194,297 research outputs found
Ophion kallanderi Johansson & Cederberg 2019, sp. nov.
Ophion kallanderi Johansson sp. nov. urn:lsid:zoobank.org:act: 9D7922E7-EC2C-40EE-8CCD-FAF3739F463E Figs 9I, 34 Diagnosis Ophion kallanderi Johansson sp. nov., which according to the barcoding results (Fig. 1) belongs to the Ophion parvulus aggregate, is one of the trickier Ophion species to identify. It is superficially similar to the species around Ophion slaviceki, but is easily distinguished by the shorter hind trochantellus, the right angled gap between the mandibular teeth, the lower number of flagellomeres and the strongly elongate second flagellomere. The latter character is only otherwise present in Ophion tenuicornis Johansson sp. nov. and Ophion scutellaris. The primary risk of confusion is with Ophion parvulus, Ophion paraparvulus Johansson sp. nov. and Ophion tenuicornis Johansson sp. nov., because of the very weakly sinuate radius in the female that sometimes can be perceived to be evenly curved. Another feature that is frequently present in O. kallanderi Johansson sp. nov., but frequently missing in the other species of the aggregate, is that the pleurosternal angles are more distinct and often slightly protruding (Fig. 9I). In all known specimens, the posterior transverse carina of the propodeum is widely interrupted or only weakly raised centrally, a feature otherwise only frequently occurring in Ophion paraparvulus Johansson sp. nov. within the O. parvulus aggregate. The best diagnostic character is, however, the shape of the flagellomeres. The second flagellomere is distinctly longer than in O. parvulus and O. paraparvulus Johansson sp. nov. and the apical flagellomeres are stouter than in O. tenuicornis Johansson sp. nov. Another distinguishing feature is the shape of the temples. In O. kallanderi Johansson sp. nov. the head is about 0.4–0.5 times as long as the compound eye in lateral view, while it is narrower, 0.3, in both O. paraparvulus and O. tenuicornis Johansson sp. nov. (Table 1). The few known males have the radius evenly curved but are distinguished from the other species in the O. parvulus aggregate on the shape of the flagellomeres. Etymology This species is named in honor of the Swedish lepidopterologist Clas Källander who by donating a large number of Ophion specimens has contributed greatly to this study. Material examined 14 ♀♀, 1 ♂ (Sweden); 1 ♂ (Great Britain); 5 ♀♀, 3 ♂♂ (Lithuania). Type material Holotype SWEDEN • ♀; Västmanland, Sala, 1.4 km W Sala Station; 59.922° N, 16.571° E; 22 Sep. 1974; S. Johansson leg.; MV-light; MZLU Type no. 6371:1. Paratypes SWEDEN • 1 ♀; same data as for holotype; 21 Sep. 1974; S. Johansson leg., MV-light; MZLU Type no. 6371:2 • 1 ♀; Västmanland, Fläckebo, 0.3 km NE Fläckebo church; 59.889° N, 16.359° E; 18 Sep. 1974; S. Johansson leg.; MV-light; MZLU Type no. 6371:3 • 1 ♀; Dalarna, Säterdalen, Näsåkerspussen; 59.734° N, 17.720° E; 4–26 Aug. 2003; SMTP leg.; Malaise trap in alder swamp wood, (Trap id. 10, coll. ev id 399); NHRS-HEVA000008675 • 2 ♀♀; Gästrikland, Gävle, Grinduga; 60.641° N, 17.299° E; 10–17 Sep. 2013; N. Ryrholm and C. Källander leg.; MV-light trap in mixed woodland/farmland; NHRS-HEVA000008676, NHRS-HEVA000008677 • 1 ♀; Skåne, Kullen, Brunkulla; 56.301° N, 12.455° E; 24 Aug. 1975; C.-H. Lindroth leg.; MV-light; MZLU Type no. 6371:4 • 1 ♀; same data as for preceding; 22 Aug. 1974; C.-H. Lindroth leg.; MV-light; MZLU Type no. 6371:5 • 2 ♀ ♀; same data as for preceding; 28 Aug. 1975; C.-H. Lindroth leg.; MV-light; MZLU Type no. 6371:6 • 1 ♀; same data as for preceding; 31 Aug. 1974; C.-H. Lindroth leg.; MV-light; MZLU Type no. 6371:7 • 1 ♂; same data as for preceding; 28 Aug. 1974; C.-H. Lindroth leg.; MV-light; MZLU Type no. 6371:8. Description Fore wing length 15 mm. Antenna with 47–50 flagellomeres. First flagellomere about 4.0–4.5 times as long as wide. Second flagellomere about 3.0 times as long as wide. Central flagellomeres about 2.0–2.5 times as long as wide. Subapical flagellomeres approximately 1.6–1.7 times as long as wide. Head relatively narrowed behind eyes, in lateral view temple 0.4–0.5 times as long as compound eye. Lateral ocellus touching compound eyes. In female the ocelli rather small, not covering the inner margin of compound eye in dorsal view. The distance between lateral ocelli about 0.5–0.6 times as wide as ocellus. Malar space about 0.2 times as long as mandibular base in female and about 0.4 times in male. Mandibular gape right-angled, with internal angles. Wing membrane clear. Ramellus usually quite short, reaching 0.2–0.4 times the width of the discosubmarginal cell. Radius weakly sinuate, sometimes even more or less straight in the basal part in the female and evenly curved in the male. Structure of mesopleuron shining or weakly shagreened with weaker, very regular punctation consisting of small punctures. Interstices between punctures about equal to their diameter up to two times their diameter. Epicnemial carina, in antero-ventral view, with pleurosternal angles obviously anterior to sternal angles. Pleurosternal angles acute, rarely right angled (Fig. 9I). Scutellum without lateral carinae (as in Fig. 6A). Structure of propodeum similar to that of O. obscuratus, posterior to anterior transverse carina mostly shining. Anterior transverse carina complete, quite strongly raised. Posterior transverse carina usually only present laterally, widely interrupted centrally. Sclerotised part of first sternite ending level to spiracle. Central longitudinal carinae weak or absent. Hind trochantellus shorter than wide in dorsal view. Legs normal with hind femur 7.0 times as long as wide. Hind trochantellus shorter than in other similar species. Inner spur of hind tibia short, about 0.3 times as long as hind metatarsus. First sternite ending in level with spiracle (as in Fig. 6H). Colour Body testaceous, inner and outer eye margins yellow. Mandibular teeth black. Ovipositor sheath testaceous. DNA barcode The DNA barcode sequences of one Swedish specimens of Ophion kallanderi Johansson sp. nov. is available at the BOLD systems database (www.boldsystems.org, Specimen code: STI-NJBC: 262). Ecology All specimens have been collected in woodlands during August–September. Distribution in Sweden Rare but probably overlooked in Central Sweden. Older records from Southern Sweden (MZLU) indicates a wider distribution. Remarks Despite the slightly sinuate radius and the more pronounced pleurosternal angles, this species is most likely a member of the O. parvulus aggregate. However the barcode sequence obtained was quite weak and the more detailed phylogenetic position is somewhat uncertain.Published as part of Johansson, Niklas & Cederberg, Björn, 2019, Review of the Swedish species of Ophion (Hymenoptera: Ichneumonidae: Ophioninae), with the description of 18 new species and an illustrated key to Swedish species, pp. 1-136 in European Journal of Taxonomy 550 on pages 70-72, DOI: 10.5852/ejt.2019.550, http://zenodo.org/record/347640
Ophion paraparvulus Johansson & Cederberg 2019, sp. nov.
