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Anafroptilum Kluge 2012
No full textGenus Anafroptilum Kluge 2012 (Figs 1–87) Anafroptilum Kluge 2012: 372. Type species: Centroptilum kazlauskasi Kluge 1983. Systematic position and characteristics of Anafroptilum. Systematic position of Anafroptilum is vague: from one hand, it shares some peculiar characters common with Cloeonini sensu Kluge 2016b; from other hand, its patella-tibial suture is characteristic for the taxon Protopatellata, while Cloeonini do not belong to Protopatellata. Character of Anafroptilum common with Protopatellata. (1) In larva, subimago and imago of both sexes patella-tibial suture is absent on fore legs and present on middle and hind legs (Kluge 2012: Figs 13, 14). This character is plesiomorphic, being the same as in the outgroup, i.e. in the most non-baetid mayflies and in the primitive baetid taxa, Siphlaenigma Penniket 1962 and Palaeocloeon Kluge 1997 (Kluge 1997). Among Baetidae in the narrow sense, i.e. Turbanoculata Kluge 1997, this character is present in the taxon Protopatellata Kluge & Novikova 2011 (= Afroptilinae sensu Kluge 1997). In contrast to this, the taxon Anteropatellata Kluge 1997 (comprising overwhelming majority of Baetidae) is characterized by presence of patella-tibial suture on all legs other than fore legs of male imago and subimago. Exceptions are made only by baetids with highly modified leg structure; unlike them, Anafroptilum has leg structure non-modified, similar to that of Cloeonini sensu Kluge 2016b and many other taxa. Characters of Anafroptilum common with Cloeonini. The taxon Cloeonini sensu Kluge 2016b includes genera Cloeon Leach 1815 (in narrowest sense), Procloeon Bengtsson 1915, Similicloeon Kluge & Novikova 1992 and related taxa (Kluge 2016b). In contrast to Anafroptilum, all representatives of Cloeonini have patella-tibial suture equally developed on all legs of larva and female, so Cloeonini undoubtedly belong to Anteropatellata (see above). Besides evident symplesiomorphies, Anafroptilum and Cloeonini share the following common characters: (2) Larval labial palp with 3rd segment truncate and 2nd segment non-projected (Figs 16, 78). The same in some other taxa. (3) Larval abdominal segments VIII and IX with spines forming one longitudinal row on each lateral margin; sometimes such spines are present also on some of previous abdominal segments. This character is constant for Cloeonini and not found in any taxa other than Cloeonini and Anafroptilum. Among Anafroptilum, these spines are either constantly present (e.g. in A. kazlauskasi, A. bifurcatum, A. victoriae), or occasionally absent (e.g., in A. album, A. conturbatum) (Lowen & Flannagan 1991). In the two new species from Thailand, these spines are either poorly developed, or absent (see below). (4) Male imaginal gonovectes are fused with penial bridge (Figs 41, 79). Among Protopatellata, the same in the Afrotropical taxon Rhithrocloeoninae (Kluge 2015). Symplesiomorphies of Anafroptilum, Cloeoninae, some Protopatellata and some other taxa. (5) Larval claw with 2 equal rows of denticles (in contrast to Baetungulata and some other taxa). (6) Fore wing with single marginal intercalaries (in contrast to Baetovectata). (7) In subimago of both sexes either all segments of all tarsi are covered with pointed microlepides, or only proximal part of tarsus is covered with microtrichiae (Fig. 49); blunt microlepides are absent on all tarsomeres (in contrast to Baetungulata). (8) Subimaginal gonostyli developing under larval cuticle are folded in « Cloeon - type » pose (Fig. 43) (in contrast to Baetovectata). Characters of Anafroptilum of unclear phylogenetic status. (9) Dorsal surface of labrum with irregularly situated setae, without constant pair of submedian setae and without regular latero-distal setal rows (Figs 35, 76). The same in some Cloeonini and in Centroptilum. (10) Both mandibles with incisor and kinetodontium deeply separated (Figs 13–14, 38–39, 61–62). The