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    FIGURES 11–14 in Three new species and six new records of Scathophagidae (Diptera) from Japan with a key to the Japanese genera

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    FIGURES 11–14. Cleigastra apicalis (Meigen). 11, Scutellum, dorsal view; 12, Male sternite V, ventral view; 13, Male terminalia, left lateral view; 14, Male terminalia, posterior view.Published as part of Iwasa, Mitsuhiro, 2021, Three new species and six new records of Scathophagidae (Diptera) from Japan with a key to the Japanese genera, pp. 531-553 in Zootaxa 4981 (3) on page 537, DOI: 10.11646/zootaxa.4981.3.6, http://zenodo.org/record/492141

    Megaphthalma Becker 1894

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    Genus Megaphthalma Becker, 1894 [Japanese name: Kiiro-funbae-zoku] Megaphthalma Becker, 1894: 105. Type-species: Scatomyza pallida Fallén, 1819, by original designation. Diagnosis. Head yellowish except for eyes; 3 fr, 3 or (1 proclinate, 2 latero-reclinate), 1 oc, 1 poc, 1 vte, 1 vti, vibrissal angle with 1 vibrissa; postpedicel short and rounded apically; palpi slender and clavate; scutum yellow to dark brown; thoracic sclerites yellow; 2 pprn, 2 npl, 1+2 ial, 1+2 sa, 2 pa, 2 +3 dc, 1 dsc; apsc absent; 1 prepst and 1 prepm; 4 anepst, 1 kepst; proepimeron sparsely with yellowish hairs; abdominal tergites yellow; male sternite V prominently bilobate posteriorly; male surstyli long and slender; male cerci clothed with setulae.Published as part of Iwasa, Mitsuhiro, 2021, Three new species and six new records of Scathophagidae (Diptera) from Japan with a key to the Japanese genera, pp. 531-553 in Zootaxa 4981 (3) on page 541, DOI: 10.11646/zootaxa.4981.3.6, http://zenodo.org/record/492141

    Spaziphora Rondani 1856

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    Genus Spaziphora Rondani, 1856 [Japanese name: Migiwa-funbae-zoku] Spaziphora Rondani, 1856: Dipt. Ital. Prodromus, 1: 99. Type-species: Cordylura hydromyzina Fallén, 1819: 7, by original designation. Diagnosis. Head whitish to black; 3-4 fr, 3 or (2 proclinate, 1 latero-reclinate), 1 oc, 1 poc, 1 vte, 1 vti, vibrissal angle with 1 vibrissa and 1-2 subvibrissae; postpedicel with acute or slightly angular upper corner; palpi distinctly spatulate and whitish; proepisternum and proepimeron with yellowish hairs; anepimeron covered with hairs in posterior half and row of black setae along posterior margin; thorax grey dusted; 2 pprn, 2 npl, 1+2 ial, 1+1 sa, 2 pa, 2+3 dc, 1 bsc, 1 apsc; 1 kepst; prepst and prepm absent; abdomen slender, elongate, and grey dusted; male sternite V with three pairs of tubercles; male surstyli short and stout, clothed with strong setae; female ovipositor short and cylindrical.Published as part of Iwasa, Mitsuhiro, 2021, Three new species and six new records of Scathophagidae (Diptera) from Japan with a key to the Japanese genera, pp. 531-553 in Zootaxa 4981 (3) on page 549, DOI: 10.11646/zootaxa.4981.3.6, http://zenodo.org/record/492141

    Chaetosa punctipes

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    Chaetosa punctipes (Meigen, 1826) [Japanese name: Kita-chibi-funbae] (Figs 7–10) Cordylura punctipes Meigen, 1826: 239. Type-locality: not given. Material examined. Hokkaido, 2 ♂, 1 ♀, Tsurui Village, Kushiro, 14. vi. 1975, leg. H. Sasaki. Diagnosis. Chaetosa punctipes Meigen can be recognized by the following characteristics: postpedicel with acute upper corner (Fig. 7); thoracic pleura greyish dusted; legs wholly yellow; posterior lobes on sternite V with minute spinules on inner parts (Fig. 8); male surstyli slender and tapering apically (Fig. 9); male cercus with 3 long setae beyond surstyli (Figs 9, 10). Body length (♂). 4.3–4.9 mm. Distribution. Europe, Mongolia, North America, Russia (the European part, Siberia, the Far East) (Bagachanova et al., 2016), and Japan (Hokkaido). New to Japan.Published as part of Iwasa, Mitsuhiro, 2021, Three new species and six new records of Scathophagidae (Diptera) from Japan with a key to the Japanese genera, pp. 531-553 in Zootaxa 4981 (3) on page 535, DOI: 10.11646/zootaxa.4981.3.6, http://zenodo.org/record/492141

