263,273 research outputs found
Pseudonannolene rosineii Iniesta & Ferreira 2014, new species
No full textPseudonannolene rosineii Iniesta & Ferreira 2014, new species (Figs. 7, 14 e) Material examined. Holotype: 1 male (ISLA 4094) from Gruta Paranoá cave (20 º 21 ’ 57.13 ”S 45 º 40 ’ 11.15 ”W), Pains/MG, Brasil, 27 /I/ 2009. Collected by R. Zampaulo. Paratypes: 1 male (ISLA 4095) from Gruta Ninfeta III cave (20 º 20 ’ 17.78 ”S 45 º 36 ’ 55.90 ”W), Pains/MG, Brasil, 25 /I/ 2009. Collected by R. Zampaulo; 1 female (ISLA 4121) from Gruta Ninfeta III cave (20 º 20 ’ 17.78 ”S 45 º 36 ’ 55.90 ”W), Pains/MG, Brasil, 25 /I/ 2009. Collected by R. Zampaulo. Etimology. The specific epithet is in honor to Rosinei de Oliveira, from the city of Pains. Rosinei has been actively contributing to increase knowledge of the subterranean fauna of the region Pains, which stands out on the national scene as a major hotspot of subterranean diversity. Comparative diagnosis. Body and eyes pigmented. Labrum with 6 supralabral setae; 24 labral setae. Mandibles with 11 rows of pectinate lamellae. P. ro s i n e i i has the internal branch of the gonopod similar those observed in the species P. t a bo a, P. leopoldoi, P. ambuatinga (Iniesta & Ferreira 2013 b), P. s pe l ae a (Iniesta & Ferreira 2013 a), P. rolamossa, P. gogo (Iniesta & Ferreira 2013 c), P. chaimowiczi, P. imbirensis, P. tocaiensis (Fontanetti 1996 a) and P. microzoporus (Mauriès 1987). The squamous portion of the solenomere is bifurcated, as in P. taboa, P. leopoldoi, P. chaimowiczi, P. i m b i re ns i s, P. rolamossa, P. gogo, P. anapophysis (Fontanetti 1996 a), P. strinatii (Mauriès 1974) and P. tricolor (Brölemann 1902). The pre-femoral process is relatively larger than prefemur, as in P. taboa, P. rolamossa, P. gogo, P. ambuatinga, P. tocaiensis, P. mesai, P. leucocephalus, P. ophiulus, P. halophila, P. chaimowiczi e P. strinatii (Fontanetti 2002; Iniesta & Ferreira 2013 b; Iniesta & Ferreira 2013 c). Description of adults. Measurements: Length from 58 up to 68 mm; maximum midbody diameter between 3.96 to 4.48 mm; body rings ranging between 62 to 65; length of antennae ranging from 4.4 to 4.7 mm (relation to diameter ranging 1.04 to 1.11); length of legs 3.52 to 4 mm (relation to diameter ranging 0.88 to 0.89); length of tarsal claw 0.24 to 0.32 mm (relation to diameter ranging 0.06 to 0.07). Color: Visualization after fixation in 70 % alcohol. Bicolor, with the anterior region of each ring darker and posterior brownish yellow. Head (Fig. 7 a): Head glabrous and pigmented. Labrum with a row containing 24 labral setae, and above a row with 6 supralabral setae. Mandibles slightly pigmented, with 2 external teeth, 4 internal teeth and 11 rows of pectinate lamellae. Eyes with 27–30 ocelli arranged in 4 rows. Antennae pigmented and densely setose. First antennomere small, second, fourth, fifth and sixth similar, being the last larger. Third antennomere largest. Presence of basiconic sensilla in latter edge of fifth and sixth antennomere. Gnatochilarium typical of the genus. Trunk: Body pigmented. Prozone dark and metazone brownish yellow. Lateral region of each ring with transverse striae. Telson, anal shield and anal valve pigmented. First male pair of legs (Fig. 7 b): Coxae (Cx) larger; densely setose; triangle-shaped. Prefemur (Prf) with shorter oral process parallel (P) to the coxae. P with bristles arranged on base and base width equal to the distal region. Gonopod (Fig. 7 c, d): Gonopod short, stout and sclerified. Coxae reduced; glabrous and adhered to basal region of gonopod. Basal section (Bs) with width little larger than half of length; basiconic bristles arranged in rows along the entire the base of gonopod. Shoulder (Sh) evident and rounded. Distal section (Ds) as long as wide and little smaller than half of length of Bs. Solenomere (S) trianguliform; distal region squamous, bifurcated, with an acute external tip and rounded internal with a seminal spine (Sp). Internal branch (Ib) starting right below the Sh line on Bs. Ib like a shield and bristles exceeding the S. Notes on the natural history and habitat. The species is also distributed in caves in the region of Pains (Fig. 15 a), though it was never recorded in sympatry with P. robsoni. However, P. ros i n ei i is less frequent, occurring in a few caves compared to P. robsoni. Their populations also tend to be small, which clearly demonstrates that populations of P. rosineii and P. robsoni also occur outside of caves, wherein the caves do not necessarily make up their main habitats. The caves where populations of P. ro s i n e i i were observed are small and generally dry, the main organic resource being guano of Desmodus rotundus.Published as part of Iniesta, Luiz Felipe Moretti & Ferreira, Rodrigo Lopes, 2014, New species of Pseudonannolene Silvestri, 1895 from Brazilian limestone caves with comments on the potential distribution of the genus in South America (Spirostreptida: Pseudonannolenidae), pp. 361-397 in Zootaxa 3846 (3) on pages 370-371, DOI: 10.11646/zootaxa.3846.3.3, http://zenodo.org/record/25010