Ophion paraparvulus Johansson sp. nov. urn:lsid:zoobank.org:act: 27180696-5ED4-4907-9603-0584ACEE7B66 Figs 8B, 38 Diagnosis Ophion paraparvulus Johansson sp. nov. is one of the species in an aggregate previously treated under the name Ophion parvulus. Ophion paraparvulus Johansson sp. nov. is morphologically intermediate between Ophion parvulus and Ophion tenuicornis Johansson sp. nov. and has the flagellomeres stouter and less pilose than O. tenuicornis Johansson sp. nov., but longer and more pilose than O. parvulus. It is also easily confused with O. kallanderi Johansson sp. nov., but distinguished by the shorter temples and the stouter central flagellomeres. The posterior transverse carina is, as in O. kallanderi, usually widely interrupted in the middle with only the lateral parts distinct (Table 1). Etymology The species is very similar and closely related to Ophion parvulus. Material examined 17 ♀♀ (Sweden); 2 ♀♀ (Estonia); 1 ♀ (Norway). Type material Holotype SWEDEN • ♀; Blekinge, Karlskrona, Tullaretorpet; 56.227° N, 15.647° E; 13–14 Jun. 2016; C. Philipsson leg.; MV-light trap in oak dominated deciduous forest; STI-NJBC40; NHRS-HEVA000008694. Paratypes SWEDEN • 1 ♀; same data as for holotype; 20–21 Jun. 2016; C. Philipsson leg.; MV-light trap; STI- NJBC36; NHRS-HEVA000008695 • 1 ♀; Bohuslän, Tossene, Stora Hultet; 58.446° N, 11.409° E; 31 May–7 Jun. 2016; N. Ryrholm and C. Källander leg.; MV-light trap in farmland; NHRS- HEVA000008696 • 1 ♀; Småland, Nybro, Bjällingsmåla; 56.930° N, 15.909° E; 1 May–1 Jul. 2015; N. Johansson leg.; Malaise trap in old mixed forest; NHRS-HEVA000008697 • 1 ♀; Blekinge, Svängsta, Bökemåla (Bökasmåla); 56.241° N, 14.736° E; 5–7 Jul. 2015; M. Sjödahl leg.; MV-light; NHRS- HEVA000008698 • 1 ♀; Skåne, Kullen, Brunkulla; 56.301° N, 12.455° E; 16 Jul. 1974; C.-H. Lindroth leg.; MV-light; MZLU Type no. 6373:1 • 1 ♀; Östergötland, Boxholm, Björneberg; 58.195° N, 14.909° E; 4–20 Jun. 2018; N. Johansson leg.; Malaise trap in old mixed forest; NHRS-HEVA000008699 • 1 ♀; Bohuslän, Uddevalla, Älje-Porsen; 58.355° N, 12.031° E; 5 Jun. 2017; M. Oomen leg.; MV-light trap; NHRS-HEVA000008700 • 1 ♀; Skåne, Kullen, Brunkulla; 56.301° N, 12.455° E; 20 Jun. 1973; C.-H. Lindroth leg.; MV-light; MZLU Type no. 6373:2. Description Fore wing length 16 mm. Antenna with 53–54 flagellomeres. First flagellomere 4.0 times as long as wide.Second–fourth flagellomere about 2.5–3.0 times as long as wide. Central flagellomeres elongate, about 2.0 times as long as wide. Apical flagellomeres 1.8–1.9 times as long as wide. Flagellomeres with quite long prominent pilosity. Length of pilosity in females at least 0.5 width of flagellomere (Fig. 8B). Head narrow behind eyes, in lateral view with temple 0.3 times as long as compound eye. Ocelli in female large, in dorsal view covering the inner margin of compound eye. The distance between lateral ocelli in female about 0.3 times the diameter of the ocellus. Malar space about 0.1 times as long as mandibular base. Mandibular gape right-angled, with internal angles. Wing membrane clear. Ramellus distinct, reaching 0.2–0.4 the width of discosubmarginal cell but sometimes small or absent. Radius evenly curved. Mesopleuron weakly shagreened with deep distinct punctures. Interstices between punctures about equal to their diameter. Epicnemial carina, in antero-ventral view, with pleurosternal angles obviously anterior to sternal angles. Pleurosternal angles rounded, slightly obtuse. Scutellum with lateral carinae only indicated basally. Propodeum with very weak rugose structure, shining with anterior transverse carina strongly raised. Posterior transverse carina usually widely interrupted centrally. Longitudinal carinae delimiting area superomedia and lateral longitudinal carinae often weak or absent, but normally clearly indicated at the junction with the posterior transverse carina. Hind trochantellus shorter than wide in dorsal view. Legs normal with hind femur about 6.0–7.0 times as long as wide. Sclerotised part of first sternite ending level or slightly posterior to spiracle. Inner spur of hind tibia as long as 0.4 times first hind metatarsus. Male unknown. Colour Body light testaceous, frequently with diffuse pale markings as in O. parvulus. Mandibular teeth black. Ovipositor sheath usually concolourous with posterior metasomal segments. DNA barcode The DNA barcode sequences of two Swedish specimens of Ophion paraparvulus Johansson sp. nov. are available at the BOLD systems database (www.boldsystems.org, BIN. BOLD: ADG1399. Specimen codes: STI-NJBC: 36, 40). Ecology Very little is known about the biology of this species. It has been collected in areas with mature semiopen mixed forests. The flight period seems to be mid June to mid July. It is active earlier in the season than its sibling species O. tenuicornis Johansson sp. nov. and O. costatus. Distribution in Sweden Rare and possibly confined to areas with favourable climate in Southern Sweden.Published as part of Johansson, Niklas & Cederberg, Björn, 2019, Review of the Swedish species of Ophion (Hymenoptera: Ichneumonidae: Ophioninae), with the description of 18 new species and an illustrated key to Swedish species, pp. 1-136 in European Journal of Taxonomy 550 on pages 86-88, DOI: 10.5852/ejt.2019.550, http://zenodo.org/record/347640
Enicospilus intermedius Johansson 2018
Enicospilus intermedius Johansson, 2018 Enicospilus intermedius Johansson, 2018: 9–12; Figs 8, 9A–C. Holotype ♀ from Sweden in NHRS, examined. Material examined: IRAN: 1♂, East Azerbaijan Province, Sharghi, Khodaafarin, Jananlou, 20 May, 2004, 360 m. a. s., Gh. / Gil. leg. (HMIM); 1♀, Golestan Province, Elburs mts, Maghazy Valley, Lux, 36˚64’92.3"N, 54˚20’70.1"E, 11 Jun. 2007 1630 m. a. s., N. Pöll leg. (OÖLM); 1♀, Kerman Province, 25 km E. of Jiroft (Mijan), 28˚42"N, 57˚57"E, 27 May 2014, Mi. Halada leg. (OÖLM); 1♀, Khorasan Province, 10 km W. Raz, Koppe Dag, 37˚54"N, 56˚55"E, 1200 m. a. s., Mi. Halada leg. (OÖLM). Remarks: Described from Southern Sweden in 2018 (Johansson 2018) this species have shown to be possibly quite common in the lowlands of Central Europe. The species is similar to Enicospilus adustus but distinguished by the wider face and the usually entirely testaceous head, without any extensive yellow markings around the compound eyes.Published as part of Johansson, Niklas, Ameri, Ali, Riedel, Matthias, Talebi, Ali Asghar & Ebrahimi, Ebrahim, 2021, Contribution to the Ophioninae (Hymenoptera: Ichneumonidae) of Iran with the description of 16 new species and an illustrated key to the Eremotylus of the Western Palaearctic, pp. 151-206 in Zootaxa 5023 (2) on pages 161-162, DOI: 10.11646/zootaxa.5023.2.1, http://zenodo.org/record/522565
Ophion norei Johansson & Cederberg 2019, sp. nov.
