30 research outputs found
FIGURE 5 in A new biogeographically disjunct giant gecko (Gehyra: Gekkonidae: Reptilia) from the East Melanesian Islands
No full textFIGURE 5. Gehyra rohan sp. nov. in life: A) holotype, RBINS 2684, near Lorengau Village, Manus Island, Papua New Guinea; and B) SAMA R69881 (SJR15105) near Nae, Mussau Island, Papua New Guinea. Photographs by M. M. T. Jocque and S. Venter.Published as part of Oliver, Paul M., Clegg, Jonathan R., Fisher, Robert N., Richards, Stephen J., Taylor, Peter N. & Jocque, Merlijn M. T., 2016, A new biogeographically disjunct giant gecko (Gehyra: Gekkonidae: Reptilia) from the East Melanesian Islands, pp. 61-76 in Zootaxa 4208 (1) on page 69, DOI: 10.11646/zootaxa.4208.1.3, http://zenodo.org/record/20201
Ocyale ghost Jocque M. & Jocque R. 2017, sp. nov.
No full textOcyale ghost Jocque M. & Jocqué R. sp. nov. urn:lsid:zoobank.org:act: 37AAF237-A0C7-40D1-9BA8-6FF3D4864D79 Figs 1, 2, 3 A–F, 4A–D, 5A–F, 6A–C, 7A–C Diagnosis Males of O. ghost sp. nov. can be recognized by details of the male palp: the tegulum is restricted to the prolateral side of the bulbus, the distal prong of the palea appendage is much narrower than the proximal one and the MA with the perpendicular prong is rectangular. Females are characterized by the epigyne in which the T-shaped posterior sclerite is fully exposed and not covered with setae as in other species in the genus. Etymology The species name ‘ghost’ refers to the fully white appearance of this spider. Additional reference is made to the large white direwolf ‘Ghost’ in Game of Thrones, the first book in the series of fantasy novels A Song of Ice and Fire by George R.R. Martin. Type material Holotype MADAGASCAR: ♂, Mahajanga Prov., Matsedroy, Lake 2, 15°29′33.9″ S, 046°39′04.1″ E, 5 Jul. 2016, 19:55–20:45, hand collected on sandy beach (Fig. 2), S. Wellens leg. (MRAC 245337). Paratypes MADAGASCAR: 1 ♀, 2 juvs, same data as holotype (MRAC 245338); 2 ♀♀, 2 juvs, same data as preceding (MRAC 245340); 2 ♀♀, 2 juvs, as preceding (MRAC 245341); 2 ♀♀, 2 juvs, as preceding (MRAC 245342); 1 ♀, as preceding (MRAC 245347); 1 ♂, 6 juvs, as preceding (MRAC 245348); 1 ♂, 3 juvs, as preceding (MRAC 245361); 2 ♀♀, 2 juvs, as preceding (MRAC 245350); 1 ♀, 5 juvs, as preceding except 12 Jul. 2016 (MRAC 245339); 1 ♀, 5 juvs, as preceding except 22 Jul. 2016 (MRAC 245351); 1 ♀, 4 juvs, as preceding except 24 Jun. 2016 (ZBA); 1 ♂, 4 ♀♀, as preceding (ZBA); 1 ♂, 2 ♀♀, 1 juv., as preceding (ZBA). Description Male (holotype) MEASUREMENTS. Total length 15.35; carapace 8.77 long, 2.27 high, 6.58 wide, narrowed to 3.95 in eye region; labium 1.14 wide, 0.79 long; sternum 3.68 long, 3.07 wide. COLOUR. Alive (Fig. 3 A–E): almost entirely white with slightly contrasting darker spots as in ethanol specimens; eyes surrounded by yellow rings; cheliceral condyle reddish brown. In ethanol (Fig. 4 A–B): fairly different from colouration of live specimens; carapace brownish cream with interrupted black radiating striae, black fovea, two pairs of spots in front of fovea and dispersed irregular spots along margins; PEQ covered with white setae, tegument dark in its anterior half; chelicerae brown with dark setae as seen in frontal view, dark brown with black setae in ventral view; labium dark brown with cream crescent shape along anterior concave margin; sternum with strongly sinuous lateral margins, ending in long, tapered point; with dispersed short dark setae; legs formula IV-III-I-II; uniform cream with dispersed short, dark setae and dark spines; pedipalp: femur, patella and tibia cream, cymbium and bulbus contrasting dark brown; abdomen: dorsum cream with dispersed small dark spots, two reddish apodemes in anterior half and dispersed dark setae, sides and venter uniform cream; spinnerets: ALS dark, PLS and PMS pale on dorsal side, dark on ventral side. EYES. AME: 0.35; ALE: 0.15; PME: 0.67: PLE: 0.61; eye rows: ALE: 1.51, PME: 1.51, PLE: 2.18. CHELICERAE. With three teeth on retromargin, one small proximal tooth and one larger distal tooth on promargin. LEGS. Spination of leg I (identical on both sides): femur pl4, d3, rl2; patella pl1, rl1; tibia pl2, d2, rl2, v2-2-2; metatarsus pl2, rl2, v2-2, dw5. Leg measurements: see Table 1. PALP (Figs 5 A–D, 6A–C, 7A–B). Tegulum ribbed, developed on prolateral basal part of bulbus; palea with two prongs, proximal one broad and slightly curved, distal one thin and strongly curved; embolus originating on retrolateral part of palea and curved ventrally around it; MA large, with prolateral part a short hook, ventral part subrectangular, perpendicular to the former. Female (paratype MRAC 245338) MEASUREMENTS. TL 16.24; carapace 7.95 long, 3.69 high, 6.31 wide, narrowed to 3.69 in eye region; labium 0.92 long, 1.08 wide; sternum 3.33 long, 2.83 wide. COLOUR. In ethanol (Fig. 4 C–D): dorsal surface of abdomen more uniform cream than in male but with similar dark spots and apodemes; pedipalp as in the male except for unmodified tarsus with dark tip. EYES. AME: 0.42; ALE: 0.21; PME: 0.67: PLE: 0.60; eye rows: ALE: 1.44, PME: 1.44, PLE: 2.14. LEGS. Spination of leg I, right (left): femur pl2(5), d3(4), rl3(3); patella pl1(1), rl1(1); tibia pl2(2), d1(1), rl2(2), v2-2-2 (3-2-2); metatarsus pl2(2), rl2(2), v2-2-2(2-2-2), dw5(5). Leg measurements: see Table 1. EPIGYNE (Figs 5 E–F, 7C). Roughly triangular area surrounded by dense mat of white setae; large and broad inverted T-shaped sclerite, 1.6 times wider than long; spermathecae large, globular; entrance ducts Z-shaped with basal portion slightly sinuous. Variation Males: TL 16.76–19.45 (n = 5); females: TL 16.47–22.01 (n = 19). Distribution Known only from the type locality (Fig. 1). Affinities We placed this species in the genus Ocyale based on the presence of the two elongate curved prongs on the palea (Figs 4B, 5 B–C, 6B), the epigyne surrounded with white hairs and with a wide inverted T-shaped sclerite (Figs 5E, 7C), the large globular spermathecae (Fig. 5F) and the conformation of the copulatory ducts (Alderweireldt 1996: fig. 27). The specimen illustrated by Siyam et al. (2015: figs 13–15) from Sudan is probably not an Ocyale, at least not O. pilosa because the palp does not fit the illustrations of Alderweireldt (1996: figs 12–13). The colour pattern described for the genus in both these papers may be inaccurate based on the observed differences between specimens in ethanol and photos of the living spiders for this new species. This indicates the value of including images of living animals in descriptions of new species. It is not clear to which species the representative of Madagascar is most closely related to. Biology Ocyale ghost sp. nov. was only found on the white sandy beaches (Fig. 3 A–B) of an inland lake in the study region. The surveys also included grassland and dry forest, but the species seems restricted to a white-sand habitat, as reflected in its habitus. Ocyale ghost sp. nov. is active at night and all specimens were caught with headlamps after sunset. Captured animals that were kept alive in large ziplock bags overnight constructed retreats in the sand, lined with silk (Fig. 3C). Possible prey include large insects such as grasshoppers (Fig. 3F) that also exhibit camouflage colours as an adaptation to the white beach they live on. Intraspecific predation is also likely to occur (Fig. 3E), a phenomenon which is not unusual among lycosids (Edgar 1969; Hallander 1970). We observed copulation and females with spiderlings (Fig. 3D) in the midst of the dry season (June–July). Juveniles of a complete range of size, from very small ones (6 mm TL) to subadults, were observed, indicating that this species might reproduce yearround. The permanent presence of water in its habitat might explain why this species is also active in the dry season when spider activity is on average very low.Published as part of Jocque, Merlijn, Wellens, Siel, Andrianarivosoa, J. D., Rakotondraparany, F., Seing, Sam The & Jocqué, Rudy, 2017, A new species of Ocyale (Araneae, Lycosidae) from Madagascar, with first observations on the biology of a representative in the genus, pp. 1-13 in European Journal of Taxonomy 355 on pages 5-11, DOI: 10.5852/ejt.2017.355, http://zenodo.org/record/383631
Cambonilla symphonia Jocqué & Jocque & Stock & Rin & Henrard 2019, gen. et sp. nov.