same in Centroptilum, selected species of Procloeon and some others. (11) Both mandibles with dense setae between prostheca and mola (Figs 13–14, 61–63). The same in all Cloeonini, Centroptilum and some others. (12) Maxilla of the « Cloeon - type », i.e. with all 3 canines and 3 dentisetae slender, pointed and bent at the same direction (Figs 17, 77). The same in many other taxa. (13) Glossa and paraglossa of the « Cloeon - type », i.e. subequal in length and width, pointed, paraglossa with no more than one regular row on outer-apical margin (Figs 36, 78). The same in many other taxa. (14) Larval femora without row of stout setae on outer margin and without stout outer-apical setae; outer-apical margin serrate, with sparse thin hairs (Fig. 59). The same in Centroptilum and selected species of Procloeon. (15) Male imaginal gonostylus with 2nd segment thickened apically, 3rd segment petiolate (Figs 41, 79). The same in Centroptilum and Cheleocloeon (Kluge 2016a: Figs 98, 106, 108, 110). In Cloeonini, second segment of gonostylus is also widened apically, and third segment is also petiolate, but smaller. In other respects external genitals of Anafroptilum are highly variable: 1st segment of gonostylus either with characteristic projection on inner side (Fig. 79), or without it (Fig. 41); 2nd segment either arched (Fig. 79), or straight (Fig. 41); median finger-like projection either present (Figs 41, 79), or absent. Variable characters of Anafroptilum. (16) Hind wings either present, or absent (in A. minor and two new species described below). If present, hind wing is either of the « Baetis -type», i.e. relatively wide, with three longitudinal veins and short costal projection (e.g., in A. bifircatum), or of the « Centroptilum -type», i.e. narrow, with two longitudinal veins and hooked costal projection (e.g., in A. kazlauskasi). (17) Tergalii either asymmetrically widened (Figs 69–75), or narrow and symmetric (Figs 28–34). Distribution. Amphipacific: Asia (East Palaearctic and Oriental Regions) and Nearctic.Published as part of Kluge, Nikita J. & Novikova, Eugenia A., 2017, Occurrence of Anafroptilum Kluge 2012 (Ephemeroptera: Baetidae) in Oriental Region, pp. 453-472 in Zootaxa 4282 (3) on pages 454-461, DOI: 10.11646/zootaxa.4282.3.2, http://zenodo.org/record/82788
Cloeodes inferior Kluge 1991
No full text<i>Cloeodes inferior</i> Kluge 1991 <p>(Fig. 142)</p> <p> <i>Cloeodes inferior</i> Kluge 1991: 133 (larva, male and female imagoes, eggs).</p> <p> <b>Material examined.</b> Holotype: L-S-I ♂ {specimen [XVIII](2)A1989}: CUBA, Prov. Guamtanamo, Rio Duaba near Baracoa, 20°22'N, 74°32'W, 15.III. 198, coll. N. Kluge. Paratypes: the same locality, 1 L-S-I ♂, 3 L-S-I ♀, 3 L- S ♂, 4 L-S ♀, 14 larvae; Prov. Santiago-de-Cuba, Rio Baconao in Las Yaguas, 20°04'N, 75°34'W, 12.II.1989, coll. N. Kluge: 2 L-S ♂, 1 larva; the same locality, 25.XII.1983, coll. C. Naranjo: 6 larvae; Trinidad mountains (border of provinces Sanctispiritus, Cienfiegos and Santa Clara), 12–18.IV.1989, coll. N. Kluge: 1 larva.</p> <p> <b> Additional descriptions. <i>Larva</i>.</b> Maxillary canines and distal dentiseta stout; distal dentiseta widened, with apex somewhat hooked toward canines (Fig. 142; as in Fig. 8). Other characters as described by Kluge 1991.</p> <p> <i>Subimago.</i> CUTICULAR COLORATION: as in <i>C. superior</i> (see above).</p> <p>HYPODERMAL COLORATION: As in imago (Kluge 1991).</p> <p>TEXTURE: On all legs of both sexes all tarsal segments entirely covered by pointed microlepides (Table 1).</p> <p> <b> <i>Tarsal spines of subimago and imago</i>.</b> Tarsus of fore leg in female with one apical spine on 3dr tarsomere (in male without apical spines). Tarsus of middle and hind leg of both sexes with 2 apical spines: one sine on 1st+2nd and another spine on initial 3rd tarsomere (Table 1). Other characters of imago as described by Kluge 1991.</p> <p> <b>Dimension.</b> Fore wing length 3.5– 4 mm.</p> <p> <b>Distribution.</b> Cuba.</p>Published as part of <i>Kluge, Nikita J., 2017, Contribution to the knowledge of Cloeodes Traver 1938 (Ephemeroptera, Baetidae), pp. 91-127 in Zootaxa 4319 (1)</i> on pages 125-126, DOI: 10.11646/zootaxa.4319.1.5, <a href="http://zenodo.org/record/888627">http://zenodo.org/record/888627</a>