Ophion norei Johansson sp. nov. urn:lsid:zoobank.org:act: 5425028E-9DCF-4D80-99F9-59ECDA17AC00 Figs 17D, F, 20E, 37 Diagnosis Most closely related and similar to Ophion perkinsi, from which it is distinguished by the more transverse head in anterior view, shorter malar space, the weaker microsculpture of the meso- and metapleuron and the stouter first tergite, usually lacking a median dorsal undulation. Also very similar to Ophion ellenae Johansson sp. nov. and Ophion matti Johansson sp. nov., from which it is separated by fewer flagellomeres, the more buccate temple, the head in dorsal view having a distinct gap between the eyes and lateral ocellus, the more distinctly punctate propodeum and the usually weaker carination of the propodeum. Also very similar to Ophion paukkuneni Johansson sp. nov., but with shorter flagellomeres, slightly less buccate temples and no dorsal undulation on the first tergite. Etymology Named after the son of the first author, Nore Nystedt. Material examined 8 ♀♀, 1 ♂ (Sweden). Type material Holotype SWEDEN • ♀; Öland, Mörbylånga, Hönstorp; 56.662° N, 16.554° E; 10 Jul. 2018; K. Alexandersson leg.; MV-light in garden; NHRS-HEVA000008685. Paratypes SWEDEN • 1 ♀; same data as for holotype; NHRS-HEVA000008686 • 1 ♂; Skåne, Ystad, Kåseberga; 55.385° N, 14.066° E; 24 May–28 Jul. 2016; N. Ryrholm and C. Källander leg.; MV-light trap in warm coastal sandslopes; STI-NJBC074; NHRS-HEVA000008687 • 2 ♀♀; same data as for preceding; STI-NJBC116, STI-NJBC223; NHRS-HEVA000008688, NHRS-HEVA000008689 • 2 ♀♀; Skåne, Ystad, Kåseberga; 55.385° N, 14.066° E; 17 Jul.–14 Sep. 2013; N. Ryrholm and C. Källander leg.; MV-light trap in warm coastal sandslopes; 1 ♀; STI-NJBC228; NHRS-HEVA000008690, NHRS- HEVA000008691 • 1 ♀; Gotland, Sundre, Hallbjäns; 56.938° N, 18.146° E; 21 Jun.–22 Jul. 2008; N. Ryrholm and C. Källander leg.; MV-light trap in rocky calcareous coastal heath; STI-NJBC225; NHRS-HEVA000008692 • 1 ♀; Skåne, Ystad, Kåseberga; 55.385° N, 14.066° E; 29 Jul.–10 Oct. 2016; N. Ryrholm and C. Källander leg.; MV-light trap in warm coastal sandslopes; STI-NJBC226; NHRS- HEVA000008693 • 1 ♀; Öland, Mörbylånga, Räpplinge; 56.827° N, 16.660° E; 26 Jul.1980; L.-Å. Janzon leg.; MV-light; MZLU Type no. 6372:1. Description Fore wing length 12–13 mm. Antenna in both sexes with 48–50 flagellomeres. First flagellomere 3.0–3.5 times as long as wide. Central flagellomeres almost square to slightly longer than wide. Head buccate behind eyes, in dorsal view with distance between inner margin of compound eye and lateral ocellus 0.2–0.3 times diameter of ocellus. Head in anterior view strongly transverse (Fig. 17D, F). Temple in lateral view about 0.7–0.8 times as long as compound eye (as in Fig. 21D). Malar space short, about 0.2 times the basal width of the mandible (Fig. 17D, F). Mandibular gape right-angled, with internal angles. Wing membrane clear. Ramellus distinct. Radius sinuous. Mesopleuron and metapleuron punctate with microsculpture weak or absent. Interstices between punctures about equal to their diameter. Pleurosternal angles rounded, often sharp, well defined and slightly anterior to sternal angles. Scutellum with distinct lateral carinae at least in basal half. Propodeum coriaceous and distinctly punctate. Anterior and transverse carina present centrally, often weak or absent laterally. Posterior transverse carina strongly raised, interrupted centrally. Longitudinal carinae delimiting area superomedia sometimes weak or absent (as in Fig. 10H). Area superomedia usually as long as wide. Hind trochantellus shorter than wide in dorsal view. First tergite in lateral view rather stout without or with only a shallow dorsal undulation centrally (Fig. 20E). Sclerotised part of first sternite ending distinctly posterior to spiracle. Inner spur of hind tibia as long as 0.5 times metatarsus. Colour Body testaceous. Mandibular teeth black. Inner and outer orbits slightly yellowish. Ovipositor sheath testaceous. DNA barcode The DNA barcode sequences of six Swedish specimens of Ophion norei Johansson sp. nov. are available at the BOLD systems database (www.boldsystems.org, BIN. BOLD: ADF8593. Specimen codes: STI- NJBC: 74, 116, 223, 225–226, 228). Ecology The species is active from late June to early September and occurs in open or semi-open grasslands in Southern Sweden. Distribution in Sweden Rare in Southern Sweden in areas with dry and hot climate.Published as part of Johansson, Niklas & Cederberg, Björn, 2019, Review of the Swedish species of Ophion (Hymenoptera: Ichneumonidae: Ophioninae), with the description of 18 new species and an illustrated key to Swedish species, pp. 1-136 in European Journal of Taxonomy 550 on pages 81-83, DOI: 10.5852/ejt.2019.550, http://zenodo.org/record/347640
Ophion ellenae Johansson & Cederberg 2019, sp. nov.