    Cleigastra apicalis

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    Cleigastra apicalis (Meigen, 1826) [Japanese name: Hizaguro-funbae] (Figs 11–14) Cordylura apicalis Meigen, 1826: 236. Type-locality: not given. Material examined. Honshu, 2 ♂, 1 ♀, Mt. Hayachine, Iwate Pref., 17-18. vi. 1971, leg. S. Shinonaga; 1 ♂, Sawauchi Vill., Iwate Pref., 19. v. 1991, leg. T. Hasegawa; 1 ♂, 2 ♀, Mt. Shiritaka, Ishikawa Pref., 23. v. 1971, leg. S. Shinonaga; 1 ♂, Mt. Hakusan, Ishikawa Pref., 24. v. 1971, leg. S. Shinonaga. Diagnosis. This species can be characterized as follows: face, parafacial, and gena distinctly silvery shiny; fore coxae yellow; scutellum with apsc and dsc setae (Fig. 11); legs very bristled; fore femur yellow often with black stripe anteriorly with long black and pale hairs ventrally; female fore femur without pale hairs; mid and hind femora with black apical rings; posterior lobes on male sternite V with long hairs (Fig. 12); male surstyli apically broad in posterior view (Fig. 14); male cerci sclerotized and projected dorsally (Figs 13, 14). Body length (♂ ♀). 7.8–8.2 mm. Distribution. Europe (widespread), Iraq, Mongolia, Russia (widespread) (Ozerov & Krivosheina, 2015), and Japan (Honshu).Published as part of Iwasa, Mitsuhiro, 2021, Three new species and six new records of Scathophagidae (Diptera) from Japan with a key to the Japanese genera, pp. 531-553 in Zootaxa 4981 (3) on page 536, DOI: 10.11646/zootaxa.4981.3.6, http://zenodo.org/record/492141

    Megaphthalmoides Ringdahl 1936

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    Genus Megaphthalmoides Ringdahl, 1936 [Japanese name: Sesuji-funbae-zoku] Megaphthalmoides Ringdahl, 1936, 179. Type-species: Cordylura unilineata Zetterstedt, 1838: 727, by original designation. Diagnosis. Head yellowish except for eyes; 2-3 fr, 3 or (1proclinate, 2 latero-reclinate), 1 oc, 1 poc, 1 vte, 1 vti, vibrissal angle with 1 vibrissa and 1 subvibrissa (small); postpedicel short and rounded apically; palpi filiform and whitish; thorax yellow to light brown; scutum and prescutum often with dark longitudinal stripes; 2 pprn, 2 npl, 1+2 ial, 1+2 sa, 2 pa, 2+3 dc, 1 dsc, 1 apsc; 1 prepst and 1 prepm; proepiperon with yellowish hairs, 2-4 anepst, 1 kepst; abdomen basally flat and apically cylindrical; male sternite V prominently bilobate posteriorly; male surstyli long and slender; male cerci covered with strong setulae.Published as part of Iwasa, Mitsuhiro, 2021, Three new species and six new records of Scathophagidae (Diptera) from Japan with a key to the Japanese genera, pp. 531-553 in Zootaxa 4981 (3) on page 542, DOI: 10.11646/zootaxa.4981.3.6, http://zenodo.org/record/492141

    Microprosopa Becker 1894

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    Genus Microprosopa Becker, 1894 [Japanese name: Hoso-funbae-zoku] Microprosopa Becker, 1894, Berl. Ent. Z., 39:147. Type-species: Cordylura haemorrhoidalis Meigen, 1826: Syst. Beschr., 5: 237, by original designation. Diagnosis. Head blackish, only gena partly whitish; 2 fr, 3 or (2 proclinate, 1 latero-reclinate), 1 oc, 1 poc, 1 vte, 1 vti, vibrissal angle with 1 vibrissa and 1 subvibrissa; postpedicel short and rounded apically; palpi spatulate and whitish; proepimeron with yellowish hairs; 2 pprn, 2 npl, 1+2 ial, 1+2 sa, 2 pa, (1-2) +3 dc, 1 bsc, 1 apsc; 1 prepst (yellow) and 1 prepm (yellow); 3 anepst, 1 kepst; abdomen slender and elongate; male sternite V prominently bilobate posteriorly; male surstyli stout and tapered apically.Published as part of Iwasa, Mitsuhiro, 2021, Three new species and six new records of Scathophagidae (Diptera) from Japan with a key to the Japanese genera, pp. 531-553 in Zootaxa 4981 (3) on page 547, DOI: 10.11646/zootaxa.4981.3.6, http://zenodo.org/record/492141