No full textCambonilla symphonia Jocqué & Henrard gen. et sp. nov. urn:lsid:zoobank.org:act: 51939E05-B3E2-41DD-BB97-E12D1D395FE7 Figs 3A, 4 G–I, 8, 9, 10, 11 Diagnosis The male of Cambonilla symphonia gen. et sp. nov. is recognized by the presence of stridulating organs on Fe I and II, and by the strong tapered RTA on the palp, the female by the epigyne with very deeply incised central plate. Etymology The species name ‘ symphonia ’ is a noun in apposition meaning ‘orchestra’ and refers to the four stridulating organs on the femora of the male. Material examined Holotype CAMBODIA • ♂; Kratie province, Base Camp 4; 12.5460° N, 105.9881° E; 5 May 2018; M. Jocque and W. Stock leg.; pitfall set 1; RBINS IG 33.889 /007 (=CA15). Paratypes CAMBODIA • 1 ♀; Kratie province, Base Camp 3; 13.0199° N, 106.0395° E; 10 May 2018; M. Jocque and W. Stock leg.; rainforest, pitfall set 3; RBINS IG 33.889 /008 • 2 ♂♂; Kratie province, Base Camp 4; 13.0198° N, 105.9268° E; 5 May 2018; M. Jocque and W. Stock leg.; rainforest, pitfall set 1; RBINS IG 33.889 /009. Description Male holotype TOTAL LENGTH. 4.83. Carapace: length 2.13, width 1.63, height 0.85. COLOUR (Fig. 8 A–D). Carapace uniform chestnut brown; chelicerae, chilum and sternum brownish orange; endites and labium pale brown with whitish distal margin; legs: Cx and Tr pale brown; Fe dark brown; remainder yellow except T IV medium greyish brown; abdomen: dorsum with two reniform pale patches in front overlaid by translucent pale reddish brown, oval scutum followed by numerous small white spots on dark background, venter sclerotized orange in front of epigastric furrow with short tubular extension around pedicel; remainder with sepia marbling on pale background; spinnerets whitish; sides with dark grey marbling. CARAPACE. Rugose, with sparse, thin, pale, branched setae; one large seta between AME. STERNUM (Fig. 8B). Shield shaped; 0.92 wide, 0.85 long. Posterior tip shallowly indented; with rounded depressions in front of coxae; no precoxal sclerites. CHILUM. A single sclerite 0.12 high, 0.25 wide, with central protrusion. EYES. ALE 0.12; AME 0.13; PLE 0.10; PME 0.13; ALE-AME 0.10; AME-AME 0.07; PLE-PME 0.13; PME-PME 0.08. Clypeus 0.41 high. LEGS. Trochanters III and IV with anterior ventral indentation; Fe I and II with basal dorsal stridulation file (Fig. 4 G–H); without hinged setae. ABDOMEN. Venter with dispersed thick setae in posterior half (Fig. 8D); with short tubular extension around pedicel, ventral margin slightly indented (Fig. 8 A–B). MALE PALP (Figs 8 E–F, 10A–B). T with a strong, straight, tapered RTA and retrobasal triangular swelling; cymbial fold slightly shorter than half the cymbium’s length; embolus with large base, originating at an angle of 270° to longitudinal axis; bifid from base onwards; MA with basal tooth and beak shaped distal extremity pointing prolaterad; C short, tapered, grooved. Female (RBINS IG 33.889/008) TOTAL LENGTH. 6.60. Carapace: length 2.56, width 1.56, height 1.14. COLOUR (Fig. 9 A–C). Carapace uniform dark brown; chelicerae medium brown without pale tip; chilum medium brown. Endites and labium medium brown with pale frontal margin. Sternum uniform medium brown. Legs: Cx yellowish brown, with white sital margin; Tr yellowish brown, F dark brown, remainder medium brown; palp medium brown, F slightly darker. Abdomen: dorsum anteriorly with two well separated reniform spots, without scutum overlay, followed by pale transverse band, small, pale patches and large patch in front of spinnerets; venter with sclerotized band in front of epigastric furrow, remainder with sepia marbling on pale background, spinnerets yellow. CARAPACE. Rugose, with sparse, thin, pale, branched setae; a few large setae between eye region and fovea. EYES. ALE 0.12; AME 0.15; PLE 0.13; PME 0.15; ALE-AME 0.10; AME-AME 0.05; PLE-PME 0.23; PME-PME 0.08. Clypeus 0.51 high. CHILUM. A single sclerite 0.12 high, 0.33 wide, with central protrusion. STERNUM (Fig. 9B) Shield shaped; 1.14 wide, 1.21 long. Posterior tip truncate; with rounded depressions in front of coxae; no precoxal sclerites. LEGS. Tr III and IV posteriorly concave. Hinged hair on T I and II. PALPAL TARSUS. With toothed claw turned inward over 30°, eight dispersed spines. ABDOMEN. Without frontal tubular extension. Venter of abdomen without PVS or thick setae (Fig. 9C). EPIGYNE (Figs 9D, 10C). A broad sclerotized area with narrow, horseshoe shaped plate, delimiting central pit. Distribution Known from two localities along the Mekong River in Cambodia (Fig. 11).Published as part of Jocqué, Rudy, Jocque, Merlijn, Stock, Willem, Rin, Naroeun & Henrard, Arnaud, 2019, The new Southeast Asian genus Cambonilla gen. nov. (Zodariidae, Araneae): ' bis repetita placent', pp. 1-24 in European Journal of Taxonomy 501 on pages 17-22, DOI: 10.5852/ejt.2019.501, http://zenodo.org/record/257829
Cambonilla securicula Jocqué & Jocque & Stock & Rin & Henrard 2019, gen. et sp. nov.