Ophion ellenae Johansson sp. nov. urn:lsid:zoobank.org:act: 66D1B37B-DDE5-44DA-8B9E-968A65 D91769 Figs 8D, 10G, 18F, 21A, C, 32 Diagnosis Superficially similar to Ophion inclinans Johansson sp. nov. and O. arenarius Johansson sp. nov., but never with the dorsal undulation on the first tergite. Usually more densely punctured face below the antennal sockets, more prominent and sharper pleurosternal angles and stouter flagellomeres. Closely related and very similar to O. perkinsi and O. norei Johansson sp. nov., but with the head more narrowed behind the eyes, no gap between the eye and lateral ocellus, complete anterior transverse carina of the propodeum and more numerous flagellomeres. Most closely related, however, to O. matti Johansson sp. nov., but distinguished by the more shining and less densely punctate mesoscutum, the shorter area superomedia and the more elongate basal flagellomeres. Etymology The species is named in honour of the authors’ wife, Ellen Nystedt. Material examined 30 ♀♀, 1 ♂ (Sweden); 2 ♀♀ (Finland); 1 ♀ (Estonia); 1 ♀ (Lithuania). Type material Holotype SWEDEN • ♀; Närke, Lerbäck, Hugghult; 58.949° N, 15.045° E; 15 Aug. 2003; A. Larsson leg.; Sweepnet; STI-NJBC188; NHRS-HEVA000008654. Paratypes SWEDEN • 1 ♀; Gotland, Sundre, Suders; 56.945° N, 18.303° E; 14 Jul.–17 Aug. 2017; N. Ryrholm and C. Källander leg.; MV-light trap; NHRS-HEVA000008655 • 2 ♀♀, 1 ♂; Gotland, Sundre, Suders; 56.945° N, 18.303° E; 29 Jul.–18 Sep. 2017; N. Ryrholm and C. Källander leg.; MV-light trap in deciduous woodland; STI-NJBC307, STI-NJBC301; NHRS-HEVA000008656 to NHRS- HEVA000008658 • 1 ♀; Gotland, Tofta, Nasume myr; 57.537° N, 18.129° E; 23 Jul. 2016; J. Törnvall leg.; MV-light by mire; NHRS-HEVA000008659 • 1 ♀; Skåne, Klippan, Bonnarpshed; 56.087° N, 13.176° E; 5–30 Aug. 2007; N. Ryrholm and C. Källander leg.; MV-light trap in open grazed heathland; STI-NJBC87; NHRS-HEVA000008660 • 1 ♀; Gotland, Sundre, Barrshage; 56.922° N, 18.186° E; 28 Aug.–22 Sep. 2017; N. Ryrholm and C. Källander leg.; MV-light trap in wet meadows surrounded by deciduous forest; NHRS-HEVA000008661 • 1 ♀; Öland, Borgholm, Halltorps hage; 56.794° N, 16.573° E; 24–30 Jul. 1938; N. Bruce leg.; NHRS-HEVA000008662 • 1 ♀; same data as for preceding; 1–5 Aug. 1939; N. Bruce leg.; NHRS-HEVA000008663 • 1 ♀; Uppland, Rådmansö, Bergholmen; 59.708° N, 18.955° E; 29 Jul.–18 Sep. 2017; N. Ryrholm and C. Källander leg.; MV-light trap in coastal mixed forest; NHRS-HEVA000008664 • 1 ♀; Öland, Mörbylånga, Räpplinge; 56.827° N, 16.660° E; 27 Jul.1975; L.-Å. Janzon leg.; MV-light; MZLU Type no. 6370:1 • 4 ♀♀; same data as for preceding; 26 Jul. 1980; L.-Å. Janzon leg.; MV-light; MZLU Type no. 6370:2 to MZLU Type no. 6370:5 • 1 ♀; Gotland, Hamra, Långmyre; 56.962° N, 18.292° E; 2–25 Aug. 1997; N. Ryrholm and C. Källander leg.; MV-light trap; STI-NJBC305; NHRS-HEVA000008665 • 1 ♀; Gotland, Hamra, Tuvlandet; 56.966° N, 18.308° E; 15 Jul.–18 Aug. 2017; N. Ryrholm and C. Källander leg. MV-light trap in abandoned farmland; STI-NJBC306; NHRS-HEVA000008666. Description Fore wing length 15–16 mm. Antenna with 52–54 flagellomeres. First flagellomere 3.5 times as long as wide. Second flagellomere about 2.2 times as long as wide. Central flagellomeres stout, about 1.2–1.3 times as long as wide. Subapical flagellomeres approximately 1.5 times as long as wide (Fig. 8D). Temple relatively short. Head in lateral view with temple 0.4–0.5 times as long as compound eye (Fig. 21C). Gap between compound eye and lateral ocellus 0.1 times the diameter of ocellus. Face below antennal sockets with dense punctures. Interstices between punctures 0.5 times their diameter. Malar space about 0.1–0.2 times as long as mandibular base in female and male. Mandibular gape rightangled, with internal angles. Wing membrane clear. Ramellus short, reaching 0.2–0.4 times the width of the discosubmarginal cell. Radius sinuous. Mesoscutum shining with irregular and weak punctures, interstices between punctures about equal to their diameter (Fig. 21A). Mesopleuron shagreened with deep, dense punctures, space between punctures 0.5 times their diameter. Epicnemial carina, in antero- ventral view, with pleurosternal angles almost in level with sternal angles, pleurosternal angles right angled. Scutellum quite wide with distinct lateral carinae in basal 0.8–0.9 (as in Fig. 6C). Propodeum with very weak rugose structure, shining with anterior and posterior transverse carina often strongly raised. Central longitudinal carinae strong, lateral longitudinal carina absent. Area superomedia relatively short, almost square (Figs 10G, 18F). Hind trochantellus shorter than wide in dorsal view. Legs normal with hind femur about 6.0 times as long as wide. Sclerotised part of first sternite ending distinctly posterior to spiracle at a distance equal to that between hind margin of the sclerotised part of the first sternite and hind margin of the first tergite. Inner spur of hind tibia as long as 0.5 times metatarsus. Colour Body testaceous. Mandibular teeth black. Head with inner and outer orbits yellow. Ovipositor sheath testaceous. DNA barcode The DNA barcode sequences of six Swedish specimens of Ophion ellenae Johansson sp. nov. are available at the BOLD systems database (www.boldsystems.org, BIN. BOLD: ADM4635. Specimen codes: STI-NJBC: 87, 188, 301, 305–307). Ecology One male and one female kept in NHRS (Sweden, Upl, Sandhamn, Ljungdahl leg.) have been reared from the rare moth Hadena bicruris (Hufnagel, 1766). The available data for the parasitoid suggests a connection to open dry, sandy or rocky grassland, which also is the main habitat of the known host and some of its close relatives. Ophion ellenae Johansson sp. nov. is active from late July to early September. Distribution in Sweden Most frequently collected in coastal areas of the eastern parts of Central Sweden where it can be locally abundant. Older records are known from a wider area in Central Sweden (Uppland, Västmanland and Dalsland) indicating a decline during the last century. Remarks This species is very closely related to O. matti Johansson sp. nov. (Fig. 4). The two species are very similar and also share the same BIN. The slight but distinct morphological differences, supported by the barcoding results motivates the description of a separate species.Published as part of Johansson, Niklas & Cederberg, Björn, 2019, Review of the Swedish species of Ophion (Hymenoptera: Ichneumonidae: Ophioninae), with the description of 18 new species and an illustrated key to Swedish species, pp. 1-136 in European Journal of Taxonomy 550 on pages 64-66, DOI: 10.5852/ejt.2019.550, http://zenodo.org/record/347640
Ophion angularis Johansson & Cederberg 2019, sp. nov.