Procloeon (Oculogaster) Kluge 2016
No full textOculogaster Kluge 2016 (Figs 1–133) Subgenus Oculogaster Kluge 2016: 494. Type species: Procloeon cylindroculum Kimmins 1956. Diagnosis. (1) Viviparous (Kluge 2016: figs 38–42, 76–79). The same in Cloeon s. str. and some other mayflies, but not in other Procloeon /g1. (2) Pterostigma normally either with 1 cross vein (Fig. 40), or with 2 cross veins (Fig. 104) (on individual wings this number varies from 1 to 3) (Figs 40–41). (3) Larval femur with two apical spine-like setae (Figs 17, 81, 110). The same in many other taxa of Baetidae, but not in other Procloeon /g1 and not in other Cloeon /fg1. Larvae of all Oculogaster have a similar structure of the labium, with a pair of submedian setae distinguishable among other setae of dorsal surface (Figs 50, 113). Mandibles of the « Baetis - type » (Kluge 2020a: table 1 and p. 578), i.e. with incisor and kinetodontium fused, left prostheca wide, and all denticles of incisor, kinetodontium, prostheca and mola visible in one view (Figs 51–52, 114–115). All species of Oculogaster lack hind wings, and larvae have no vestiges of hind protoptera (Müller-Liebenau & Hubbard 1985: fig. 9h). Distribution. Paleotropical (Afrotropical and Oriental Regions). Discussion. In the previous diagnosis, the number of cross veins in pterostigma was reported as one, with individual variability; this diagnosis was based on P. (O.) cylindroculum and P. (O.) album, whose pterostigma normally has one cross vein. Examination of P. regularum reveals that it should be placed in Oculogaster, while its pterostigma normally has 2 cross veins; this fact expands the diagnosis of Oculogaster. While pterostigma with one cross vein is not found in any taxa other than Oculogaster, pterostigma with two veins occurs in some other taxa, particularly in some species of Cloeon s. str., which were confused with species belonging to Oculogaster. Species composition and classification. Oculogaster can be divided into two species groups: the African group, which includes P. (O.) cylindroculum Kimmins 1956, P. (O.) barnardi sp. n., P. (O.) niger sp. n., P. (O.) sp. «Wolfkloof» and presumably P. (O.) silvicola Gillies 1997, and the Asian group, which includes P. (O.) album Kluge 2016, P. (O.) regularum Müller-Liebenau & Hubbard 1985, P. (O.) malabarensis sp. n., presumably P. julia (Gillies 1949) comb. n. and unnamed P. sp. «Thailand-3», P. sp. «Sulawesi» and P. sp. « Philippines ».Published as part of Kluge, Nikita J., 2020, Review of Oculogaster Kluge 2016 (Ephemeroptera, Baetidae, Procloeon Bengtsson 1915), pp. 401-437 in Zootaxa 4820 (3) on pages 402-403, DOI: 10.11646/zootaxa.4820.3.1, http://zenodo.org/record/439786
Securiops : Kluge 2020
No full textAutapomorphies of Securiops Larva is psammophylous, with adaptation for inhabitancy on sandy ground: legs are long and slender, with long claws (Figs 28, 67, 112–114); mouth apparatus is modified, with labium greatly elongated; labium, maxillae and labrum having peculiar shape and setation: Labrum (Figs 31–33, 100). Distal margin is nearly straight (neither sharply concave, nor sharply convex); median incision is wide and shallow; other distal margin is distinctly differentiated into paired submedian and paired lateral portions. The submedian portion has a groove bordered by a more prominent ventral flange and a less prominent dorsal flange; the regular row of marginal setae is attached inside this groove, between these two flanges. The lateral portion has no groove, and the regular row of marginal setae is attached openly on its distal margin. This structure of distal margin is different both from sharply concave in Pseudocentroptiloides, and from convex or straight with