No full text<i>Cambonilla securicula</i> Jocqué gen. et sp. nov. urn:lsid:zoobank.org:act: 00D14C53-6330-42BA-A2F1-63C760959CF3 <p>Figs 2, 3 B–F, 4A, 5, 6, 7, 11</p> Diagnosis <p> The male of <i>Cambonilla securicula</i> sp. nov. is recognized by the presence of small lateral scuta behind the epigastric fold and by the male palp in which both the MA and the C are hatchet shaped. The female has a remarkable, small, modified spine on the tibia of the palp and a characteristic epigyne in which the central plate has a concave anterior and a straight posterior margin.</p> Etymology <p> The species name ‘ <i>securicula</i> ’ meaning hatchet, is a noun in apposition referring to the shape of both the MA and the C.</p> Material examined <p> <b>Holotype</b> CAMBODIA • ♂; Kratie Province, Base Camp; 13.00986° N, 106.0640° E; 16 May 2018; M. Jocque & W. Stock leg.; Herpetology pitfall in forest; RBINS IG 33.889 /001.</p> <p> <b>Paratypes</b></p> <p>CAMBODIA • 3 ♂♂; same data as for holotype; RBINS IG 33.889 /002(=CA3) • 2 ♂♂, 1 ♀; 17 May 2018; same data as preceding; RBINS IG 33.889 /003 • 4 ♂♂ 2 ♀♀; 13 May 2015; same data as preceding; RBINS IG 33.889 /004 • 4 ♂♂; Kratie Province, Base Camp 4; 13.0198° N, 105.9268° E; 5 May 2018; M. Jocque and W. Stock leg.; pitfall set 1; RBINS IG 33.889 /005 • 1 ♂; Kratie Province; 13.00986° N, 106.06395° E; 10 May 2018; M. Jocque and W. Stock leg; pitfall set 3; RBINS IG 33.889 /006.</p> <p> <b>Other material</b></p> <p>CAMBODIA • 1 ♂, 1 ♀; Kratie Province, Base Camp; 13.00986° N, 106.0650° E; 6–11 May 2018; M. Jocque and W. Stock leg.; forest, by hand; RBINS IG 33.889 /010 • 2 ♂♂; Kratie Province, Base Camp 1; 12.61788° N, 106.0 0 296° E; 10 Apr. 2018; M. Jocque and W. Stock leg.; flooded forest on sand, Malaise trap, by hand; RBINS IG 33.889 /011.</p> <p> LAOS – <b>Vientiane Province</b> • 1 ♀; Vientiane, Ban Don Ma Khay; 18.08384° N, 102.1713° E; 13 Jun. 2013; P. Jäger leg.; 186 m, secondary forest, at night by hand; SMF – <b>Champasak Province</b> • 1 ♀; Muang Bachieng, That Paxuam; 15.17653° N, 105.9225° E; 3.Aug. 2016; P. Jäger and S. Munnich leg.; 197 m,secondary forest with single primary trees, by night, by hand; SMF – <b>Khammouan Province</b> • 1 ♀; Ban Tathot; 17.62361° N, 105.1466° E; 18–19 Fev. 2003; P. Jäger leg.; ca 180 asl, village and neighbourhood, shrubs and rocks, night and evening; SMF.</p> Description <p> <b>Male holotype</b></p> <p>TOTAL LENGTH. 6.96. Carapace: length 3.48, width 2.70, height 2.00.</p> <p>COLOUR (Fig. 5 A–D). Carapace uniform dark chestnut brown; chelicerae dark brown with orange extremity and white tip; chilum medium brown; sternum medium brown; labium and endites medium brown, paler towards anterior white tip; legs: Cx and Tr medium brown, Fe dark brown, P and T greyish brown, Mt and t pale brown.Abdomen: dorsum with pale reniform patches fused in front and at posterior end, overlaid by reddish orange, translucent oval scutum; further backward with five transverse bands and pale patch above spinnerets on dark background; venter sclerotized in front of epigastric furrow and yellow sclerotized area on either side behind epigastric furrow; remainder marbled with sepia on pale background; spinnerets pale brown.</p> <p>CARAPACE GRANULATED. With sparse cover of thin, pale, branched setae.</p> <p>EYES. ALE 0.16; AME 0.20; PLE 0.13; PME 0.13; ALE-AME 0.10; AME-AME 0.07; PLE-PME 0.12; PME-PME 0.12. Clypeus 0.66 high.</p> <p>CHILUM. 0.21 high, 0.43 wide, with slightly concave inferior side, centre protruding.</p> <p>STERNUM (Fig. 5B). Shield shaped, granular; 1.42 wide, 1.49 long. Posterior tip shallowly indented; with rounded depressions in front of coxae; no precoxal sclerites.</p> <p>LEGS. Trochanters III and IV with anterior ventral indentation; dorsal hinged hair on base of T I and II. Cx and ventral side of F granular.</p> <p>ABDOMEN. With PVS in ill-defined group and dispersed thick setae in posterior half (Fig. 5D); with short tubular extension around pedicel, its anterior margin ventrally indented (Fig. 5B).</p> <p>MALE PALP (Figs 4 C–F, 5E–F, 7A–B). T with strong, slightly sinuous, downcurved, blunt RTA; cymbial fold as long as half the cymbium’s length; embolus with large base, originating at an angle of 270° to longitudinal axis, halfway with short, flat bifurcation, distal end corkscrew shaped; MA hatchet-shaped with short ventral tooth at base; C with short, sharp, flat, distal protrusion, pointing prolaterad.</p> <p> <b>Female</b> (RBINS IG 33.889/003)</p> <p>TOTAL LENGTH. 7.31. Carapace: length 3.05, width 2.06, height 1.42.</p> <p>COLOUR (Fig. 6 A–C). Carapace uniform dark brown; chelicerae medium brown with pale tip; chilum medium brown. Endites and labium medium brown with pale frontal margin. Sternum uniform medium brown. Legs I: Cx medium brown, Tr pale brown, F dark brown, remainder yellowish orange; palp medium brown except T yellowish orange. Abdomen: dorsum with two ill-defined reniform spots touching in front, without scutum overlay, followed by small, faint, pale patches and large patch in front of spinnerets; venter with sclerotized band in front of epigastric furrow, without tubular extension, remainder marbled with sepia on pale background, spinnerets pale yellow.</p> <p>CARAPACE. Rugose with sparse cover of branched setae.</p> <p>EYES. ALE 0.18; AME 0.18; PLE 0.15; PME 0.13; ALE-AME 0.15; AME-AME 0.07; PLE-PME 0.23; PME-PME 0.08. Clypeus 0.51 high.</p> <p>CHILUM. A single sclerite 0.12 high, 0.33 wide, with central protrusion. STERNUM (Fig. 6B). Shield shaped; 1.28 wide, 1.35 long. Posterior tip rounded; with rounded depressions in front of coxae; no precoxal sclerites.</p> <p>LEGS. Tr III and IV posteriorly indented. Hinged hair on T I and II.</p> <p>FEMALE PALP. Tarsus with toothed claw turned inward over 30°, eight dispersed spines; tibia with short, sinuous prolateral spine (Fig. 6F).</p> <p>ABDOMEN. Venter without PVS or thick setae (Fig. 6C).</p> <p>EPIGYNE (Figs 6 D-E, 9D). With wide rectangular plate, concave in front, posterior margin convex and thickened in the middle. Copulatory ducts with few whorls, ending in poorly delimited spermathecae near posterior end.</p> Variation <p>The abdominal venter of the female specimen from Ban Tathot (17.62361 °N, 105.1466 °E) is not marbled but has three dark, faint longitudinal lines.</p> Distribution <p>Known from lowland forest in Cambodia and Laos; all localities are along the Mekong river (Fig. 11).</p>Published as part of <i>Jocqué, Rudy, Jocque, Merlijn, Stock, Willem, Rin, Naroeun & Henrard, Arnaud, 2019, The new Southeast Asian genus Cambonilla gen. nov. (Zodariidae, Araneae): ' bis repetita placent', pp. 1-24 in European Journal of Taxonomy 501</i> on pages 12-17, DOI: 10.5852/ejt.2019.501, <a href="http://zenodo.org/record/2578299">http://zenodo.org/record/2578299</a>
Amphibolocypris arida Jocque, Brendonck, Riddoch & Martens, 2010, sp.nov.