    Chaetosa Coquillett 1898

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    Genus Chaetosa Coquillett, 1898 [Japanese name: Chibi-funbae-zoku] Chaetosa Coquillett, 1898: 163. Type-species: Cordylura punctipes Meigen, 1826: 239, by original designation. Diagnosis. Frons anteriorly yellow and posteriorly black; face, parafacial, and gena yellow; postpedicel yellow with acute apical corner, about 2.5 times as long as wide; arista pubescent; vibrissal angle with 1 vibrissa and 1 subvibrissa; palpus yellow without strong subapical seta; 2 fr, 3 or, 1 oc, 1 poc, 1 vte, 1 vti; thorax black; scutum and prescutum grey microtrichose: thoracic pleura greyish dusted; 1-2 pprn, 2 npl, 1+2 ial, 2+3 dc, 1+2 sa, 2 pa; 1 bsc, 1 apsc, 1 prepst, 1 prepm, 3 anepst, 2 kepst; vein A 1 +CuA 2 extending to wing margin; legs yellow; male sternite V bilobate posteriorly; male surstyli long and slender; male cerci with distinct long setae.Published as part of Iwasa, Mitsuhiro, 2021, Three new species and six new records of Scathophagidae (Diptera) from Japan with a key to the Japanese genera, pp. 531-553 in Zootaxa 4981 (3) on pages 534-535, DOI: 10.11646/zootaxa.4981.3.6, http://zenodo.org/record/492141

    Megaphthalmoides japonicus Ozerov 2008

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    Megaphthalmoides japonicus Ozerov, 2008 [Japanese name: Sesuji-funbae] (Figs 30–33) Megaphthalmoides japonicus Ozerov, 2008: 420. Type-locality: ”Izumi Katswiagi”, Osaka, Japan. Material examined. Hokkaido, 7 ♂, 1 ♀, Shirogane-onsen, Biei, 26. viii. 1979, leg. M. Iwasa; 1 ♂, Mt. Ponyaoromappu, Hidaka range, 11. viii. 1998, leg. M. Iwasa; 1 ♀, Mt. Tsurugi, Hidaka range, 11. viii. 1998, leg. M. Iwasa. Honshu , 3 ♂, Mt. Hayachine, Iwate Pref., 17-18. vi. 1971, leg. S. Shinonaga; 1 ♀, Karuizawa, Nagano Pref., 11. viii. 1970, leg. R. Kano; 1 ♀, Mt. Houou, Yamanashi Pref., 6. vi. 1972, leg. S. Shinonaga. Kyushu , 1 ♂, Izumimachimoniki, Yatsushiro City (alt. 1350 m), Kumamoto Pref., 11. v. 2016, leg. D. Kato. Diagnosis. This species is a large species and characterized as follows:characteristics: head yellow except for eyes; postpedicel with long-haired arista (Fig. 30); thorax yellow with 2 longitudinal stripes on prescutum to scutum; abdominal tergites blackish, covered with hair-like setulae and strong lateral setae; male sternite V posterior lobes which (Fig. 31) are basally broadened; male surstyli (Figs 32, 33) long and slender, with rows of setulae posteriorly; male cerci clothed with strong setulae. Body length (♂ ♀). 7.8–10.0 mm. Distribution. Japan (Hokkaido, Honshu, and Kyushu).Published as part of Iwasa, Mitsuhiro, 2021, Three new species and six new records of Scathophagidae (Diptera) from Japan with a key to the Japanese genera, pp. 531-553 in Zootaxa 4981 (3) on pages 542-543, DOI: 10.11646/zootaxa.4981.3.6, http://zenodo.org/record/492141

    Megaphthalma pallida

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    <i>Megaphthalma pallida</i> (Fallén, 1819) <p>[Japanese name: Seobi-kiiro-funbae]</p> <p>(Figs 28–29)</p> <p> <i>Cordylura pallida</i> Fallén, 1819: 8. Type-locality: “Westrogothis… Scania ” [Sweden].</p> <p> <b>Material examined.</b> <i>Hokkaido,</i> 2 ♀, Iwanai Riv., Obihiro, Tokachi, 19. ix. 1995, leg. M. Iwasa. <b>Diagnosis.</b> This species is recognized by the following characteristics: frons, face, parafacial, and gena yellow; prescutum and scutum dark brown; postpronotum and thoracic pleura yellow; scutellum yellow with a pair of apical setae (Fig. 28); legs wholly yellow; abdominal tergites wholly yellow with narrow black line on each posterior margin (Fig. 29).</p> <p> <b>Body length</b> (♀). 5.5–6.0 mm.</p> <p> <b>Distribution.</b> Europe, Japan (Hokkaido), Russia (the European part, Siberia, the Far East) (Bagachanova <i>et al</i>., 2016), North America. New to Japan.</p> <p> <b>Remarks.</b> Available specimens of this species are females, but as a result of my examination of specimens of <i>M. pallida</i> preserved in the Museum National d’Histoire naturelle (Paris), the characteristics agreed with those of female <i>M. pallida</i>.</p>Published as part of <i>Iwasa, Mitsuhiro, 2021, Three new species and six new records of Scathophagidae (Diptera) from Japan with a key to the Japanese genera, pp. 531-553 in Zootaxa 4981 (3)</i> on pages 541-542, DOI: 10.11646/zootaxa.4981.3.6, <a href="http://zenodo.org/record/4921413">http://zenodo.org/record/4921413</a&gt
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