Ophion angularis Johansson & Cederberg sp. nov. urn:lsid:zoobank.org:act: BB40BE24-344A-40B6-AF63-078ED2FFCDB3 Figs 9F, 22 Diagnosis The combination of the shape of the pleurosternal angles, the buccate temples and the number of flagellomeres separates this species from all other known Swedish species of Ophion. Ophion angularis Johansson & Cederberg sp. nov. is most likely to be confused with Ophion crassicornis Brock, 1982 and Ophion borealis Johansson sp. nov., but has more prominent pleurosternal angles, usually more numerous flagellomeres, slightly less buccate temples, entirely uncarinated scutellum and more densely punctate face. Etymology The pleurosternal angles are very prominent in this species. Material examined 27 ♀♀, 10 ♂♂ (Sweden); 2 ♀♀ (France); 7 ♂♂ (Estonia). Type material Holotype SWEDEN • ♀; Östergötland, Norrköping, Kimstad; 58.551° N, 15.962° E; 1 May–1 Jul. 2012; M. Stahre leg.; Yellow pan trap in gravelpit close to pine and oak forest; STI-NJBC79; NHRS- HEVA000008581. Paratypes SWEDEN • 1 ♀; Öland, Borgholm, 0.6 km SW Borgholm church; 56.876° N, 16.648° E; 22 May 1974; S. Johansson leg.; MV-light in garden surrounded by oak forest; MZLU Type no. 6366:1 • 1 ♀; Öland, Mörbylånga, Räpplinge; 56.827° N, 16.660° E; 18 Jun. 1964; S. Johansson leg.; MV-light; MZLU Type no. 6366:2 • 1 ♀; Skåne, Ystad, Kåseberga; 55.385° N, 14.066° E; 21 May–28 July 2016; N. Ryrholm and C. Källander leg.; MV-light trap in warm coastal sandslopes; NHRS-HEVA000008582 • 1 ♀; Skåne, Simrishamn, Örnahusen; 55.450° N, 14.261° E; 4–26 Jun. 2006; N. Ryrholm and C. Källander leg.; MVlight trap in coastal meadow; NHRS-HEVA000008583 • 2 ♀♀; Blekinge, Karlskrona, Tullaretorpet; 56.227° N, 15.647° E; 2–3 Jun. 2016; C. Philipsson leg.; MV-light in deciduous forest; 1 ♀ STI-NJBC78; NHRS-HEVA000008584, NHRS-HEVA000008585 • 1 ♀; Gotland, Roleks; 57.536° N, 18.339° E; 2–18 Jun. 2004; SMTP leg.; Malaise trap in grazed calcareous pine forest (Trap id 28, coll ev. id. 497); NHRS-HEVA000008586 • 1 ♀; Gotland, Roleks; 10 Apr.–6 Jun. 2005; SMTP leg.; Malaise trap in grazed calcareous pine forest (Trap id 28, coll ev. id. 1464); NHRS-HEVA000008587 • 1 ♂; Öland, Mörbylånga, Strandskogen; 56.702° N, 16.494° E; 22 May 2016; B. Andersson leg.; MV-light trap in garden on sand close to deciduous forest; NHRS-HEVA000008588 • 1 ♀; Skåne, Ystad, Spraggehusen; 55.442° N, 14.066° E; 27 May–28 Jul. 2016; N. Ryrholm and C. Källander leg.; MV-light trap in coastal sanddunes; NHRS-HEVA000008589 • 1 ♀; Gotland, Sundre, Hallbjäns; 56.938° N, 18.146° E; 21 Jun.–22 Jul. 2006; N. Ryrholm and C. Källander leg.; MV-light trap in coastal rocky, calcareous heath; NHRS-HEVA000008590 • 1 ♀; Gotland, Hamra, Tuvlandet; 56.966° N, 18.378° E; 24 Aug.–27 Aug. 2007; N. Ryrholm and C. Källander leg.; MV-light trap in abandoned farmland/gardens; NHRS- HEVA000008591 • 3 ♀♀; same locality as for preceding; 56.966° N, 18.308° E; 26Aug.–24 Sep. 2007; N. Ryrholm and C. Källander leg.; MV-light trap in abandoned farmland/gardens; NHRS-HEVA000008592 to NHRS-HEVA000008594 • 2 ♀♀; Småland, Ljungby, Agunnaryd, Nockarp; 56.767° N, 14.167° E; 10 Jul. 1966; N. Burreau leg.; MZLU Type no. 6366:3, MZLU Type no. 6366:4 • 1 ♂; same data as for preceding; 19 Jun. 1966; N. Burreau leg.; MZLU Type no. 6366:5 • 4 ♂♂, 2 ♀♀; Skåne, Klippan, Bonnarpshed; 56.087° N, 13.176° E; 8–13 Jun. 2007; N. Ryrholm and C. Källander leg.; MV-light trap in grazed heathland; NHRS-HEVA000008595 to NHRS-HEVA000008600 • 2 ♂♂; Skåne, Höganäs, Mölle; 56.289° N, 12.498° E; 1 Jun. 1965; N. Burreau leg.; MZLU Type no. 6366:6, MZLU Type no. 6366:7 • 1 ♂; Halland, Halmstad, Steninge; 56.758° N, 12.637° E; 29 Jun. 1941; B. Hanström leg.; MZLU Type no. 6366:8 • 1 ♀; Uppland, Väddö, Ytterskär; 59.938° N, 18.920° E; 23 May–26 Jun. 2017; N. Ryrholm and C. Källander leg.; MV-light trap; STI-NJBC186; NHRS-HEVA000008601 • 1 ♀; Öland, Mörbylånga, Torslunda Ekologiska Station; 56.619° N, 16.497° E; 5 Jun. 1977; B. Cederberg leg.; Sweepnet in dry grassland, NHRS-HEVA000008602 • 1 ♂; same data as for preceding; 12 Jun. 1977; B. Cederberg leg.; Sweepnet in dry grassland; NHRS-HEVA000008603. Description Fore wing length 15–17 mm. Antenna in both sexes with 59–67 flagellomeres. First flagellomere 2.5–3.0 times as long as wide. Central flagellomeres stout, about 1.2–1.3 times as long as wide. Apical flagellomeres approximately 1.5 times as long as wide. Temple in female and male strongly buccate, in lateral view 0.7–0.8 times as long as compound eye. Head with distinct gap between lateral ocellus and inner margin of compound eye. Face below antennal sockets very densely punctate, interstices about 0.2 times diameter of punctures. Malar space about 0.2 times as long as mandibular base in female and male. Mandibular gape right-angled, with internal angles. Wing membrane clear. Ramellus reaching 0.2–0.3 times the width of the discosubmarginal cell. Radius sinuous. Mesopleuron shagreened with very dense punctures which often merge centrally. Epicnemial carina, in antero-ventral view, with pleurosternal angles nearly in level with sternal angles, pleurosternal angles acute (Fig. 9 F). Epicnemial carina between tip of pleurosternal angle and sternal part strongly concave. Scutellum without distinct lateral carinae in basal half. Propodeum distinctly punctate and coriaceous in front of anterior transverse carina, between transverse carina more often quite shiny, weakly rugose. Anterior transverse carina sometimes absent or weak laterally and posterior transverse often interrupted centrally, in continental specimens the anterior carina sometimes more strongly raised. Longitudinal carinae delimiting area superomedia and petiolar carinae often strong. Area superomedia quite narrow, about two times as long as wide. Hind trochantellus shorter than wide in dorsal view. Legs normal with hind femur about 7.0 times as long as wide. Sclerotised section of first sternite ending level to spiracle (as in Fig. 5 H). Inner spur of hind tibia 0.4 times as long as hind metatarsus. Colour Body testaceous. Mandibular teeth black. Head with inner and outer eye margins widely yellow. Ovipositor sheath black or dark brown, contrasting in colour with posterior metasomal segments. DNA barcode The DNA barcode sequences of four Swedish specimens of Ophion angularis Johansson & Cederberg sp. nov. are available at the BOLD systems database (www.boldsystems.org, BIN. BOLD:ADG2027. Specimen codes: STI-NJBC: 78–79, 186, 267). Ecology The species seems to occur in open or semi-open habitats and is active during late June–August. Other than that, nothing is known about the biology. Distribution in Sweden Rare but widespread in Southern Sweden. Remarks This species has been confused with Ophion crassicornis Brock, 1982 and was also probably partly included in the description of that species. Ophion angularis Cederberg & Johansson sp. nov. is, however, both morphologically and genetically distinct.Published as part of Johansson, Niklas & Cederberg, Björn, 2019, Review of the Swedish species of Ophion (Hymenoptera: Ichneumonidae: Ophioninae), with the description of 18 new species and an illustrated key to Swedish species, pp. 1-136 in European Journal of Taxonomy 550 on pages 37-40, DOI: 10.5852/ejt.2019.550, http://zenodo.org/record/347640