No full textAmphibolocypris arida sp.nov. Jocque & Martens (Figs 2–5) Type locality: Rock pools near Thamaga, south-eastern Botswana (24 ° 40 ’ 30 ” S, 25 ° 31 ’00” E). The pools are situated on granite rocky outcrops, surfacing in a mainly savannah matrix of grasslands and shrubs. The pools are generally shallow (max 20 cm deep) and small (max. 2 m radius), with water temperature reaching up to 37.1 °C at mid day. Based on measurements in 8 pools, habitat waters were characterized by a basic pH varying between 7.3 and 8.8 and were always low in conductivity with values ranging from 27.6 to 69.1 µSm/cm (Jocque, unpubl.). Accompanying ostracod fauna: Heterocypris sp.nov., Heterocypris incongruens, Sarscypridopsis cf gregaria, Potamocypris spec. Type material: Holotype: a male, with soft parts dissected in glycerine in a sealed slide and with valves stored dry (after use for SEM) in a micropalaeontological slide (OC. 3167) Allotype: a female, with soft parts dissected in glycerine in a sealed slide and with valves stored dry (after use for SEM) in a micropalaeontological slide (OC. 3168) Paratypes: 2 males and 2 females, with soft parts dissected in glycerine in a sealed slide and with valves lost (OC. 3166, OC. 3169 -OC. 3171) Derivation of name: as the species is thus far only known from rock pools in a (semi-) arid area, the specific name refers to this characteristic of its habitat. Diagnosis: Valves elongated, with anterior margin more broadly rounded, posterior margin bluntly pointed, posterior calcified inner margin narrow, but present. External surface set with elongated microridges. Valves sexually dimorphic, with female valves being more elongated. Natatory setae on A 2 of medium length. T 2 with subapical seta claw-like, reaching middle of elongated end claw. CR without proximal seta. Hemipenis with one shield with one club-like expansion, the other shield with two elongated lobes, one pointed, basally inserted, and one distally rounded, more apically inserted. Prehensile palps almost symmetrical, relatively short, with broad base and sickle-shaped. Additional description of male: Valves in lateral view (Figs 2 A,B) elongated, weakly calcified, externally set with longitudinal micro-ridges and sparse rimmed pores (Figs 2 E,F = from female valves). with rounded dorsal margin, anterior margin more broadly rounded than posterior margin, ventral margin slightly sinuous. Valve outline antero-dorsally with a dent. Anterior calcified inner lamella broad, dorsally ending at dent; posterior calcified inner lamella narrow, but present. Muscle scars situated in front of the middle. Carapace in dorsal view very narrow, lancet shape. No overlap between LV and RV. A 1 (Fig. 3 A) with 7 segments. First segment with 2 ventral setae, no dorsal seta. Second segment with one medio-dorsal seta, Rome and Wouters organs not visible. Third segment more than twice as long and wide, with one shorter ventral and one longer dorsal seta. Fourth segment almost as long as wide, with 2 shorter ventral setae and 2 long dorsal natatory setae. Fifth segment slightly longer than wide, also with two shorter ventral and 2 longer dorsal natatory setae. Sixth segment almost twice as long as wide, with 1 short ventral setae and 4 long natatory setae. Terminal (seventh) segment more than twice as long as wide, with 1 shorter seta, one long aesthetasc Ya (not drawn) and 2 long natatory setae. A 2 (Fig. 3 B) with exopodite reduced to a small plate, bearing 1 long and 2 short setae. Endopodite 3 - segmented. First segment elongated and stout, aesthetasc Y short and club-like, distal part swollen. Natatory setae of unequal length, but all relatively short, mostly not reaching tip of second segment; ventral-most seta the longest. Second segment with 2 dorso-lateral and 3 ventro-lateral setae (2 long, 1 short), distal chaetotaxy typical of male Cyprididae, with setae z 2 and z 3 changed into claws, z 1 a seta and claw G 1 turned into a long seta, extending beyond tips of all claws, these claws stout, relatively short and set with strong spines. Terminal segment with claws G M and G m, seta g and aesthetasc y 3, basally fused with an accompanying seta. Md (compare to Figs 3 D,E,E’ = female) with coxal plate distally set with rows of spines and small setae. Palp with alpha-seta short, narrow and smooth, beta-seta short, stout and hirsute, gamma-seta short, broad and hirsute in distal fourth of its length. Second segment dorsally with a group of 3 smooth setae, ventrally with 3 long and smooth and 1 shorter and hirsute setae. Third segment dorsally with 4 subapical setae, ventrally with 1 subapical seta, distally with 3 normal setae and a gamma-seta. Terminal segment with 3 stout claws and 3 small setae. Mx 1 (compare to Fig. 4 A = female) with second palp-segment slightly spatulate, Zahnborsten on third endite smooth. T 1 (Figs 5 A, B) with 1 a-seta, 1 b- and 1 d-setae. Distal chaetotaxy of coxal plate consisting of 16 setae of sometimes very different shape and length. Prehensile palps almost symmetrical; first segments with large ventral protuberance, set with 1 large sensory organ; second segments sickle-shaped, tapering towards the end, distally with one long and stout sensory organ. T 2 (compare to Fig. 4 B= female) with elongated segments and unusually long setae. First segment with seta d 1 reaching into second segment, this (knee-) segment with seta d 2 missing. Third segment with 1 long ventro-apical seta, reaching almost to fifth segment. Fourth segment divided into two elongated subsegments: segment 4 a with a relatively short ventro-apical seta, not reaching tip of segment 4 b, this latter segment with 2 subequal, ventro-apical setae. Fifth (terminal) segment with 1 subapical seta and 1 subapical claw (generic character) and 1 long and thin apical claw. T 3 (compare to Figs 4 C,C’= female) a cleaning limb. First segment with 3 setae. Second segment with 1 long apical seta. Third segment with 1 short lateral seta. Distal part of third and fourth segment fused to a pincer shaped organ, bearing 1 long seta, 1 seta of medium length, set with two rows of setulae and 1 very short seta. CR (Fig. 4 E) distally with 2 claws and 1 apical seta, proximal seta missing (generic character). Attachment (Fig. 4 D = female) slender, with simple distal bifurcation. Hemipenis (Fig. 5 C) with a complex of ventral lobes of medial and lateral shields: one shield with one club-like expansion, the other shield with two elongated lobes, one pointed, basally inserted, one distally rounded, more apically inserted. Additional description of female: Female valves more elongated, dorsal margin bluntly pointed, with greatest height less than ½ the length and situated almost in the middle. Anterior margin more rounded than posterior margin, but both less so than in the male. Calcified inner lamellae and position of muscle scars as in the male. A 1, Md (Figs 3 D,E,E’), Mx 1 (Fig. 4 A) and T 3 (Fig. 4 C,C’) as in the male. A 2 (Fig. 3 C) with natatory setae slightly longer than in the male, z 1-3 all setae, reaching up to or beyond tips of end claws, the latter more slender and longer than in the male and set with more delicate spines. T 1 (Fig. 4 F) with palp short and wide, distally with 3 relatively short setae. T 2 (Fig. 4 B) with segments and setae slightly longer and more slender than in the male. CR slightly more slender than in the male. Remarks: Most species in the subfamily Isocypridinae are known to have weakly calcified valves. In the present study, most specimens of A. arida sp.nov. have either weakly calcified, or completely decalcified valves. This means that only few specimens had valves reflecting the true shape, hence the limited number of illustrations of these valves. The longitudinal ridges on the valve surfaces (Figs 2 E,F) could function as reinforcement of weakly calcified valves. Measurements (in mm): Male, holotype: RV, L = 1.69, H= 0.65; LV, L= 1.67, H= 0.65 Female, allotype: RV, L= 1.58, H= 0.63; LV, L = 1.58, H= 0.64 Differential diagnosis: Amphibolocypris arida sp.nov. differs from the other described species in the genus, the type species A. exigua, by the more elongated valves, with more pointed posterior margin, especially in the female; by the shape of the hemipenis (one large rounded lobe and one narrower lobe in A. exigua, one subquadrate lobe and three more elongated lobes in A. arida) and of the prehensile palps (proximal segments with more elongated ventral protuberances and more elongated and narrower distal segments with shorter distal sensory organs in A. exigua). Two further, as yet undescribed, species from Namibia (Martens, unpubl.) differ from both A. exigua and A. arida sp.nov. in the shape of the hemipenis (see below). Ecology and distribution: The species is presently known from its type locality only, namely rock pools on a granite outcrop in south-eastern Botswana. From other, as yet unpublished records of other species of Amphibolocypris from Namibia (see above), it would seem that this once monospecific genus might be more speciose than was previously assumed, in which case the individual species could have rather limited distributions. Other species investigated: As a comparison, two further new species of Amphibolocypris from the western part of southern Africa are mentioned here, and the outlines of the hemipenis given. Due to limitation of material (single specimen, completely decalcified valves), these species are left in open nomenclature.Published as part of Jocque, Merlijn, Brendonck, Luc, Riddoch, Bruce J & Martens, Koen, 2010, On Amphibolocypris arida sp. nov. (Crustacea, Ostracoda), from rock pools in Botswana (southern Africa), pp. 47-58 in Zootaxa 2408 on pages 49-56, DOI: 10.5281/zenodo.19426
Ocyale Audouin 1823