Pseudonannolene lundi Iniesta & Ferreira 2015
No full textPseudonannolene lundi Iniesta & Ferreira, 2015 (Fig. 1–3) Material examined: Holotype: 1 ♂ (ISLA 8684) from Lapa Sem Fim cave (UTM 23 K – 539.855 – 8.214.614), Luislândia/MG, Brazil, 17 /IV/ 2014. Collected by R. L. Ferreira. Paratypes: 2 ♂ (ISLA 8685, 8686) and 3 ♀ (ISLA 8687, 8688, 8689) from Lapa Sem Fim cave (UTM 23 K – 539.855 – 8.214.614), Luislândia/MG, Brazil, 17 /IV/ 2014. Collected by R. L. Ferreira. Etimology. The specific epithet honors the Danish naturalist Peter Wilhelm Lund, considered the founder of speleology as a science in Brazil. Peter Lund has worked for decades in the caves of Minas Gerais state, and made remarkable contributions to Brazilian paleontology. In addition, we intend to extend the honor to a caving group of the same name (Espeleo Grupo Peter Lund) for their contributions to our knowledge regarding the caves in the north of Minas Gerais state, as well as the support given to us during our collections in the area. Comparative diagnosis. P. lundi is similar to the species P. ambuatinga and P. spel aea in relation to depigmentation and reduction of body size, which have evolved as a result of restriction in the subterranean environment (Iniesta & Ferreira 2013 a, b). The gonopod of P. lundi resembles P. strinatii (Areias Cave, São Paulo), mainly in the structure of the distal section. The internal brach is more robust and apparent. The solenomere is sligthly trianguliform, varying in the format as in P. ambuantinga and P. saguassu (more trianguliform) in relation to P. strinatii, with a lateral/diagonal tip. Description of adults. Measurements: Length from 54 up to 61 mm; maximum midbody diameter between 2.4 and 2.8 mm; body rings ranging between 62 to 68; length of antennae ranging from 2.6 to 2.9 mm (relation to diameter ranging 1.03 to 1.08); length of legs 2.10 to 2.3 mm (relation to diameter ranging 0.82 to 0.87); length of tarsal claw 0.12 to 0.16 mm (relation to diameter ranging 0.05 to 0.06). General characteristics: Body whitish (Fig. 2). Trunk with metazona distinguished of prozona for a transversal suture and a weak difference in tonality. Tergites and collum depigmented. Lateral region of rings with transversal striae. Anal ring, valve and hypoproct depigmented. Head glabrous and depigmented. Labrum with a row of 15 labral setae and 6 supralabral setae. Mandibles depigmented with 2 external teeth evident, 4 internal teeth and 8 pectinate lamellae. Eyes depigmented with 27 to 33 ocelli; Antennae depigmented and densely setose. First antennomere smaller than others. Groups of basiconic sensilla on the edge of the fifth and sixth antennomeres (difficult to see). Four sensory cones on apical of sixth antennomere. Male characteristics: The first male pair of legs with elongated coxae (Cx), two times longer than wide; prefemoral process elongated and larger than pre-femur (Pf) (Fig. 3 C). Process (P) densely setose from the basis up to distal portion (Fig. 3 D). Gonopod elongated and well-sclerotized (Fig. 3 A, B). Coxae reduced (Cx); basal section (Bs) with length larger than width. Basiconic sensilla (B) arranged in rows along the internal edge of Bs. A short shoulder (not visualized in SEM). Distal section (Ds) with length similar to half of Bs. Internal branch (Ib) robust and enlarged; similar to a shield of solenomere (S). Some bristles of Ib exceeding the apex of S. S slightly trianguliform; a long spine (Sp) on apex and lateral tip rounded; evident squamous surface on external side. Notes. P. lundi n. sp. comprises the third troglobitic species of the genus described from Brazilian caves. This species presents some specific troglomorphic traits (morphological adaptation to the subterranean environment), such as pronounced depigmentation and a reduction of body size. Regarding these adaptations, some aspects deserve comments. Regarding the lack of pigmentation, it is worth noting that is undoubtedly a result of restriction in an aphotic environment, although some troglobitic species belonging to the genus do not necessarily have a notable depigmentation (personal observation). In addition to the depigmentation, the reduction of body size can be an indicator of the restrictions in the subterranean realm. Indeed, the three troglobitic species from the genus do not exceed 44 mm in length (maximum of 66 body rings) (Iniesta & Ferreira 2013 a, b), while some Brazilian nontroglobitic