Knowledge sharing strategies for Project Knowledge Management in the automotive sector
Project Knowledge Management is regarded as a field of increasing importance for both researchers and organisations. This led Volvo Technology, the innovation business unit within a Swedish automotive multinational corporation, to explore through a qualitative case study how Project Knowledge Management could be improved to support knowledge sharing between projects within the organisation. The current situation of Project Knowledge Management is described through a developed theoretical framework and with input from thirteen semi-structured interviews conducted and analysed in an iterative fashion. The description shows that the contributor employs the codification strategy to share knowledge with other projects while the receiver adapts a personalisation strategy to retrieve knowledge from other projects. This description was analysed and compared with current research through brainstorming. Since there are current initiatives within the organisation to improve the codification strategy, this research focuses on improving the personalisation strategy. The recommendation was to promote Communities of Practice. Six semi-structured interviews were conducted to evaluate the relevance of the recommendations. The recommendation was found to be relevant for improving Project Knowledge Management within Volvo Technology
Ophion vardali Johansson & Cederberg 2019, sp. nov.
Ophion vardali Johansson sp. nov. urn:lsid:zoobank.org:act: 2395488B-9203-4CB7-BE81-62510D5989BA Figs 19 A–B, 45A–B Diagnosis Similar to Ophion pteridis, but besides the obvious differences in colour, with the head more rounded in anterior view and the face wider in relation to the compound eyes. The occipital carina is centrally raised forming an acute angle, the head is more buccate with a distinct gap between the lateral ocellus and the compound eye and the central flagellomeres are stouter. Also similar to Ophion inclinans Johansson sp. nov. and O. arenarius Johansson sp. nov., but the head has a distinct gap between the lateral ocellus and compound eye, the head in anterior view is more rounded and the face wider in relation to the compound eyes. According to the molecular analysis the species is closely related to Ophion pteridis, but based on the distinct morphological characters, both species are regarded as valid. Etymology Named in honour of Hege Vårdal, curator at the Swedish Museum of Natural History, who has been of great help during this study. Material examined 2 ♀♀, 3 ♂♂ (Sweden); 7 ♀ (Norway). Type material Holotype NORWAY • ♀; Nordland, Bodø, Ausvika; 67.336° N, 14.496° E; 10 Aug. 2018; G. Ørsnes leg.; coastal meadow adjacent to forest; BOLD: DS-ICHNN; NHRS-HEVA000008745. Paratypes SWEDEN • 2 ♀♀, 2 ♂♂; Skåne, Ystad, Järahusen; 55.396° N, 14.207° E; 8 Jul.–28 Aug. 2016; N. Ryrholm and C. Källander leg.; MV-light trap in coastal sanddunes in mixed oak and pine forest; NHRS-HEVA000008746 to NHRS-HEVA000008749 • 1 ♂; Skåne, Kristianstad, Åhus; 55.944° N, 14.262° E; 22–26 Jul. 1964; unknown leg.; MV-light; MZLU Type no. 6375:1. Description Fore wing length 16–17 mm. Antenna in both sexes with 54–55 flagellomeres. First flagellomere 3.5–4.0 times as long as wide. Central flagellomeres about 1.5–1.7 times as long as wide. Head quite narrow behind eyes, with or without small gap between lateral ocellus and compound eye. Temple in lateral view about 0.7–0.8 times as long as compound eye as a result of the very short compound eye in lateral view (Fig. 19B). Head in frontal view rounded (width/height measured from the apical margin of clypeus to the top of head = 1.15) with face wide in relation to compound eyes (Fig. 19A). Head with occipital carina centrally raised forming an acute angle (as in Fig. 7B). Face below antennal sockets with quite scarce punctures, polished or weakly shagreened. Malar space about 0.4 times as long as mandibular base in female and about 0.4–0.5 times in male. Mandibular gape right-angled, with internal angles. Wing membrane yellowish. Ramellus distinct, reaching 0.2–0.3 times the width of the discosubmarginal cell. Radius sinuous. Mesopleuron shagreened and distinctly punctate, spaces between punctures about equal to their diameter. Pleurosternal angles weakly obtuse to right angled, obviously anterior to sternal angles. Scutellum with distinct lateral carinae (as in Fig. 6C). Propodeum weakly shagreened, often quite polished with anterior transverse carina strong, but sometimes partly absent laterally. Posterior transverse carina often widely interrupted centrally. Area superomedia often present. Central longitudinal carinae often distinct. Sclerotised part of first sternite ending obviously posterior to spiracle. First tergite in lateral view with slight or distinct median undulation (as in Fig. 6E). Hind trochantellus shorter than wide in dorsal view. Inner spur of hind tibia as long as 0.5 times hind metatarsus. Colour Body testaceous, the lower part of mesopleuron, mesosternum, propleuron, coxae mostly and propodeum strongly infuscate, black or dark brown (Fig. 45 A–B). Head with inner and outer orbits yellow. Mandibular teeth black. Ovipositor sheath testaceous, concolourous with posterior metasomal segments. DNA barcode The DNA barcode sequence of four specimens of Ophion vardali Johansson sp. nov. is available at the BOLD systems database (www.boldsystems.org, BOLD: DS-ICHNN; STI-NJBC 24–26). Ecology No host records are known for this species and it is active mainly during July. The habitat seems to be open or semi-open coastal meadows. Distribution in Sweden Ophion vardali Johansson sp. nov. is very rare and known from only a couple of localities in the southeastern part of the country. Remarks On the website www.artsobservasjoner.no there are also pictures of specimens from several different coastal localities in Central and Northern Norway. One Norwegian female, kindly provided by Geir Ørsnes, is designated as holotype. The rather vague description of Ophion praecinctus Meyer, 1935 describes a species from the desert country Turkmenistan with similar colouration to O. vardali Johansson sp. nov. The type material is lost (Townes et al. 1965; Andrey Khalaim, ZIN, pers. com.) and there seems to be no remaining additional material determined by the Meyer. The description (Meyer 1935) and a simplified translated key (Meyer 1937) refers to a very small species (body length 12 mm), with transverse head, long malar space, vague notauli, deep concavities along the inner eye margin and two distinct transverse ridges on the propodeum, all of which are characters that cannot be attributed to O. vardali Johansson sp. nov.Published as part of Johansson, Niklas & Cederberg, Björn, 2019, Review of the Swedish species of Ophion (Hymenoptera: Ichneumonidae: Ophioninae), with the description of 18 new species and an illustrated key to Swedish species, pp. 1-136 in European Journal of Taxonomy 550 on pages 106-108, DOI: 10.5852/ejt.2019.550, http://zenodo.org/record/347640
Ophion tenuicornis Johansson & Cederberg 2019, sp. nov.