No full textOcyale Audouin, 1823 Key to the Afrotropical species of Ocyale Letters refer to characters shown by arrows in key figures. Species included: O. dewinterae Alderweireldt, 1996 ♂ ♀ O. discrepans Roewer, 1960 ♀ O. ghost Jocque M. & Jocqué R. sp. nov. ♂ ♀ O. grandis Alderweireldt, 1996 ♂ ♀ O. guttata Karsch, 1878 ♂ ♀ O. pilosa Roewer, 1960 ♂ ♀ 1. Males................................................................................................................................................. 2 – Females............................................................................................................................................. 6 2. Small species: TL 6–8 mm; distal part of cymbium shorter than bulbus (b) and bulbus clearly broadened (a).............................................................................. O. dewinterae Alderweireldt, 1996 – Larger species, TL> 16 mm; distal part of cymbium longer than bulbus (c) and bulbus clearly not broadened (d).................................................................................................................................... 3 3. Distal prong of palea much narrower than proximal prong (e); ventral process of MA subrectangular with long parallel margins (f)............................................ O. ghost Jocque M. & Jocqué R. sp. nov. – Prongs of palea of similar size (g); long margin of ventral MA process with bulge (h).................. 4 4. Lateral process of MA slender, tapered to sharp tip (i); TL> 23 mm........................................................................................................................................................... O. grandis Alderweireldt, 1996 – Lateral process of MA, shorter and broader, blunt (j); TL 23 mm; epigyne with small and short septum (p).......................................................................................................................................................... O. grandis Alderweireldt, 1996 – Smaller species, TL <23 mm; epigyne with well-developed inverted T-shaped septum (q, r)........ 8 8. Longitudinal part of inverted T longer than transverse part (q); colour pattern uniform......................................................................................................................... O. dewinterae Alderweireldt, 1996 – Longitudinal part of inverted T as long as or shorter than transverse part (r); brown to yellow carapace pattern (in ethanol)............................................................................................................................ 9 9. Lateral epigyne margins along median septum swollen and shiny; longitudinal part of inverted T slender, longer than transverse part.............. O. guttata Karsch, 1878, O. discrepans Roewer, 1960 – Lateral epigyne margins along median septum not swollen and shiny; longitudinal part of inverted T broad, only half as long as transverse part (r)................................................ O. pilosa Roewer, 1960Published as part of Jocque, Merlijn, Wellens, Siel, Andrianarivosoa, J. D., Rakotondraparany, F., Seing, Sam The & Jocqué, Rudy, 2017, A new species of Ocyale (Araneae, Lycosidae) from Madagascar, with first observations on the biology of a representative in the genus, pp. 1-13 in European Journal of Taxonomy 355 on pages 3-5, DOI: 10.5852/ejt.2017.355, http://zenodo.org/record/383631
Pristina terrena Collado & Schmelz 2000
No full textPristina terrena Collado & Schmelz, 2000 Pristina terrena Collado & Schmelz, 2000: 513 –514, figs 8, 9. Material examined. Five specimens, sexually immature, IG 327183 - 23 to IG 327183 - 27; one specimen with budding zone and primordial testes in VIII and primordial ovaries in IX. Cusuco National Park, Honduras, [15.505813 -88.21473, 15.506299 -88.23704, 15.522537 -88.27598, 15.513245 -88.28807, 15.526152 -88.27662], respectively 2061 m, 1457 m, 1558 m, 1468 m and 1587 m asl, M. Jocque 07/07/ 2013 to 06/08/ 2013. Further 6 specimens in the 3 rd author's collection, [15.52425 -88.28853, 15.50731 -88.29428], respectively 1659 m, 1347 m asl, M. Jocque 07/08/ 2006. Description. Body length 1–2 mm, diameter 0.1–0.14 mm. Segment number of complete specimens 17–24 (17, 17, 19, 20, 24). Budding zone present in two specimens, between chaetae of XVII and XVIII (specimen with 24 segments) or XIV and XV (specimen with 20 segments). Posterior zooids with 6 or 7 segments. Dorsal bundles with 1 or 2 elongate, serrate hairs and 1 or 2 simple-pointed needles; more often just one hair and one needle per bundle. Hairs up to or more than three times as long as body diameter, increasing from II to VII to maximum length. From VII on, hair length 240–340 Μm, varying among bundles, segments and specimens. Hair length slightly reduced in hindmost 2 segments, not as much as in foremost segments. Hairs c. 2 Μm thick at base, tapering continuously ectad, less than 1 Μm thick at tip; serration distinct at x 250 magnification on convex side of bent chaeta, serration 'teeth'> 1 Μm apart, distance between teeth wider at base than towards the tip. Needles tightly attached to proximal hair shaft and not always well-distinguished, c. 40 Μm long, straight without nodulus, about 1.5 Μm thick in ental 4 / 5, strongly thinning out in ectal 1 / 5, bent backwards towards hair, ectal tip parallel to hair shaft. Tip simple-pointed at x 1000 magnification but somewhat widened. Chaetal follicle conspicuous, diameter c. 15 Μm. Ventral chaetae all alike, 37–40 Μm long, bifid, nodulus as a faint swelling at c. 2 / 5 from distal tip, teeth short, about 1.5 Μm long, upper tooth minutely shorter and thinner than lower; chaetae straight distally, only teeth bent here; proximal third of chaetae bent in opposite direction of teeth; in II 6–9 chaetae per bundle, varying among specimens, 5–7 in II–XI, in posterior body half 3–5 per bundle, decreasing posteriad from 5 to 4 or from 4 to 3. Prostomium without proboscis, rounded, longer than wide in one specimen, wider than long in all others (c. 50–70 Μm long and 70–80 Μm wide). Prostomial epithelium with several bilateral-symmetrical protuberances projecting entad into prostomial lumen, protuberances larger ventrally near mouth opening than dorsally, absent mid-ventrally. Epidermal gland cells seen in some specimens, laterally of and level to dorsal chaetal bundles. Pharyngeal pad with tenuous protractor and retractor muscles. Pharyngeal glands in separate packages at 3 /4, 4/ 5, and 5 / 6, enclosing septa. Gut diameter abruptly widened behind septum 6 / 7, from c. 80 to 180 Μm, not constricted in following segment (i.e. no stomach distinguishable), no cells with intracellular 'stomachal' canals distinguished. Dorsal blood vessel large in VI (and V), not seen posteriorly. Three simple and unbranched commissural vessels observed near 2 /3, 3/4, 4/ 5. Pars tumida of midgut present. First nephridium in VII, unpaired. Most of following nephridia unpaired as well, on alternating sides, 10 nephridia counted altogether in a specimen with 20 segments. Nephrostome present, on anterior face of 6 / 7 ventro-laterally; wide and densely packed loops in all of VII, dorsally and ventrally. Nephropores ventral, anterior to ventral chaetal bundles. Coelomocytes spherical, diameter 6–10 Μm, with glassy irregular texture, vesicles not distinguished. Remarks. Our specimens agree in all diagnostic details with Pristina terrena Collado & Schmelz, 2000. Noteworthy are the long serrate hairs that increase in length from II to VII and vary in length from VIII on, seemingly simple-pointed needles without nodulus and a tapering distal fifth bent towards the hair, and ventral chaetae that differ very little in size and shape between anterior and posterior bundles, with short and more or less equal-sized teeth. Further similiarities of P. terrena extend to body size, segment number, blood vessels, blood commissurals, and location of the first nephridium. The only difference of possible taxonomic importance is the widening of the intestine, described as gradual in P. t errena and abrupt in our specimens. However, the latter is a fixation artefact due to strong contraction of the animals when fixed in ethanol. We reinvestigated non-type ethanol-preserved reference specimens of the original series from Collado's personal collection (see Collado & Schmelz 2000), and there the intestine shows an abrupt widening as well. A stomach was not seen in the Honduras material and is absent in P. t e r ren a as originally described. The coelomocyte granulation, conspicuous in living specimens, is no longer seen in ethanol-preserved material, but the non-type reference material (see above) has the same irregular glassy texture without distinct vesicle; such a pattern can be indicative of coarse refractile granules as seen in live P. t errena. The same correspondence has been observed in enchytraeids (comp. Rota 2013). This is the first record of P. t er re n a after the original description from rain forest soils near Manaus, Brazil. The distance of approx. 3500 km between the two localities may suggest an extremely good dispersal ability of the species, but P. t e r re n a may also be common and widespread in Central and South America. Records of Pristina in Central and South America are scarce and mostly restricted to limnic, river and groundwater habitats, where this soil-dwelling species may not occur. Pristina species were regularly found in a non-flooded ("terra firme") primary forest plot of Amazonia, with six species described or recorded so far (Collado & Schmelz 2000, 2001, 2002; Augustsson 2001) and also in the Mata Atlântica of the Brazilian State Paraná (Römbke et al. 2005).Published as part of Schmelz, Rüdiger M., Jocque, Merlijn & Collado, Rut, 2015, Microdrile Oligochaeta in bromeliad pools of a Honduran cloud forest, pp. 508-526 in Zootaxa 3947 (4) on pages 518-519, DOI: 10.11646/zootaxa.3947.4.3, http://zenodo.org/record/24261
Gehyra rohan Oliver, Clegg, Fisher, Richards, Taylor & Jocque, 2016, sp. nov.