species, in average, have lengths around 61.5 mm (Iniesta & Ferreira 2013 a, b, Iniesta & Ferreira 2014). According to Culver et al. (1995, 2010), the reduction of body size in some cave-dwelling arthropod is known, in which the size depends of the specific habitat in caves, or its interactions with other groups of arthropods (e.g. predation). Regarding to the decrease on the number of ocelli, it did not prove to be necessarily a differential feature for the troglobitic species. Both troglobitic species, P. l u nd i and P. ambuatinga, have numbers of ocelli similar to other non-troglobitic Brazilian species (Mauriés 1974, 1987; Fontanetti 1996 a, b; Iniesta & Ferreira 2013 a, b; Iniesta & Ferreira 2014). The exception is P. s p e l a e a, which has a maximum of 15 ocelli (Iniesta & Ferreira 2013 a). The number of ocelli is quite variable between non-troglobitic species, varying from 20 to almost 50 (Iniesta & Ferrera 2013 b; Iniesta & Ferrera 2014). Accordingly, it is quite difficult to assume the presence of a reduced number of ocelli in any species, since for all cases the closest relative species is unknown, given the lack of a phylogenetic analysis for the genus. So, if the ancestor of P. lundi had around 30 ocelli, we could consider that a reduction did not occur. However, if this species descends from an epigean species with originally around 50 ocelli, the reduction would be obvious. Unfortunately, lack of knowledge regarding the phylogenetic relationships among the known species prevents us from actually assessing whether there was (or not) a reduction in the number of ocelli in troglobitic species from this genus. According to Iniesta & Ferreira (2013 a), another possible troglomorphic trait in the genus is the elongation of the sensilla basiconica on the antennae. In the species P. s p el ae a the elongation is remarkable in comparison to other non-troglobitic species from Brazil. However, to confirm the modification of this trait, some future work will be required. For P. lundi, the sensilla are larger than in other species, but no comparative measurement was made. Ecologial remarks. For many troglobitic myriapods, as centipedes, the knowledge regarding the environment in which the species live certainly contributes to establishing if the species is actually restricted to the subterranean environment (Ázara & Ferreira 2014). This is also true for Pseudonannolene, since there are many species (mostly non-troglobitic) living in caves. Accordingly, the species’ habitat should be characterized, since in some cases, the population can only be found in specific inner chambers of a cave. The Lapa Sem Fim cave comprises the largest cave in Minas Gerais state, with around 15 km of explored galleries. The cave morphology is quite unusual, with an intricate system of conduits. There are only two known entrances (Figure 1 A), each one located in the extremities of the only drainage (which is intermittent) existent in the cave. Most of the galleries are extremely dry (Figure 1 B), being located in an upper level in relation to the drainage conduit. In a few areas, there are some moist chambers, in which the humidity comes from percolating water. Such chambers are usually full of speleothems. Specimens of P. lundi were observed only in moist areas inside the cave, though in very distinct regions (Figure 1 C). No specimen was observed close to entrances. The biggest aggregation was observed in the middle part of the drainage conduit, although specimens were usually located in upper areas. In the inner portions of the cave, specimens were only found on those moist chambers, full of speleothems. Specimens seem to feed on distinct organic debris, as vegetal debris (mainly near the drainage conduit) or bat guano (all over the cave), though those resources were very rare, especially in areas far from the drainage. At least 5 other caves were sampled in the vicinity, but specimens of P. lundi were only observed in Lapa Sem Fim cave, suggesting that the species might be endemic to that cave. Although Brazilian caves are under risk due to the new legislation, the population of P. lundi seems to be relatively safe. The cave is rarely visited by locals and is well preserved. Furthermore, the large size of the cave, allied to its atypical morphology, are attributes that certainly enhance the relevance of the cave.Published as part of Iniesta, Luiz Felipe Moretti & Ferreira, Rodrigo Lopes, 2015, Pseudonannolene lundi n. sp., a new troglobitic millipede from a Brazilian limestone cave (Spirostreptida: Pseudonannolenidae), pp. 123-128 in Zootaxa 3949 (1) on pages 124-127, DOI: 10.11646/zootaxa.3949.1.6, http://zenodo.org/record/23904
Pseudonannolene marconii Iniesta & Ferreira 2014, new species