Ophion tenuicornis Johansson sp. nov. urn:lsid:zoobank.org:act: C1A3799F-344E-49E3-AA66-3083A9D3FF27 Figs 8C, 10L, 13A, 44 Diagnosis Ophion tenuicornis Johansson sp. nov. is one of the species in an aggregate previously treated under the name Ophion parvulus. Ophion tenuicornis Johansson sp. nov. is easily distinguished from the other species in the aggregate currently known from Sweden by the more elongate and strongly pilose flagellomeres, and from O. costatus and O. parvulus also by the considerably narrower temple in lateral view. Etymology The species has very slender antennae. Material examined > 100 ♀♀, 27 ♂♂ (Sweden); 3 ♀♀ (Germany); 1 ♀, 6 ♂♂ (Norway); 3 ♀♀ (Lithuania). Type material Holotype SWEDEN • ♀; Öland, Mörbylånga, Strandskogen; 56.702° N, 16.494° E; 7 Sep. 2016; B. Andersson leg.; MV-light in garden on sandy soil close to deciduous forest; STI-NJBC252; NHRS-HEVA000008727. Paratypes SWEDEN • 2 ♀♀; Uppland, Rådmansö, Strömsborg; 59.711° N, 18.962° E; 14 Aug.–7 Sep. 2016; N. Ryrholm and C. Källander leg.; MV-light trap in coastal mixed forest; STI-NJBC243, STI-NJBC248; NHRS-HEVA000008728, NHRS-HEVA000008729 • 1 ♀; Uppland, Norrtälje, Väddö Skjutfält; 59.939° N, 18.914° E; 4 Sep.–16 Oct. 2016; N. Ryrholm and C. Källander leg.; Military shooting range, MV-light trap; STI-NJBC244; NHRS-HEVA000008730 • 1 ♀; Skåne, Ystad, Spraggehusen; 55.442° N, 14.246° E; 29 Jul.–28 Aug. 2016; N. Ryrholm and C. Källander leg.; MV-light trap in sanddunes; STI- NJBC245; NHRS-HEVA000008731 • 1 ♀; Gotland, Öja, Gisle; 57.049° N, 18.289° E; 29 Sep.–23 Oct. 2007; N. Ryrholm and C. Källander leg.; MV-light trap; STI-NJBC246; NHRS-HEVA000008732 • 1 ♀; Gotland, Sundre, Suders; 56.945° N, 18.303° E; 29 Sep.–26 Oct. 2007; N. Ryrholm and C. Källander leg.; MV-light trap in deciduous woodland; STI-NJBC247; NHRS-HEVA000008733 • 1 ♀; Gotland, Sundre, Barrshage; 56.922° N, 18.186° E; 10 Sep.–9 Oct. 2006; N. Ryrholm and C. Källander leg.; MVlight trap in wet meadows; NHRS-HEVA000008734 • 1 ♀; Öland, Borgholm, Hälludden Byxelkrok; 57.356° N, 17.058° E; 22–24 2017; R. Isaksson leg.; MV-light; NHRS-HEVA000008735 • 1 ♀; Norrbotten, Övertorneå, Soukolojoki; 66.472° N, 23.632° E; 2 Aug.–5 Oct. 2017; N. Ryrholm and C. Källander leg.; MV-light trap; NHRS-HEVA000008736 • 1 ♀, 2 ♂♂; Uppland, Norrtälje, Väddö Skjutfält; 59.939° N, 18.914° E; 29 Jul.–18 Sep. 2017; N. Ryrholm and C. Källander leg.; MV-light trap in coastal military shooting range; NHRS-HEVA000008737 to NHRS-HEVA000008739 • 2 ♀♀; Skåne, Ystad, Spraggehusen; 55.442° N, 14.246° E; 28 Aug.–30 Sep. 2006; N. Ryrholm and C. Källander leg.; MV-light trap in sanddunes; NHRS-HEVA000008740, NHRS-HEVA000008741 • 1 ♂; same data as for holotype; 1 Aug.–28 Jul. 2016; B. Andersson leg.; MV-light in garden on sandy ground close to deciduous, oak dominated forest, NHRS-HEVA000008742 • 1 ♀; Gotland, Öja, Petesviken; 57.004° N, 18.295° E; 5 May–13 Oct. 2013; N. Ryrholm and C. Källander leg.; MV-light trap; NHRS- HEVA000008743 • 1 ♂; Öland, Mörbylånga, Nedra Västerstad; 56.416° N, 16.411° E; 10 Jul. 2018; N. Johansson leg.; MV-light in garden surrounded by deciduous forest; NHRS-HEVA000008744. Description Fore wing length 15–16 mm. Antenna with 46–53 flagellomeres. First flagellomere 4.5 times as long as wide. Second flagellomere very elongate about 3.5–4.0 times as long as wide. Central flagellomeres elongate, about 2.5–3.0 times as long as wide. Apical flagellomeres approximately 2.5 times as long as wide (Fig. 8C). Apical flagellomeres in female, less so in males, with long prominent pilosity. Length of pilosity in females at least 0.5 width of flagellomere (Fig. 8C). Head narrow behind eyes, in lateral view with temple 0.2–0.3 times as long as compound eye in females, slightly longer in males, 0.4–0.5 times the lenght of the compound eye. Ocelli in female large, partly covering the inner margin of compound eye in dorsal view, the distance between ocelli about 0.3 times the diameter of ocelli. Malar space about 0.1 times as long as mandibular base in female and about 0.3 times in male. Mandibular gape right-angled, with internal angles. Wing membrane clear. Ramellus usually distinct, reaching 0.2–0.3 the width of discosubmarginal cell but sometimes small or absent. Radius evenly curved (Fig. 13A). Mesopleuron weakly shagreened with deep, distinct punctures. Interstices between punctures about equal to their diameter. Epicnemial carina, in antero-ventral view, with pleurosternal angles obviously anterior to sternal angles. Pleurosternal angles rounded, slightly obtuse. Scutellum with lateral carinae only indicated basally. Propodeum with very weak rugose structure, shining with anterior and posterior transverse carina strongly raised (Fig. 10L). Longitudinal carinae delimiting area superomedia and lateral longitudinal carinae sometimes weak or absent, but normally clearly indicated at the junction with the posterior transverse carina. Hind trochantellus shorter than wide in dorsal view. Legs normal with hind femur about 6.0–7.0 times as long as wide. Sclerotised part of first sternite ending level or slightly posterior to spiracle. Inner spur of hind tibia as long as 0.4 times metatarsus. Colour Body testaceous. If infuscate or pale markings occur, they are usually far less distinct than in other species of the aggregate. Mandibular teeth black. Ovipositor brownish, slightly contrasting with the more testaceous posterior metasomal segments. DNA barcode The DNA barcode sequences of nine Swedish specimens of Ophion tenuicornis Johansson sp. nov. are available at the BOLD systems database (www.boldsystems.org, BIN. BOLD: ACK3000. Specimen codes: STI-NJBC: 243–248, 250–252). Ecology The flight period is late summer and autumn. One female in NHRS was reared from Thyatira batis (Linnaeus, 1758) and one female was reared from the same host in HSC. This is notable since only the two species of the subgenus of Platophion are known to parasitize the subfamily Thyatirinae. Both reared specimens are rather small, which suggests the existence of additional hosts. Distribution in Sweden Occurs commonly all over the country, but rarer in the north.Published as part of Johansson, Niklas & Cederberg, Björn, 2019, Review of the Swedish species of Ophion (Hymenoptera: Ichneumonidae: Ophioninae), with the description of 18 new species and an illustrated key to Swedish species, pp. 1-136 in European Journal of Taxonomy 550 on pages 104-106, DOI: 10.5852/ejt.2019.550, http://zenodo.org/record/347640
Ophion vardali Johansson & Cederberg 2019, sp. nov.