No full textGehyra rohan sp. nov. Figs 2–5 Holotype. RBINS 2684 (Field No. PNG 14-88), adult male, near Lorengau, Manus Island, Papua New Guinea (2.0388S, 147.2371E), collected 3 December 2014 by J. R. Clegg, P. N.Taylor and M. M. T. Jocque. Paratypes. AMS R129497 – 129498, adult females, Lombrum, Los Negros Island, Admiralty Islands, Papua New Guinea (2.01S, 147. 42E), collected 15 December 1951 by N. C. Goddard; CAS 252881, adult female, south bank of Lorengau River, 2.5 km southwest of Lorengau, Manus Island, Papua New Guinea (2.0415S, 147.2594E), collected 30 May 2010 by R. Fisher; PNGNM 25220 (PNG 14-158), adult female, Yiringou Village, Manus Island, Papua New Guinea (2.0833S, 147.1167E), collected 9 December 2014 by J. R. Clegg, P. N. Taylor and M. M. T. Jocque); RBINS 2685 (PNG 14-157), adult male, Yiringou Village, Manus Island, Papua New Guinea (2.0833S, 147.1167E), collected 9 December 2014 by J. R. Clegg, P. N. Taylor and M. M. T. Jocque; SAMA R69881 (SJR15105), adult male, near Nae, Mussau Island, Papua New Guinea (1.524S, 149.739E), collected 18 October 2015 by K. Aplin. Diagnosis. Gehyra rohan sp. nov. is distinguished from other Gehyra species by the following suite of characters: very large size (adult SVL 130–150 mm), large head (HW/SVL 0.18–0.22, HD/SVL 0.11–0.14), prominent skinfolds on the anterior forelimbs and posterior hind limbs, weak lateral fold, heterogeneous dorsal scalation consisting of large rounded scales bordered by numerous much smaller rounded or triangular scales, massive digital discs with high number of wide undivided subdigital lamellae (finger IV 23–25, toe IV 22–26) that are not deeply notched or divided, rostral with near horizontal dorsal edge and not deeply notched, precloacal and femoral pores in a moderately long single continuous chevron of up to at least 40 pores, original tail without lateral serrations, slightly compressed and with a prominent medial row of enlarged subcaudals, and a prominent ring of orange scales around the eye in life. Description of holotype. Adult male. Habitus, large (SVL 140.3 mm) and robust (Fig. 2). Head triangular, robust (HW/HL 0.87), moderately long (HL/SVL = 0.23) and deep (HD/HL = 0.47). Snout long and robust (EN/ HL = 0.35). Rostral large, broadly rectangular with rounded corners, rostral groove descends approximately 35% of rostral height, bifurcates, and extends to almost contact both nares (Fig. 3). Supranasals ovoid but with distinct points at ventrolateral edges, separated by three much smaller asymmetrical (becoming smaller from right to left) squarish internasals in a transverse series. Nares bordered by rostral, first supralabial, one large supranasal and three or four postnasals. Supralabials large and squarish with rounded dorsal edges, total number to inflexion of mouth 14 (left) and 13 (right), and total number to midpoint of eye 11 (both sides). Supralabials bordered dorsally by 2–3 rows of enlarged scales. Infralabials large and squarish, total number to rictus of jaw 14 on both sides. Mental triangular, bordered by two infralabials and two large rounded postmentals (Fig. 3). Scales on dorsal surface of head tiny, irregular and slightly conical, becoming larger and flatter laterally and anteriorly. Superciliaries forming a brillar fold of small spiniform scales extending along the dorsal border of the orbit from anteroventral to posterodorsal corners. Pupil partially dilated, somewhat elliptical with limited crenulations. Body robust (TrK/SVL 0.50). Dorsal and lateral scales distinctly heterogeneous in both shape and size, generally consisting of irregularly arranged large rounded scales, bordered by numerous much smaller rounded or triangular scales, often forming a ‘Star of David’ pattern; scales on nape much smaller than those on snout and torso. Ventral scales imbricate, arranged more regularly than those on dorsum, larger towards middle and posterior of venter, and tiny and granular on throat. Skin along ventrolateral edge of body loose and forming a weak fold along axilla-groin interval (Fig. 4). Precloacal and femoral pores (n = 40) arranged in a single recurved series terminating halfway along each femur. Hemipenal bulge present, moderately pronounced. Limbs robust and fleshy, with prominent lateral folds along anterior and posterior edge of forelimb, posterior edge of hindlimb, and less prominently on anterior edge of hindlimb (Fig. 4). Digits on both the fore- and hind limbs with prominent and expanded pads; terminal phalanges free and with well developed claws on all digits except finger I and toe I. Subdigital lamellae undivided, wide under expanded portion of disk, tapering and becoming narrower than toe proximal to the expanded disc (Fig. 3); total lamellar counts for all digits as follows (left/right): fingers I = 23/23, II = 24/25, III = 26/26, IV = 30/31, V = 27/26; toes I = 21/19, II = 24/26, III = 28/28, IV = 32/27, V = 24/22; total number of lamellae under expanded portion of the disk (left/right): fingers I = 22/21, II = 18/21, III = 19/19, IV = 21/24, V = 21/22; toes I = 21/19, II = 20/21, III = 22/23, IV = 23/23, V = 23/22. Webbing extending to base of disc on all digits, folded in preservative. Tail original, thin and short (107.8 mm in length) with blunt tip, much narrower than body at base; dorsal and lateral caudal scales granular and arranged irregularly, similar to dorsal scales on body; subcaudal scales distinctly enlarged, rounded, with a single medial row of 54 dilated (0.35–0.50 width of tail) ovoid scales extending full length of tail. Coloration in preservative. Dorsal and lateral surfaces grey with an indistinct chestnut wash and scattered darker grey maculations, generally corresponding to a single scale. Dorsal surfaces of limbs with coloration similar to dorsum of body, becoming distinctly darker distally and on posterior skinfolds of hindlimbs. Ventral surfaces of body, limbs and tail predominantly plain light tan. Paired pinkish-brown regions on anterior lateral edges of throat, and also forming two series of faint bars extending along lateral edges of venter, from posterior edge of insertion of the forelimbs to anterior edge of insertion of hindlimbs. Subdigital lamellae under expanded discs of all digits beige proximally, tending distinctly darker greyish-brown distally. Tail coloration as for body, but with more dense dark grey maculations on dorsal surfaces. Coloration in life. When initially captured: dorsal and lateral surfaces of head, body, limbs and tail dark chestnut brown mottled with patches of orange, light brown and off white, and with extensive black maculations, especially towards posterior region of dorsum and tail (Fig. 4). After capture: base dorsal color faded towards greyish but with same basic pattern (Fig. 4). Ventral surfaces of torso and limbs yellow, brightest anteriorly, undersurface of head brownish and digits white, transition between white of digits and yellow coloration of limbs relatively sharply defined, throat and torso both with regions of diffuse brown barring. Ventral surface of tail yellowish white with extensive dark-brown flecks. Scales around orbit forming a distinct orange ring, larger spiniform superciliary scales around dorsal edge bright reddish orange, grading to paler orange on smaller scales around ventral edge of eye. Details of holotype. Measurements (in mm): SVL 140.3; TL 107.8; TrK 69.8; HW 27.5; HL 31.6; HD 14.8; EN 11; EYE 6.5; IORB 12.4; POM 3.4; FA 16.9; CS 18.8. Meristic data: IN 3; SUPR 14; INFR 14; LAMF4 23; LAMT4 25; POR 40. Variation. Summary meristic values for all adults (2 males, 4 females) in the type series are as follows (mean, with the range in parentheses): SVL 138.4 (131.1–150.0); TL 107.2 (93.0–141.0); TrK 64.9 (58.0–72.6); HW 26.9 (24.5–29.8); HL 32.0 (30.2–35.0); HD 16.3 (14.8–18.80); EN 11.5 (10.9–13.2); EYE 7.6 (6.5–9.6); IORB 12.5 (11.4 –13.7); POM 3.4 (3.0–3.8); FA 16.5 (15.5 –17.3); CS 20.0 (17.7 –22.7). Summary scalation information for these same 6 individuals are as follows: SUPR (to midpoint of eye) 10.5 (9–11); SUPR (rictus of mouth) 