No full textPseudonannolene marconii Iniesta & Ferreira 2014, new species (Figs. 8, 14 d) Material examined. Holotype: 1 male (fragmented, ISLA 4106) from Gruta da Pedra Suspensa cave (15 º 23 ’ 31.42 ”S 39 º 44 ’ 18.41 ”W), Pau Brasil /BA, Brasil, 21 /I/ 2005. Collected by R.L. Ferreira. Etimology. The specific epithet is in honor of the biospeleologist Marconi Souza Silva for his contribution to our knowledge of Brazilian subterranean biology. It is noteworthy that this biologist developed work in caves of the Brazilian Atlantic Forest and he has contributed significantly to the expansion of knowledge of the subterranean fauna of this biome. Comparative diagnosis. Body and eyes pigmented. Labrum with 6 supralabral setae; 28 labral setae. Mandibles with 10 rows of pectinate lamellae (difficult to see). P. marconii also of the internal branch of gonopod like a shield, as those observed in the species P. ro s i n e i i, P. taboa, P. leopoldoi, P. ambuatinga (Iniesta & Ferreira 2013 b), P. s pe l ae a (Iniesta & Ferreira 2013 a), P. rolamossa, P. gogo (Iniesta & Ferreira 2013 c), P. chaimowiczi, P. imbirensis, P. tocaiensis (Fontanetti 1996 a; Fontanetti 1996 b) and P. microzoporus (Mauriés 1987). However, the solenomere is slightly bifurcated, being similar to P. rolamossa, P. gogo, P. anapophysis (Fontanetti 1996 a), P. strinatii (Mauriès 1974) and P. tricolor (Brölemann 1902). A notable difference in the gonopod is the elongated shape, being the basal region with length two times longer than the width. The pre-femoral process is proportionally longer than pre-femur, as in the species P. taboa, P. ros i n ei i, P. imbirensis, P. silvestri and P. tricolor (Fontanetti 2002). Description of adults. Measurements: Length from 58 mm; maximum midbody diameter between 5.63 mm; 60 body rings; length of antennae ranging 3.32 mm (relation to diameter 0.59); length of legs 3.82 mm (relation to diameter 0.68); length of tarsal claw 0.24 mm (relation to diameter 0.04). Color: Visualization after fixation in 70 % alcohol. Bicolor, with the anterior region of each ring darker and posterior yellow reddish. Head (Fig. 8 a): Head glabrous and pigmented. Labrum with a row containing 28 labral setae, and above a group of 13 supralabral setae. Mandibles slightly pigmented, with 2 external teeth, 4 internal teeth and 10 rows of pectinate lamellae. Eyes with 34 ocelli arranged in 4 rows. Antennae pigmented and densely setose. First antennomere small, second and third similar, fourth, fifth and sixth lower and similar, being the last larger. Presence of basiconic sensilla in latter edge of fifth and sixth antennomere. Gnatochilarium typical of the genus. Trunk: Body pigmented. Prozone dark and metazone yellow reddish. Lateral region of each ring with transverse striae. Telson, anal shield and anal valve pigmented. First male pair of legs (Fig. 8 b): Coxae (Cx) larger; densely setose; triangle-shaped. Prefemur (Prf) with elongated oral process parallel (P) to the coxae. Bristles arranged on base of P to the middle region; base wide and distal region short. Gonopod (Fig. 8 c, d): Gonopod elongated; thin and slightly sclerified. Coxae reduced; glabrous and adhered to basal region of gonopod. Basal section (Bs) with length little larger than two times the width. Presence of a short process between the coxae supporting a seta. Basiconic bristles arranged in rows along the entire the base of gonopod. Shoulder (Sh) short and rounded. Distal region (Ds) as wide as long and little short than half of length of Bs. Solenomere (S) with a short trunk, distal portion squamous; slightly bifurcated, with an acute external tip and rounded internal. Internal branch (Ib) stout, wide, and like a shield of S. Bristles arranged in almost all edge and exceeding the S. Notes on the natural history and habitat. The Pedra Suspensa cave is located in the carbonate group Rio Pardo, which occurs in the southeastern state of Bahia. This formation covers several caves; however, most of them are dry. Although the area is in Atlantic forest areas, deforestation is intense. The cave has a single entry and its conduit is almost entirely covered with guano produced by bats from different diets, among which stands out guano produced by Desmodus rotundus, a hematophagous species. The population of P. marconii is extremely large, dozens of specimens were observed throughout the cave, except for the area near the entrance. This condition differs from most species found in caves of Brazil, for which extremely large populations are not often observed. The species was found in the same karst region as P. tricolor (Trajano et al. 2000), though in different caves. The species was also found in Milagrosa (Fig. 15 f) and Califórnia caves, located in the same area.Published as part of Iniesta, Luiz Felipe Moretti & Ferreira, Rodrigo Lopes, 2014, New species of Pseudonannolene Silvestri, 1895 from Brazilian limestone caves with comments on the potential distribution of the genus in South America (Spirostreptida: Pseudonannolenidae), pp. 361-397 in Zootaxa 3846 (3) on pages 371-373, DOI: 10.11646/zootaxa.3846.3.3, http://zenodo.org/record/25010
