum and venter, Heterobrevennia is somewhat similar to Paramacoccus Foldi & Cox but Paramacoccus is unique in lacking ostioles. On the other hand, in having trilocular pores on the dorsum and venter, and spine-like setae on dorsum, Heterobrevennia is somewhat similar to Phenacoccus Cockerell but new genus differs in having many quinquelocular pores on the dorsum. Etymology. The name Heterobrevennia refers to its similarity to the genera Heterococcus and Brevennia. Gender female.Published as part of Kaydan, Bora, 2011, Revision of Heterococcopsis Borchsenius (Hemiptera: Coccoidea: Pseudococcidae), with description of a new genus with two new species from Turkey, pp. 49-62 in Zootaxa 2970 on pages 53-54, DOI: 10.5281/zenodo.20396
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