13.9 (12– 14); INFR 12.7(11–14); LAMF4 22.9 (21–25); LAMT4 24.1 (22–26); POR 36.5 (33–40). The single male paratype SAMA R69881 (SJR15105) (Fig. 5) has a presumed developmental anomaly in which the pore-bearing and surrounding large scales have been shifted to the left, such that the right end of the pore series starts in the precloacal region, and extends almost fully along the left tibia, while the ventral scales on the right tibia are heterogeneous and do not appear to have formed properly. This specimen has two large, welldeveloped testes and in other respects appears to be a normal adult male. All adult specimens are similar to the holotype and share the key diagnostic traits including a bright orange ring extending around the eye, wide subcaudals under the original tail and prominent skinfolds on the arms and legs, although the prominence of the latter character varies with the angle of limb preservation. Smaller specimens tend to have a plainer ventral coloration and less obvious brown barring and mottling on the throat and ventrum in preservative, suggesting that this pattern is most pronounced in adult specimens. The largest specimens in the type series are all female (max SVL 150 versus maximum of 140 mm for the males), raising the possibility that the species is sexually dimorphic like several other Pacific Gehyra (Zug 2013), but given the very small number of males, more material is needed to confirm this. Based on photographs and field notes the dorsal coloration in life varies from quite dark chestnut brown to light grey, and the pattern of extensive but indistinct orange, greyish and brown mottling also varies in intensity. However much of this variation appears to be temporal, and single specimens vary extensively in appearance over short timescales (several hours) (Fig. 4). Comparisons. Only four other species of Gehyra approach the large size (adult SVL consistently> 120 mm) of Gehyra rohan sp. nov.: G. georgepottshaasti (Vanuatu), Gehyra marginata Boulenger, 1887 (Maluku), G. membranacruralis (Papua New Guinea) and G. vorax (Fiji). Based on published descriptions (Flecks et al. 2012) and field observations (RNF, SJR and Fred Kraus pers. com.) these taxa all lack a complete, bright orange ring around the eye in life, although occasional specimens of Gehyra vorax do have a yellow contour (as opposed to orange) around the dorsal edge of the eye (RNF pers. obs., for an example see page 172 in Ryan 1998). Gehyra rohan sp. nov. further differs from Gehyra marginata in having enlarged subcaudals under the original tail (versus absent), in having a chestnut brown iris (versus light green), and in having a less prominent ventrolateral dermal fringe on the body; from G. georgepottshaasti in having rounded postmentals (versus distinctly elongate) (Flecks et al. 2012); from Gehyra vorax in having a lower number of femoral pores in adult males (up to 40 versus 58–90) (Beckon 1992); and from Gehyra membranacruralis by its heterogeneous dorsal scalation consisting of large rounded scales separated by numerous much smaller rounded or triangular scales (versus large rounded scales only), and by having larger enlarged subcaudals (maximum anteroposterior length on adults> 2.5 mm versus 100 mm versus 100 versus <65 mm), and the absence of minute serrations along the lateral edges of the original tail (versus present). Distribution and ecology. Gehyra rohan sp. nov. is recorded from several localities across Manus Island. While most type material is from the east, one author (SJR) observed a very large Gehyra that is most likely this species at in lowland rainforest at Yeri River (2.001S, 146.819E) in north-western Manus (Fig. 6). Older material has also been collected from nearby Los Negros Island (see paratypes). This species has also been recorded from a single site on Mussau Island. The extent of its distribution, if any, beyond these islands remains unknown. Beckon (1992) reported a large Gehyra supposedly from Nauna Island near Manus (UPNG 5772), but noted that as it was collected from a banana box so its ultimate provenance was uncertain. Based on morphology, especially its high number of pores (62) Beckon further suggested that this animal is consistent with specimens from Fiji. Given uncertainty about provenance and morphology at this stage do not consider this a confirmed record of Gehyra rohan sp. nov. Gehyra rohan sp. nov. appears to be largely arboreal and is generally found in primary or disturbed lowland tropical rainforest (Fig. 6) on the trunks of large trees. It is also found on around human habitation in forested areas. Three of the authors (MJ, JRC, PT) found it to be reasonably common around Yiringou village in the interior of Manus, and two specimens were found on the same night on wooden beams below houses. The holotype was found running across a road in forest at night. The specimen from Mussau was found in a cave in disturbed forest close to the coast. One paratype (CAS 252881), from forest on the bluffs above the Lorengau River, that was initially ~6 meters high on the trunk of a tree, “glided” approximately 3 meters to an adjacent tree trunk when disturbed. Similar gliding or parachuting behavior has been observed in Gehyra mutilata (Heyer & Pongsapipatana 1970), and in numerous other genera of arboreal lizards, including many that lack obvious adaptations for gliding ( McGuire & Dudley 2011). Eytmology. Rohan is the Sohoniliu Village (Nali language) ‘tok ples’ (local language) name for this gecko. The community of Sohoniliu Village requested that this name be used for the formal description of this species, and we thank them for their support of this work.Published as part of Oliver, Paul M., Clegg, Jonathan R., Fisher, Robert N., Richards, Stephen J., Taylor, Peter N. & Jocque, Merlijn M. T., 2016, A new biogeographically disjunct giant gecko (Gehyra: Gekkonidae: Reptilia) from the East Melanesian Islands, pp. 61-76 in Zootaxa 4208 (1) on pages 65-71, DOI: 10.11646/zootaxa.4208.1.3, http://zenodo.org/record/20201
Zodariidae Thorell 1881
No full textKey to the genera of Zodariidae from tropical Asia 1. Femoral organ present on all legs, or at least legs II-IV (A)............................................................. 2 – Femoral organ absent......................................................................................................................... 5 2. Legs with dense cover of flattened incised setae (C); femoral organ single, on retro-lateral side (A)......................................................................................... Tropizodium Jocqué & Churchill, 2005 – Legs not covered with flattened incised setae; femoral organ double, on pro- and retrolateral side (B)...................................................................................................................................................... 3 3. AME less than half their diameter apart (E); anterior femora with prolateral wrinkles (F); chilum faint (females unknown)....................................................... Malayozodarion Ono & Hashim, 2008 – AME their diameter or slightly less apart (D); femora without wrinkles; chilum well developed... 4 4. Cymbium narrow, with strong retrolateral fold (G); ducts in epigyne intricately wound (H)...................................................................................................................... Asceua Thorell, 1887 – Cymbium wide, without retrolateral fold (I) but sometimes with pit; epigyne with few loops (J)..................................................................................................................... Suffasia Jocqué, 1991 5. Tarsi with two claws and claw tuft (K)......................................................... Hermippus Simon, 1893 – Tarsi with three claws (L), unpaired claw may be hidden by thick scopula in Storenomorpha........ 6 6. Eyes in three rows, 2-2-4 (M)............................................................................................................ 