narrow median incision in other Procloeon /g1 and most other Baetidae (Kluge 2016b: fig. 43; Kluge 2020a: fig. 16; Kluge 2020b: figs 49–50). Maxillae (Figs 36–38). The 2nd and the 3rd dentisetae are located at a distance from the 1st dentiseta, nearer to the proximal angle of biting margin; a few small setae are located between the 1st and the 2nd dentiseta. In other respects, maxilla retains plesiomorphic structure: apex is pointed, with 3 long, slender, arched maxillary canines; the 1st dentiseta has the same shape as the canines and is pressed to the canines. In other Procloeon s. l., the 2nd and the 3rd dentisetae are located close to the 1st dentiseta (e.g., Kluge 2016b: fig. 46; Kluge 2020a: fig. 17; Kluge 2020b: fig. 53). Labium (Figs 40–46, 116–119). Paraglossae are greatly enlarged, three times longer than glossae; glossae and paraglossae are highly modified, with shape and setation different from the initial one (see below); labial palp is greatly widened apically (see below). Glossa is short, with apex rounded. Ventral side of glossa proximally with a regular transverse row of very long and straight setae; being pressed on slide, these setae can be turned either proximally (Fig. 43), or distally (Fig. 41, 42, 45). Dorsal side of glossa distally with a regular, arched, transverse row of stout, pointed setae directed distally (Fig. 46). Rounded apex of glossa is smooth, without setae. Glossa of Securiops differs from the initial one (see Kluge 2016a: 137–138 and figs 4–5; Kluge 2016b: figs 47–48), in that its apex is not pointed and lacks setae, but the marginal setal row is shifted to the dorsal side. Origin of the peculiar transverse ventral-proximal row of straight setae is unclear. Paraglossae are very large and flattened, either oval (Figs 40–43), or with inner margins angulate and diverging (Fig. 116). Most surface of ventral and dorsal sides is smooth, lacking setae; lateral margin with a regular longitudinal row of long setae; median side with two regular longitudinal rows of larger setae and one longitudinal row of smaller setae dorsad of them. Paraglossa of Securiops differs from the initial one (see Kluge 2016a: 137–138 and figs 4–5; Kluge 2016b: figs 47–48), in that its apex is not pointed, and median margin is not evenly concave; possibly, two of the three regular setal rows along the median margin are homologous to the ventro-median and the dorso-median setal rows of the initial structure, and the setae on lateral margin correspond to the latero-apical setae of the initial structure. Labial palp (with 3rd segment initially truncate) has 3rd+2nd segments greatly widened, triangular, with straight distal margin of the 3rd segment exceeding median margin of the 3rd+2nd segments; angle formed by lateral and distal margins is bent and stretched forming a small projection (Fig. 44, 116–117). Authors of the previous publications (Gillies 1988, Lugo-Ortiz & McCafferty 1996, Kaltenbach et al. 2023) erroneously regarded labial palp of Securiops as 2-segmented taking the 2nd and the 3rd segments as one; actually, border between these segments is retained, as well as the muscle going from the 2nd segment to the base of 3rd segment (Figs 40, 44).Published as part of Kluge, Nikita J., 2023, Redescription of the subgenus Securiops Jacobus, McCafferty & Gattolliat 2006 (Ephemeroptera, Baetidae, Procloeon Bengtsson 1915), pp. 243-272 in Zootaxa 5343 (3) on pages 246-249, DOI: 10.11646/zootaxa.5343.3.2, http://zenodo.org/record/833418
Procloeon Kluge, 2020, s.l.