Pseudonannolene rolamossa Iniesta & Ferreira 2013
No full text<i>Pseudonannolene rolamossa</i> Iniesta & Ferreira, 2013 <p>Figs 30D, 117–118, 164I, 166H, 172A, 178K, 187; Supp. file 4: Figs 217C, 219B</p> <p> <i>Pseudonannolene rolamossa</i> Iniesta & Ferreira, 2013c: 77, figs 2a–c.</p> <p> <i>Pseudonannolene rolamossa</i> – Gallo & Bichuette 2019: 48.</p> Diagnosis <p> Males of <i>P. rolamossa</i> differ from those of all other species of the genus by having a solenomere with a subtriangular and elongated ectal process exceeding in length the rounded apicomesal process (Fig. 118D).</p> Etymology <p>Noun in apposition, taken from the State Park “Parque Estadual do Rola Moça” where the species was found (Iniesta & Ferreira 2013c).</p> Material examined <p> <b>Holotype</b> BRAZIL • ♂; Minas Gerais, Nova Lima, Rola Moça I cave; [-20.020857, -43.812518]; 22 Mar. 2012; R.L. Ferreira and M. Souza-Silva leg.; ISLA 4004.</p> <p> <b>Paratypes</b> (total: 1 ♀) BRAZIL • 1 ♀; same collection data as for holotype; ISLA 4005.</p> <p> <b>Other material</b> (total: 7 ♂♂, 2 ♀♀, 4 immatures)</p> <p> BRAZIL – <b>Minas Gerais</b> • 1 ♂; Brumadinho, Serrinha 02 cave; [-20.151476, -44.201095]; 784 m a.s.l.; M.P. Oliveira leg.; ISLA 15054 • 1 ♂; PBR_18 cave; 15–20 Mar. 2010; R. Bessi <i>et al.</i> leg.; IBSP 5903 • 1 ♂, 2 immatures; Nova Lima, TUTA-14 cave; [-19.993344, -43.849412]; 763 m a.s.l.; M.P. Oliveira leg.; ISLA 15038 • 1 ♂; Rio Acima, ABOB_0028 cave; [-20.087775, -43.790650]; 743 m a.s.l.; 13 Jun. 2019; Equipe Spelayon leg.; IBSP 7766 • 1 immature; same collection data as for preceding; IBSP 7767 • 2 immatures; same collection data as for preceding; IBSP 7769 • 1 ♂; same collection data as for preceding; IBSP 7772 • 1 ♂, 2 ♀♀; same collection data as for preceding; IBSP 7773 • 1 ♂; Mariana, GS_25 cave; [-20.365015, -43.414773]; 780 m a.s.l.; 16 Jan.–11 Feb. 2011; Bessi <i>et al.</i> leg.; ISLA 6594.</p> Descriptive notes <p>MEASUREMENTS. 60–62 body rings (1–2 apodous + telson). Males: body length 56 mm; maximum midbody diameter 4 mm. Females: body length 58 mm; maximum midbody diameter 4.1 mm.</p> <p>COLOR. Body color brownish grey; head, antennae, collum, and legs darker; prozonites anteriorly greyish; metazonites with a medial darker band and a posterior lighter one.</p> <p>HEAD. Antennae long (Fig. 164I), just reaching back to end of ring 6 when extended dorsally; antennomeres elongated; relative antennomere lengths 1<2<3>4>5≈6>7. Mandibular cardo with ventral margin narrow. Ommatidial cluster well-developed, elliptical; ca 38 ommatidia in 6 rows.</p> <p>BODY RINGS. Collum with lateral lobes rounded, with ca 10 striae, strongly curved ectad (Fig. 117A). Very faintly constricted between prozonite and metazonite; prozonites smooth; metazonites laterally with transverse striae above ozopore in anterior body rings. Anterior sterna in midbody rings subrectangular, without transverse striae (Fig. 172A).</p> <p> FIRST LEG-PAIR OF MALES. Coxae (<i>cx</i>) short (less than half of remaining podomere lengths), subtriangular, with the base arched and expanded, densely setose (Fig. 118A); prefemoral process (<i>prf</i>) about as wide as half of prefemur, subcylindrical, densely setose up to its median region (Fig. 118B); remaining podomeres with setae along the mesal region.</p> <p> SECOND LEG-PAIR OF MALES. Coxa (<i>cx</i>) large and rounded; penis (<i>pn</i>) located at proximal region, rounded, not extended basally (Fig. 118C); prefemur compressed dorsoventrally; remaining podomeres setose.</p> <p> GONOPODS. Gonocoxa (<i>gcx</i>) elongated, twice longer than telopodite, with the base slightly arched; antero-posteriorly flattened (Fig. 118D–F); with rows of papillae mesally. Seminal groove (<i>sg</i>) curved; arising medially on mesal cavity and terminating apically on the seminal apophysis (<i>sa</i>). Shoulder (<i>sh</i>) long, subtriangular. Telopodite (<i>tp</i>) almost as wide as <i>gcx</i> (Figs 118D, 217C, 219B); solenomere (<i>sl</i>) with apicomesal process (<i>amp</i>) short, rounded; ectal process (<i>ep</i>) subtriangular, elongated, exceeding in length the <i>amp</i>; <i>sa</i> located at mesal portion, nearly not visible apically. Internal branch (<i>ib</i>) subtriangular, narrow, surrounding basally <i>tp</i> as a shield; with torsion of 180° in the distal portion but without projection; <i>ib</i> with setae along its entire margin slightly exceeding apically seminal region of <i>sl</i> (Fig. 118D–F).