Ophion vardali Johansson sp. nov. urn:lsid:zoobank.org:act: 2395488B-9203-4CB7-BE81-62510D5989BA Figs 19 A–B, 45A–B Diagnosis Similar to Ophion pteridis, but besides the obvious differences in colour, with the head more rounded in anterior view and the face wider in relation to the compound eyes. The occipital carina is centrally raised forming an acute angle, the head is more buccate with a distinct gap between the lateral ocellus and the compound eye and the central flagellomeres are stouter. Also similar to Ophion inclinans Johansson sp. nov. and O. arenarius Johansson sp. nov., but the head has a distinct gap between the lateral ocellus and compound eye, the head in anterior view is more rounded and the face wider in relation to the compound eyes. According to the molecular analysis the species is closely related to Ophion pteridis, but based on the distinct morphological characters, both species are regarded as valid. Etymology Named in honour of Hege Vårdal, curator at the Swedish Museum of Natural History, who has been of great help during this study. Material examined 2 ♀♀, 3 ♂♂ (Sweden); 7 ♀ (Norway). Type material Holotype NORWAY • ♀; Nordland, Bodø, Ausvika; 67.336° N, 14.496° E; 10 Aug. 2018; G. Ørsnes leg.; coastal meadow adjacent to forest; BOLD: DS-ICHNN; NHRS-HEVA000008745. Paratypes SWEDEN • 2 ♀♀, 2 ♂♂; Skåne, Ystad, Järahusen; 55.396° N, 14.207° E; 8 Jul.–28 Aug. 2016; N. Ryrholm and C. Källander leg.; MV-light trap in coastal sanddunes in mixed oak and pine forest; NHRS-HEVA000008746 to NHRS-HEVA000008749 • 1 ♂; Skåne, Kristianstad, Åhus; 55.944° N, 14.262° E; 22–26 Jul. 1964; unknown leg.; MV-light; MZLU Type no. 6375:1. Description Fore wing length 16–17 mm. Antenna in both sexes with 54–55 flagellomeres. First flagellomere 3.5–4.0 times as long as wide. Central flagellomeres about 1.5–1.7 times as long as wide. Head quite narrow behind eyes, with or without small gap between lateral ocellus and compound eye. Temple in lateral view about 0.7–0.8 times as long as compound eye as a result of the very short compound eye in lateral view (Fig. 19B). Head in frontal view rounded (width/height measured from the apical margin of clypeus to the top of head = 1.15) with face wide in relation to compound eyes (Fig. 19A). Head with occipital carina centrally raised forming an acute angle (as in Fig. 7B). Face below antennal sockets with quite scarce punctures, polished or weakly shagreened. Malar space about 0.4 times as long as mandibular base in female and about 0.4–0.5 times in male. Mandibular gape right-angled, with internal angles. Wing membrane yellowish. Ramellus distinct, reaching 0.2–0.3 times the width of the discosubmarginal cell. Radius sinuous. Mesopleuron shagreened and distinctly punctate, spaces between punctures about equal to their diameter. Pleurosternal angles weakly obtuse to right angled, obviously anterior to sternal angles. Scutellum with distinct lateral carinae (as in Fig. 6C). Propodeum weakly shagreened, often quite polished with anterior transverse carina strong, but sometimes partly absent laterally. Posterior transverse carina often widely interrupted centrally. Area superomedia often present. Central longitudinal carinae often distinct. Sclerotised part of first sternite ending obviously posterior to spiracle. First tergite in lateral view with slight or distinct median undulation (as in Fig. 6E). Hind trochantellus shorter than wide in dorsal view. Inner spur of hind tibia as long as 0.5 times hind metatarsus. Colour Body testaceous, the lower part of mesopleuron, mesosternum, propleuron, coxae mostly and propodeum strongly infuscate, black or dark brown (Fig. 45 A–B). Head with inner and outer orbits yellow. Mandibular teeth black. Ovipositor sheath testaceous, concolourous with posterior metasomal segments. DNA barcode The DNA barcode sequence of four specimens of Ophion vardali Johansson sp. nov. is available at the BOLD systems database (www.boldsystems.org, BOLD: DS-ICHNN; STI-NJBC 24–26). Ecology No host records are known for this species and it is active mainly during July. The habitat seems to be open or semi-open coastal meadows. Distribution in Sweden Ophion vardali Johansson sp. nov. is very rare and known from only a couple of localities in the southeastern part of the country. Remarks On the website www.artsobservasjoner.no there are also pictures of specimens from several different coastal localities in Central and Northern Norway. One Norwegian female, kindly provided by Geir Ørsnes, is designated as holotype. The rather vague description of Ophion praecinctus Meyer, 1935 describes a species from the desert country Turkmenistan with similar colouration to O. vardali Johansson sp. nov. The type material is lost (Townes et al. 1965; Andrey Khalaim, ZIN, pers. com.) and there seems to be no remaining additional material determined by the Meyer. The description (Meyer 1935) and a simplified translated key (Meyer 1937) refers to a very small species (body length 12 mm), with transverse head, long malar space, vague notauli, deep concavities along the inner eye margin and two distinct transverse ridges on the propodeum, all of which are characters that cannot be attributed to O. vardali Johansson sp. nov.Published as part of Johansson, Niklas & Cederberg, Björn, 2019, Review of the Swedish species of Ophion (Hymenoptera: Ichneumonidae: Ophioninae), with the description of 18 new species and an illustrated key to Swedish species, pp. 1-136 in European Journal of Taxonomy 550 on pages 106-108, DOI: 10.5852/ejt.2019.550, http://zenodo.org/record/347640
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