No full textProcloeon /g1, or genus <i>Procloeon</i> Bengtsson 1915 s.l. <p>Branch «11»: Kluge & Novikova 1992: Fig. 11;</p> <p>Procloeon /g1: Kluge 2012: 362;</p> <p> genus <i>Procloeon</i>: Kluge 2016: 494.</p> <p> <b>Type species:</b> <i>Cloeon bifidum</i> Bengtsson 1912.</p> <p> <b>Autapomorphy of Procloeon /g1.</b> At distal part of larval cercus, each cercomere bears apically on its outer (lateral) side one large spine directed posteriorly; usually this spine is spindle-like thickened and its length exceeds length of cercomere (Fig. 7).</p> <p> <b>Composition.</b> The taxon Procloeon /g1 includes the plesiomorphon Procloeon /g2 (or subgenus <i>Procloeon</i>), holophyletic subgenera <i>Oculogaster</i> Kluge 2016, <i>Pseudocentroptiloides</i> Jacob (in Jacob & Glazaczow) 1987, <i>Securiops</i> Jacobus, McCafferty & Gattolliat 2006 and <i>Monilistylus</i> subgen. n. Characters of these taxa are briefly given in the Table 1.</p> <p> <b>Discussion.</b> Among species of the taxon <i>Procloeon</i> s. l. and among its subordinated taxa <i>Procloeon</i> s. str, <i>Pseudocentroptiloides</i> and <i>Securiops</i>, some species have no hind wings, and some species have hind wings of the « <i>Centroptilum</i> - type », i.e. narrow, with two longitudinal veins and with hooked costal process. The same variability occurs within many baetid taxa, so recently presence/absence of hind wings is regarded to be purely a species-specific character. In the past, this character was used as a generic or subgeneric one. Particularly, some authors placed four-winged representatives of <i>Procloeon</i> s.str. in a separate genus or subgenus under the name « <i>Cloeoptilum</i> » (e.g., Kazlauskas 1972) or « <i>Pseudocentroptilum</i> » (e.g., Jacob 1991). The name « <i>Cloeoptilum</i> » remains to be unavailable, because type species have not been formally designated (Hubbard 1979). The name <i>Pseudocentroptilum</i> Bogoescu 1947 belongs to the monotypic taxon opposed to <i>Procloeon</i> s. l. as a whole, while <i>Procloeon</i> s. l. includes both twowinged and four-winged species (Kluge & Novikova 1992; Kluge <i>et al</i>. 2014: 490). Both known species of the new subgenus <i>Monilistylus</i> have no hind wings.</p>Published as part of <i>Kluge, Nikita J., 2020, New subgenus Monilistylus subgen. n. and a new species Procloeon (Monilistylus) ornatipennis sp. n. (Ephemeroptera: Baetidae: Procloeon), pp. 573-587 in Zootaxa 4742 (3)</i> on page 577, DOI: 10.11646/zootaxa.4742.3.11, <a href="http://zenodo.org/record/3677917">http://zenodo.org/record/3677917</a>
Procloeon (Monilistylus) monilistylus monilistylus Kluge, Tiunova & Novikova 2014
No full textProcloeon (Monilistylus) monilistylus Kluge, Tiunova & Novikova 2014 (Figs 1–2, 24–25, 52–54) Cloeon (Centroptilum) sp. 1: Kluge & Novikova 1992: 64 (larva). Cloeon monilistylus: Tiunova 2003: 9 (nomen nudum). Procloeon /g 1 sp.: Kluge 2011: 375. Procloeon monilistylus Kluge, Tiunova & Novikova 2014: 484 (larva, subimago, male imago, female imago, egg). Material examined. Holotype and paratypes (Kluge et al. 2014). Comments. Larva, subimago, male imago, female imago and egg of this species are described in our previous paper (Kluge et al. 2014). In the original description, mandibles were shown only in dorsal view (Kluge et al. 2014: figs 5–6). In such view, shapes of kinetodontia and left prostheca are unclear, because their flatness is perpendicular to the general flatness of the mandible. Here, the left kinetodontium and the left prostheca are shown as they are visible on the mandible, if pressed on a slide (Figs 24–25). The shape of right kinetodontium of P. (M.) monilistylus is the same as in P. (M.) ornatipennis sp. n. (as in Fig. 19). The fine setae of the larval leg were not shown (Kluge et al. 2014: fig. 22). Judging by the setal sockets visible on slide in Canadian balsam, long setae of the tibial base are arranged in the same manner as in P. (M.) monilistylus (as in Fig. 31). Genital muscles of P. (M.) monilistylus were inaccurately figured and wrongly interpreted in the original description. Corrected drawings are given here (Figs 52–53).Published as part of Kluge, Nikita J., 2020, New subgenus Monilistylus subgen. n. and a new species Procloeon (Monilistylus) ornatipennis sp. n. (Ephemeroptera: Baetidae: Procloeon), pp. 573-587 in Zootaxa 4742 (3) on page 580, DOI: 10.11646/zootaxa.4742.3.11, http://zenodo.org/record/367791
Tricorythis tener Kluge, 2016, sp.n.