7 – Eyes in two rows 4-4 (N)................................................................................................................... 8 7. Highest part of carapace near fovea (O); body covered with sticky setae, covered with mud and soil debris......................................................................................................... Cryptothele L. Koch, 1872 – Highest part of carapace in eye region (P); without sticky setae..................... Cydrela Thorell, 1873 8. Posterior eye row strongly recurved (M); tarsal claws strongly curved and in alveolus; tarsi with well-developed scopulae ....................................................................... Storenomorpha Simon, 1884 – Posterior eye row straight or procurved (N); tarsal claws as usual; tarsi without scopulae.............. 9 9. Posterior ventral spines (PVS) in front of spinnerets absent (females only)............................................................................................................................................ Cambonilla Jocqué gen. et sp. nov. – Posterior ventral spines in front of spinnerets present (Q).............................................................. 10 10. Posterior ventral spines arranged in a single row (Q), normally situated on weakly sclerotized area................................................................................................................. Mallinella Strand, 1906 – Posterior ventral spines arranged in a group or more than one row (R)...........................................11 11. Carapace granulated with tiny branched setae; venter of abdomen with marbled pattern (T); anterior margin of sternum straight; apical surface of PVS serrated, consisting of two rows of minute denticles (visible in SEM only)....................................................................................................................... 12 – Carapace smooth or when granulated with simple setae; venter abdomen with pattern of longitudinal dark stripes (V); anterior margin of sternum with central concavity accommodating labium (U); apical surface of PVS smooth (visible in SEM only) (W)............................................................... 13 12. Carapace with thick longitudinal band of setae (S); without sclerotized protrusion around the pedicel............................................................ Heliconilla Dankittipakul, Jocqué & Singtripop, 2012 – Carapace without longitudinal band of setae; with sclerotized protrusion around pedicel (X) (males only)............................................................................................ Cambonilla Jocqué gen. et sp. nov. 13. Chelicerae with two small teeth on promargin; PLE more than three times their diameter from PME (Y); AER strongly pro-curved; carapace strongly domed and strongly granulated; dorsal scutum on opisthosoma large and ovoid, heavily sclerotized............. Euryeidon Dankittipakul & Jocqué, 2004 – Chelicerae without teeth; distance between PLE and PME not so wide (<2 times diameter); AER slightly procurved or straight; carapace not elevated and not strongly granulated; dorsal scutum on opisthosoma longitudinal, weakly sclerotized or absent; legs elongated........................................ 14 14. Carapace smooth and shiny; setae, if present, restricted to margins; F with basal, dorsal, tipped swelling bearing a spine (Z); dorsal scutum on opisthosoma longitudinal, weakly sclerotized........................................................................... Heradion Dankittipakul & Jocqué, 2004 – Carapace reticulated or rugose and provided with dispersed setae; F without basal swelling; dorsal scutum on abdomen indistinct ...................... Workmania Dankittipakul, Jocqué & Singtripop, 2012Published as part of Jocqué, Rudy, Jocque, Merlijn, Stock, Willem, Rin, Naroeun & Henrard, Arnaud, 2019, The new Southeast Asian genus Cambonilla gen. nov. (Zodariidae, Araneae): ' bis repetita placent', pp. 1-24 in European Journal of Taxonomy 501 on pages 5-7, DOI: 10.5852/ejt.2019.501, http://zenodo.org/record/257829
Paraphlebia hunnal Ortega-Salas & Gonzalez-Soriano 2022, sp. nov.
No full textParaphlebia hunnal Ortega-Salas & González-Soriano sp. nov. Figs. 8 (head ♂), 21 (thorax ♂), 77, 89 (appendages ♂), 98, 107 (posterior lobe of prothorax ♂), 151 (map). urn:lsid:zoobank.org:act: 74E83C0A-4560-4836-A57B-278697908431 Holotype. NICARAGUA • 1 ♂; Jinotega, Wiwilí, Cerro Kilambé, below double falls upstream of trailcrossing, near camp site; [13.587°N, 85.710°W]; 1250 m; 30 Jul 2001; González-Soriano, R. Beckemeyer leg.; IBUNAM; IBUNAM:CNIN:OD22722. Paratypes. NICARAGUA • 1 ♂; Jinotega, Wiwilí, Cerro Kilambé, below double falls upstream of trailcrossing, near camp site; [13.587°N, 85.710°W]; 1250 m; 30 Jul 2001; González-Soriano, R. Beckemeyer leg.; IBUNAM; IBUNAM:CNIN:OD22721 • same data but 1 ♂; Matagalpa, San Ramón, Stream 10 km SE of San Ramón; [12.862°N, 85.831°W]; 700 m; 18–20 Jun 1974; T. Donnelly leg.; TWD:CNIN:OD22720 • same data but 4 ♂♂; IBUNAM:CNIN:OD22723 to 22726. Etymology. Named hunnal /̠hʌnˈnɑl/ (L. noun in apposition), after Hun Nal (also Hun Hunahpú or Hun Nal Yeh), the Mayan god of maize, father of the twin gods Hunahpú and Ixbalanqué. Although no evidence of any strong Mayan influence in Nicaragua has been found, there is no doubt about the existence of an extensive commerce network which linked both regions. Description of holotype Head. Chiefly black, labrum dark glossy blue, anteclypeus with diffuse pale medial spot, postclypeus with pale blue on anterior half, frons with pale spots between suture with postclypeus and eye margin, vertex with two pale spots anterolateral to lateral ocelli, occiput with complete pale occipital bar, eyes dark brown to black; antennal scape with pale distal margin; antennae pedicel and flagellum brown. Thorax. Prothorax: chiefly pale on dorsum; middle lobe with medial black spot; see Figs. 98, 107 for morphology; propleuron black. Pterothorax: chiefly black with diffuse green metallic reflections over dark colouration on mesepisternum and mesepimeron; mediodorsal carina pale; see Fig. 21 for colour pattern; sternum pale with lateral black marks; legs pale, femora with light brown distal rings, tibiae light brown, tarsi light brown, claws reddish. Wings. Hyaline, slightly amber-tinted, FW with the vein that descends from the subnodus proximal to first post-quadrangular Vx by a distance of one-quarter of the first post-quadrangular cell; CuA in HW not forked, field between CuA and posterior margin with one supplementary vein. Px: FW 39/39, HW 36/34 Abdomen. Black with pale areas as follow: S2 with ventrolateral horizontal lines; S3–8 with basolateral spots; S9–10 and cerci pruinose on dorsum. Caudal appendages. Cerci: see Fig. 77 (paratype) for morphology. Paraprocts: superior lobe rudimentary, only distinguished by a transverse groove; inferior lobe in lateral view with an acute projection. Measurements. Abdomen: 41.5, FW: 35.3, HW: 34. Variation in males.In one paratype from Kilambé, prothorax with an extra diffuse pale line parallel to mediodorsal carina and second line on pterothorax wider, covering four-fifths of metepisternum towards antealar carina. Measurements. Abdomen: 42.3–47.3, FW: 36–38, HW: 33.3–38; Px: FW 37–42; Px: HW 28–36. Description of female. Female unknown. Diagnosis. This species is related to P. kinich and P. flinti and is diagnosed under the latter. Natural history. This species has been collected along small streams within montane cloud forest. Distribution. West Nicaragua: it has been collected in two localities on the Jinotega and Matagalpa departments in Nicaragua from 700–1250 m.a.s.l. This is the southernmost species of the genus.Published as part of Ortega-Salas, Héctor, González-Soriano, Enrique & Jocque, Merlijn, 2022, Untangling the waterfall damsels: a review of the Mesoamerican genus Paraphlebia Selys in Hagen, 1861 (Odonata: Thaumatoneuridae) with descriptions of 11 new species, pp. 1-66 in Zootaxa 5089 (1) on pages 14-15, DOI: 10.11646/zootaxa.5089.1.1, http://zenodo.org/record/583606