</p> <p>VULVAE. As typical for the genus. Bursa subtriangular, glabrous (Fig. 178K); internal valve subtriangular; operculum narrow; external valve wide, subtriangular.</p> Distribution <p>The species is widely distributed in iron ore caves and surrounding forests in the central region of Minas Gerais State, Brazil (Fig. 187).</p>Published as part of <i>Iniesta, Luiz Felipe Moretti, Bouzan, Rodrigo Salvador & Brescovit, Antonio Domingos, 2023, A reassessment of the Neotropical genus Pseudonannolene Silvestri, 1895: cladistic analysis, biogeography, and taxonomic review (Spirostreptida: Pseudonannolenidae), pp. 1-312 in European Journal of Taxonomy 867 (1)</i> on pages 77-79, DOI: 10.5852/ejt.2023.867.2109, <a href="http://zenodo.org/record/7891021">http://zenodo.org/record/7891021</a>
Pseudonannolene saguassu Iniesta & Ferreira 2013
No full text<i>Pseudonannolene saguassu</i> Iniesta & Ferreira, 2013. <p>(Figs. 7–10)</p> <p> <b>Material examined:</b> Holotype: 1 Male (Fragmented, ISLA 2266), from Gruta do Éden, Pains/MG, Brazil, 15/III/ 2012.</p> <p> <b>Etimology.</b> <i>Saguassu (saguaçú)</i> comes from the Tupi-Grarani origin (Brazilian Indian languages), meaning “those with big eyes”. This name obviously refers to the numerous ocelli observed in this species. It is to be treated as a noun in apposition.</p> <p> <b>Diagnosis.</b> Body with 69 rings and brownish coloration. About 45 Ocelli, pigmented. 30 labral setae and mandible with 11 pectinate lamellae. Gonopod with rhomboid solonomere and a central spine. Basal section 0.75 times longer than wide and distal section about 0.5 times longer than wide.</p> <p>Measurements: Length of 65.16 mm; maximum midbody diameter of 3.68 mm; 69 body rings; length of antennae of 3.67 mm (relation to diameter 0.99); length of legs 2.60 mm (relation to diameter 0.71); length of tarsal claw 0.2 mm (relation to diameter 0.05).</p> <p>Color: Brownish.</p> <p> <b>Description of adults.</b> <i>Head</i> (Fig. 7): Head glabrous and pigmented. Three small labral teeth, a row of 30 labral setae and above a row of 6 supra labral setae (one apparently missing). Mandibles pigmented, glabrous and with 2 external teeth above, 5 internal teeth and 11 pectinate lamellae. Eyes pigmented with 45 ocelli, distributed into 5–6 rows. Antennae pigmented and densely setose. First antennomere small, second and third antennomeres of similar sizes, fourth and fifth antennomeres shorter than third. Sixth antennomere longer and wider than the fourth and fifth. Groups of basiconic sensilla on the edge of the fifth and sixth antennomeres. Four terminal sensory cones.</p> <p> <i>Gnathochilarium</i> (Fig. 8 A; B): <i>Gula</i> (<b>Gu</b>) with short setae. <i>Mentum</i> (<b>Me</b>) rounded and with base wider than the latter. <i>Stipes</i> (<b>St</b>) with base shorter than distal region. <i>Promentum</i> (<b>Pme</b>) with two trianguliform parts and <i>Laminae linguales</i> (<b>Ll</b>) entirely separated by <b>Pme</b>.</p> <p> <i>Trunk:</i> Body with 69 rings. Tergites and collum pigmented (homogeneously brownish). Lateral transverse striae present (striae variables into individuals). Anal shield and anal valve pigmented like the trunk.</p> <p> <i>First male pair of legs</i> (Fig. 9 A): The first leg pair is modified and densely setose. Coxae (<b>Cx</b>) with basis longer than distal region. Distal region with a shoulder. Sternum not visible. Prefemur (<b>Prf</b>) with thin oral process parallel (<b>P</b>) to the coxa, more rounded in distal (Fig. 9 B). Post-femur (<b>Psf</b>) and tíbia (<b>Tb</b>) reduced. Femur (<b>F</b>) elongated. Terminal claw (<b>Tc</b>) not modified.</p> <p> <i>Gonopod</i> (Fig. 10 A; B): Gonopod short and stout. Coxae (<b>Cx</b>) little visible and glabrous. The basal section (<b>Bs</b>) of telopodite glabrous, 0.75 times longer than wide. Shoulder absent. Basiconic sensilla (<b>B</b>) extending along the membranous margin from basal section. Distal section (<b>Ds</b>) about 0.5 times longer than width and with two well-developed processes, the solenomere (<b>S</b>) and an internal (seminal) branch (<b>Ib</b>). <b>S</b> rhomboid and with a squamous surface coating the structure. Spine (<b>Sp</b>) present in center of the apex. <b>Ib</b> (coxosternal branch?) digitiform and supporting setae at distal region that exceeds the length of <b>S</b>. Short seminal groove (<b>Sg</b>) between <b>S</b> and <b>Ib</b>.