No full textTricorythis tener sp.n. (Figs 15 –25, 28, 29) Etymology. Tener (Lat.), delicate. Material examined. Holotype: L-S-I♂ {specimen [VI](18) 2014}: ZAMBIA, river Luangwa near Luangwa Bridge, 8.VIII. 2014, coll. N. Kluge & L. Sheyko. Paratypes: the same locality, 2–7.VIII. 2014: 2 L-S-I♂, 2 L-S♂, 4 L-A♀, 30 larvae; river Mutanda at Mutanda Falls 30 km SSW Solwezi, 10–12.VIII. 2014, coll. N. Kluge & L. Sheyko: 2 I ♂; river West Lunga near Mwinilunga, 14–20.VIII. 2014, coll. N. Kluge & L. Sheyko: 1 L-S♂, 3 I ♂; river Zambezi above Victoria Falls, 25–31.VIII. 2014, coll. N. Kluge & L. Sheyko: 73 I ♂, 10 A♀, 5 larvae. Additional material: alate females without eggs, collected at light in localities where both T. furcifer sp.n. and T. tener sp.n. were collected, so systematic position of these females is unclear; these are 73 ♀ alates from river West Lunga and 2 ♀ alates from river Mutanda. Descriptions. Larva. CUTICULAR COLORATION: Cuticle entirely ocher, nearly unicolor, without maculation. HYPODERMAL COLORATION (Figs 17–19): Head and thorax dorsally light ocher with dark gray maculae, thorax ventrally light ocher. Legs light ocher; anterior side of each femur with four dark gray longitudinal stripesstripe on outer margin, stripe on inner margin and two stripes connected distally (Fig. 18), or with fragments of these stripes (Fig. 17). Protoptera different in male and female: in male light ocher or whitish with costal and subcostal fields darker gray (Fig. 17), in female entirely darkened with gray (Figs 18, 19). Abdominal terga from light ocher to dark gray, sterna lighter. Tergalii with dorsal lamellate lobe gray with margins colorless, ventral lobe gray. Caudalii either unicolor light, or with dark maculae in proximal part. SHAPE AND SETATION: Frons and clypeus lacking projections (as in Kluge 2010: Fig. 12). Eyes of male as small as in female. Mandibles with incisors not elongated, curved (as in Kluge 2010: Figs 4–6); left prostheca asymmetrically or symmetrically widened apically, with 3 (occasionally 4) bristle-like processes at base (as in Kluge 2010: Fig. 5); shape of right prostheca usual for Tricorygnatha (as in Kluge 2010: Fig. 6). In last larval instar, transverse row on ventral side of maxilla consists of 9 (occasionally 10 or 11) bristles. Maxilla with long palp. Pronotum moderately wide; anterolateral angles rounded and only slightly projected forward; anterior margin between them in both sexes nearly straight; mesonotum not shortened (Figs 17–19; as in Kluge 2010: Figs 12). Femora moderately wide (Figs 17–19; 23). Stout setae on femora (which form rows characteristic for Pantricorythi) elongate and spatulate (i.e., apically widened and blunt), irregularly arranged: on fore femur they form irregular transverse row, not continuous on outer margin, on middle and hind femora they form longitudinal row on outer margin (Fig. 23; as in Kluge 2010: Figs 7–9). Anterior surface of middle and hind femora with irregularly arranges stout spatulate setae much smaller than spatulate setae on outer margin (Fig. 23). Tibia of each leg with three longitudinal rows of stout setae: on inner-anterior side, on inner side and on inner-posterior side; tibia of middle and hind leg, besides it, with dense regular longitudinal row of spatulate setae on outer-posterior side (Fig. 23). Fore tarsus in male larva not widened, the same as in female larva. Abdominal terga with tergalial cavities not bordered by long setae. Denticles on hind margins of abdominal terga irregular, weak, elongate, either pointed, or blunt, or terminated by several points. Vestiges of tergalii VII absent. Protopenis not large, far not reaching posterior margin of genital plate, cleft up to middle, with paired lobed pressed together (Fig. 25). Subimago, male. CUTICULAR COLORATION AND TEXTURE: Cuticle of pronotum light brown with blanks of composite branched shape; densely covered with microtrichiae except for blanks. Cuticle of pterothorax with sclerites light brown and membranes colorless. Mesonotum with antelateroparapsidal suture and anterolateral scutal costa dark brown; anteronotal protuberance, medioscutum and submedioscutum light brownish; sublateroscutum a little darker; most part of lateroscutum and posterior scutal protuberance nearly colorless (Fig. 20). Microtrichiae densely cover all areas of mesonotum except for posterior scutal protuberances; posterior scutal protuberances without microtrichiae, with net-like relief. Basal plate of wing has distinctly outlined subimaginal sclerite of characteristic shape and brown color (Fig. 20). Cuticle of fore leg partly colorless, with contrasting brown longitudinal stripes (Fig. 21). Cuticle of middle and hind legs either as on fore leg, or entirely colorless. Cuticle of abdominal terga and sterna light brown, with colorless median stripe on terga and small, contrasting, paired blanks; sternum IX has a T-shape blank on anterior margin and midline; gonostyli entirely light (Fig. 22). Cuticle of caudalii colorless. HYPODERMAL COLORATION: As in imago. Imago, male. HYPODERMAL AND CUTICULAR COLORATION: Head and thorax light ocher with dark gray hypodermal maculae; cuticular pigmentation poorly developed. Wings colorless, with proximal parts of costal and subcostal fields tinged with gray. Legs partly colorless, partly with contrasting dark brown hypodermal coloration: femur with longitudinal stripes or/and apical band; tibia with or without dark middle part. Abdomen light ocher, terga tinged with light gray except lateral areas corresponding to larval tergalial cavities. Abdominal sterna, styliger, gonostyli and penis entirely lacking hypodermal and cuticular pigmentation. Caudalii pale ocher with contrasting dark gray proximal half of each segment. SHAPE: Eyes small, as in female (as in Kluge 2010: Fig. 28). Mesonotum with lateroparapsidal suture strongly curved (Fig. 20). Genitals as in Figs 15, 16, 24: Styliger short, with posterior margin either nearly straight (Fig. 16), or shallowly concave (Figs 15, 24) or shallowly convex. Papillae on posterior margin of styliger narrow and sparse. Penis non-sclerotized, narrow, nearly parallel-sided, bifurcates very close to apex, with apices brought together (Fig. 24). Alate female. One alate stage, without molt from subimago to imago (as in other Tricorygnatha). Texture of mesonotum as in male subimago: posterior scutal protuberances without microtrichiae, with net-like relief. Egg. Chorion with characteristic relief (Figs 28, 29). One polar cap, divided to central part and collar. Dimension. Wing length (and approximate body length) of male 4.5 mm, of female 6 mm. Comparison. Male imago and subimago of T. tener sp.n. differ from most other species by narrow penis with distal paired lobes very short and pressed together. Among African species of Tricorythus, similar shape of penis is present in T. discolor (Burmeister 1839), from which T. tener sp.n. differs by small eyes of male. Larva of T. tener sp.n. is very similar to larva of T. varicauda (Pictet 1843), differs by short protopenis of male (Table 1).Published as part of Kluge, Nikita J., 2016, Two new species of Tricorythus Eaton 1868 (Ephemeroptera, Tricorythidae) from Zambia, pp. 273-285 in Zootaxa 4092 (2) on pages 279-282, DOI: 10.11646/zootaxa.4092.2.9, http://zenodo.org/record/26789
FIGURES 52–53 in Occurrence of Anafroptilum Kluge 2012 (Ephemeroptera: Baetidae) in Oriental Region
No full textFIGURES 52–53. Anafroptilum orthostylus sp. n., eggs.Published as part of Kluge, Nikita J. & Novikova, Eugenia A., 2017, Occurrence of Anafroptilum Kluge 2012 (Ephemeroptera: Baetidae) in Oriental Region, pp. 453-472 in Zootaxa 4282 (3) on page 461, DOI: 10.11646/zootaxa.4282.3.2, http://zenodo.org/record/82788
Procloeon (Monilistylus) Kluge 2020, subgen. n.
No full textMonilistylus subgen. n. (Figs 1–54) Type species: Procloeon monilistylus Kluge, Tiunova & Novikova 2014. Autapomorphies of Monilistylus. (1) In male imago, sterno-styligeral muscle (initially unpaired) has enlarged paired lateral portions, which are attached to the bases of unistyligers (Figs 51, 53), so that each lateral portion can serve as adductor of the unistyliger. In our previous paper (Kluge et al. 2014) the lateral portions of the sterno-styligeral muscle were wrongly interpreted as «penial muscles» (Kluge et al. 2014: legend to fig. 25) or as parts of the «penial muscles» (ibid: legend to fig. 31). This error was caused by the fact that in P. (M.) monilistylus the lateral portions of the sterno-styligeral muscle run parallel to the gonovectal (penial) muscles and completely overlap them in ventral view, so that these two pairs of muscles look as one pair (compare left and right halves of Fig. 52). In the new species, P. (M.) ornatipennis sp. n., the lateral portions of the sterno-styligeral muscle are divergent caudally (Fig 51), while the gonovectal (penial) muscles are convergent caudally (Fig. 50), so that in ventral view these two pairs of muscles cross one another, being well distinguishable (Fig. 49). Initially for Ephemeroptera, the sterno-styligeral muscle is unpaired and serves for