</p> <p> <b>Remarks</b>. Morphological analysis was performed by stereoscopic observations and light microscopy in a single male specimen. The specimen was found in an area between the twilight and aphotic zones of Gruta do Éden, near the vertical entrance (Fig. 1 B). In the same cave, there is a huge population of <i>P. ambuatinga</i>, which is distributed especially in the main conduit of the cave, however, cut by a stream. Accordingly, this cave represents the first case of syntopic species of <i>Pseudonannolene</i> occurring in Brazilian caves. However, it is very likely that <i>P. saguassu</i> actually represents an “accidental” species, since only a single specimen was found during many visits paid to the cave. Since the external (epigean) environment was not properly sampled, it is plausible to assume that the main habitat of this species is the external semi-deciduous forest which covers the area. Therefore, it is not possible to consider the species as “rare” based on the fact that only a single specimen was collected.</p>Published as part of <i>Iniesta, Luiz Felipe Moretti & Ferreira, Rodrigo Lopes, 2013, Two new species of Pseudonannolene Silvestri, 1895 from Brazilian limestone caves (Spirostreptida: Pseudonannolenidae): synotopy of a troglophilic and a troglobiotic species, pp. 357-369 in Zootaxa 3702 (4)</i> on pages 363-364, DOI: 10.11646/zootaxa.3702.4.3, <a href="http://zenodo.org/record/217312">http://zenodo.org/record/217312</a>
Pseudonannolene leopoldoi Iniesta & Ferreira 2014, new species
No full textPseudonannolene leopoldoi Iniesta & Ferreira 2014, new species (Figs. 3, 4, 14 b) Material examined. Holotype: 1 male (ISLA 4123) from Lapa do Zu cave (17 º01’ 17.79 ”S 44 º 29 ’ 59.84 ”W), São João da Lagoa/MG, Brasil, 25 /IX/ 2013. Collected by R. L. Ferreira. Paratypes: 2 males (ISLA 4124, 4125) from Lapa do Zu cave, São João da Lagoa/MG, Brasil, 25 /IX/ 2013. Collected by R. L. Ferreira. 3 females (ISLA 4126, 4127, 4128) from Lapa do Zu cave, São João da Lagoa/MG, Brasil, 25 /IX/ 2013. Collected by R. L. Ferreira. Etimology. The specific epithet is in honor of the biospeleologist Leopoldo Ferreira de Oliveira Bernardi for his contribution to our knowledge of Brazilian subterranean biology, especially cave mites. Comparative diagnosis. Body and eyes pigmented. Labrum with 6 supralabral setae; 18–22 labral setae. Mandibles with 10–11 rows of pectinate lamellae (difficult to see). P. leopoldoi has an internal branch of the gonopod similar to those observed in the species P. taboa, besides the P. ambuatinga (Iniesta & Ferreira 2013 b), P. spelaea (Iniesta & Ferreira 2013 a), P. rolamossa, P. gogo (Iniesta & Ferreira 2013 c), P. chaimowiczi, P. imbirensis, P. tocaiensis (Fontanetti 1996 b) and P. microzoporus (Mauriès 1987). The bifurcated form of the solenomere and the presence of a seminal spine on the internal apex of the branch are similar to what is observed in the species P. taboa, P. chaimowiczi, P. imbirensis, P. rolamossa, P. gogo, P. anapophysis (Fontanetti 1996 a), P. strinatii (Mauriès 1974) and P. tricolor (Brölemann 1902). About the pre-femoral process, that is rounded: its size is shorter when compared to the pre-femur, and similar to those occurring in the species P. imbirensis, P. silvestri e P. tricolor (Fontanetti 2002) and P. s p e l a e a (Iniesta & Ferreira 2013 a). Description of adults. Measurements: Length from 48 up to 55 mm; maximum midbody diameter between 2.9 to 3.3 mm; body rings ranging between 62 to 64; length of antennae ranging from 3 to 3.2 mm (relation to diameter ranging 1 to 1.03); length of legs 2.3 to 2.5 mm (relation to diameter ranging 0.75 to 0.79); length of tarsal claw 0.14 to 0.16 mm (relation to diameter ranging 0.04 to 0.05). Color: Visualization after fixation in 70 % alcohol. Reddish color, with the anterior region of each ring darker and posterior ranging from light brown to reddish. Head (Fig. 3 a): Head glabrous and pigmented. Labrum with a row containing 18–22 labral setae, and above a row with 6 supralabral setae. Mandibles slightly pigmented, with 2 external teeth, 4 internal teeth and 10–11 rows of pectinate lamellae. Eyes with 28–36 ocelli arranged in 5 rows. Antennae pigmented and densely setose. First antennomere small, second and third similar, fourth, fifth and sixth lower and similar, being the last larger. Presence of basiconic sensilla in latter edge of fifth and sixth antennomere. Gnatochilarium typical of the genus. Trunk: Body pigmented. Prozone dark and metazone ranging from light brown to reddish. Lateral region of each ring with transverse striae. Telson, anal shield and anal valve pigmented. First male pair of legs (Fig. 3 b): Coxae (Cx) larger; densely setose; square-shaped. Prefemur (Prf) with shorter oral process parallel (P) to the coxae. Bristles arranged on base of P. Gonopod (Fig. 3 c, d): Gonopod stout and sclerified. Coxae reduced; glabrous and adhered to basal region of gonopod. Presence of a process supporting a seta. Basal section (Bs) with width little larger than