rotation of the unpaired styliger as a whole to the ventral direction; when this muscle is contracted, the styliger becomes directed perpendicular to the body and the penis becomes protracted (Kluge 2003: figs 3–7; 2004: figs 10–11). In Baetidae, the styliger is divided to a pair of unistyligers separated by a deep incision, and the sterno-styligeral muscle is attached to the integument of this incision (Kluge & Novikova 2011: fig. 3); as a result of this, contraction of the sterno-styligeral muscle leads not to a rotation of the styliger ventrally, but to inclining the unistyligers toward one another. Primitively for Baetidae, the sterno-styligeral muscle is wide and strong, the place of its attachment between unistyligers is sclerotized, and its lateral parts touch the bases of unistyligers (Kluge 2018: fig. 46). In many baetid taxa, the median sclerite between unistyligers is lost; the sterno-styligeral muscle is often weak and unable to move the unistyligers; in selected taxa this muscle is completely lost (e.g., in Labiobaetini Kluge & Novikova 2016 and Rhodobaetis Jacob 2003). In Monilistylus, the sterno-styligeral muscle evolved in opposite direction, so that its lateral portions become strong and look as paired muscles. (2) The second segment of the gonostylus has a composite shape: its proximal 2/3 has a groove on its concave median side; the distal 1/3 has no groove and forms a blunt angle with the grooved proximal portion (Figs 49, 53). Characters of Monilistylus of unclear phylogenetic status. (3) Mandibles of the « Centroptilum - type »: the kinetodontium is deeply separated from the incisor and rotated perpendicular to the plane of the mandible, so that its denticles hide one another in dorsal or ventral view; the left prostheca is bifurcate and rotated in the same manner (Figs 4, 18–25). Among Procloeon /g1, such « Centroptilum - type » of mandibles, besides Monilistylus, occurs in Pseudocentroptiloides and Securiops as well; it is also present in some other baetid taxa (Kluge, in press). In other taxa of Procloeon /g1, the mandibles are of the « Baetis - type »: the kinetodontium is fused with the incisor; the left prostheca is wide, terminated by 3–6 blunt, more distal denticles and 2–4 pointed, more proximal denticles; all denticles of incisor, kinetodontium, prostheca and mola are visible in one view. (4) Larval femur without stout apical setae on outer margin (Figs 30–31) (in contrast to Oculogaster and various taxa not belonging to Cloeon /fg1, which have 2 stout apical setae). Plesiomorphies of Monilistylus. In contrast to Oculogaster, pterostigma with several cross veins (Fig. 44), eggs with sculptured chorion (known for P. (M.) monilistylus only). In contrast to Pseudocentroptiloides, the maxillae are narrowed distally (Fig. 17), the labrum is not deeply emarginated (Fig. 16). In contrast to Pseudocentroptiloides and Securiops, the glossae are not much longer than the paraglossae (Figs 28–29). Distribution. Oriental Region and south-east of Palaearctic. Species composition. Two species: Procloeon (Monilistylus) monilistylus Kluge, Tiunova & Novikova 2014 (south of Russian Far East) and Procloeon (Monilistylus) ornatipennis sp. n. (Indonesia).Published as part of Kluge, Nikita J., 2020, New subgenus Monilistylus subgen. n. and a new species Procloeon (Monilistylus) ornatipennis sp. n. (Ephemeroptera: Baetidae: Procloeon), pp. 573-587 in Zootaxa 4742 (3) on pages 578-579, DOI: 10.11646/zootaxa.4742.3.11, http://zenodo.org/record/367791
Alexander Kluge, « Seize histoires pour Anselm Kiefer »
No full textVersion preprint de Alexander Kluge, « Seize histoires pour Anselm Kiefer ». Traduites de l’allemand par Hilda Inderwildi et Vincent Pauval. Parues dans le catalogue d'exposition Die Ungeborenen. Galerie Thaddaeus Ropac. Octobre 2012.16 histoires dédiées à Anselm Kiefer À propos de son tableau « The Unborn » et à l'occasion de son exposition parisienne chez Thaddeus Ropac du 14 octobre 2012 au 23 février 2013. Traduites par Hilda Inderwildi et Vincent Pauval.« Dans son texte LE NON-NÉ OU LES REPAIRES CÉLESTES D'ANSELM KIEFER (Der Ungeborene oder Die Himmelsareale des Anselm Kiefer), Christoph Ransmayr a décrit les séries peintes par Anselm Kiefer. Il n'y a pas lieu de remettre cela. Mais on peut en reprendre le passage de fin et, avec lui, le concept des non-nés ou de non-naissance. Telle est mon intention en consacrant à Anselm Kiefer cette suite d'histoires au coeur desquelles se situent ses tableaux. » (Alexander Kluge
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