half of length; Basiconic bristles arranged in rows along the entire the base of gonopod. Shoulder (Sh) evident and rounded. Distal section (Ds) little shorter than the length of Bs and with width little larger than half of length. Solenomere (S) elongated and with trunk wide; shorter squamous region; bifurcated, with an acute external tip and rounded internal. Internal branch (Ib) wide, starting right below the Sh line on Bs; like a shield of S and bristles not exceeding the S. Notes on the natural history and habitat. The Lapa do Zu cave (Fig. 15 c) comprises a limestone cave of considerable volume. It presents a single entry and receives organic material from the external environment, especially due to floods that occur during the rainy season. Thus, a large amount of plant debris is noticeable in various parts of the cave. In its distal portion, there is a large gap that leads to a drain (a small stream) that appears and disappears amid blocks of rock. This considerably increases the humidity of this portion of the cave. Individuals of P. leopoldoi were observed throughout the entire cave, and were associated with plant debris and deposits of bat guano. The fact that several specimens were associated with the large carcasses of a grasshopper (Caelifera), left in the cave by insectivorous bats (Fig. 3) is noteworthy. Such individuals, in many cases, clustered on one carcass, apparently feeding on fungus that developed in profusion in these carcasses. Pseudonannolene robsoni Iniesta & Ferreira 2014 , new species (Figs. 5, 6, 14 c) Material examined. Holotype: 1 male (ISLA 4080) from Gruta Água Limpa cave (20 º 27 ’06.99”S 45 º 39 ’ 10.58 ”W), Pains/MG, Brasil, 28 /V/ 2009. Collected by R. Zampaulo. Paratypes: 1 male (ISLA 4083) from Gruta do Bicho Desconhecido cave (20 º 24 ’ 20.04 ”S 45 º 35 ’ 24.70 ”W), Pains/MG, Brasil, 04/IV/ 2009; 1 male (ISLA 4084) from Gruta Loca dos Negros cave (20 º 24 ’05.99”S 45 º 39 ’ 49.49 ”W), Pains/MG, Brasil, 21 /III/ 2009; 1 male (ISLA 4085) from Gruta Duas Bocas cave (20 º 22 ’07.15”S 45 º 41 ’ 10.10 ”W), Pains/MG, Brasil, 01/IV/ 2009. Collected by R. Zampaulo. Other material examined. 1 male (ISLA 4079) from Gruta Zé da Fazenda cave (20 º 22 ’ 10.78 ”S 45 º 40 ’07.44”W, Pains/MG, Brasil, 09/III/ 2009; 1 male (ISLA 4081) from Gruta das Cerâmicas cave (20 º 24 ’ 15.21 ”S 45 º 35 ’ 51.15 ”W), Pains/MG, Brasil, 28 /V/ 2009; 1 male (ISLA 4082) from Gruta Fumaça III cave (19 º 28 ’ 29.68 ”S 44 º 19 ’ 41.31 ”W), Pains/MG, Brasil, 12 /II/ 2009; 1 male (ISLA 4086) from Gruta Tio Rafa II cave (20 º 24 ’ 47.04 ”S 45 º 39 ’ 52.73 ”W), Pains/MG, Brasil, 24 /I/ 2009; 1 male (ISLA 4087) from Gruta Ninfeta de Baixo cave (20 º 20 ’ 18.69 ”S 45 º 36 ’ 55.67 ”W), Doresópolis/MG, Brasil, 25 /I/ 2009; 1 male (ISLA 4088) from Gruta Ninfeta de Baixo cave (20 º 20 ’ 18.69 ”S 45 º 36 ’ 55.67 ”W), Doresópolis/MG, Brasil, 25 /I/ 2009; 1 male (ISLA 4089) from Gruta Cinderela cave (20 º 26 ’ 45.51 ”S 45 º 35 ’ 59.83 ”W), Pains/MG, Brasil, 18 /IX/ 2009; 1 male (ISLA 4090) from Gruta Dolina dos Angicos cave (20 º 25 ’05.92”S 45 º 40 ’ 43.74 ”W), Pains/MG, Brasil, 25 /VI/ 2009; 1 male (ISLA 4091) from Gruta Capoeirão cave (20 º 21 ’ 55.34 ”S 45 º 40 ’ 15.”W), Pains/MG, Brasil, 22 /I/ 2009. Etimology. The specific epithet is in honor of the biospeleologist Robson de Almeida Zampaulo for his contribution to our knowledge of Brazilian subterranean biology. It is noteworthy that the work of this biologist, developed in the region of Pains, has contributed significantly to the expansion of knowledge of the subterranean fauna of this important Brazilian karst region. Comparative diagnosis. Body and eyes pigmented. Labrum with 6 supralabral setae; 19–23 labral setae. Mandibles with 10 rows of pectinate lamellae (difficult to see). P. robsoni has the internal branch of the gonopod slightly similar to a shield, being its medium-distal portion twisted over the solenomere, besides the presence of an elongated apical projection on the branch. The pre-femoral process is similar to that observed in the species P. leopoldoi, P. imbirensis, P. silvestri, P.tricolor e P. spelaea due to the reduced size compared to the pre-femur (Fontanetti 2002; Iniesta & Ferreira 2013 a). Description of adults. Measurements: Length from 70 up to 88 mm; maximum midbody diameter between 4.8 to 5.8 mm; body rings ranging between 65 to 70; length of antennae ranging from 5.2 to 5.8 mm (relation to diameter ranging 1.07 to 0.99); length of legs 3.2 to 4 mm (relation to diameter ranging 0.66 to 0.68); length of tarsal claw 0.16 to 0.32 mm (relation to diameter ranging 0.03 to 0.05). Color: Visualization after fixation in 70 % alcohol. Bicolor, with the anterior region of each ring darker and posterior reddish yellow. Head (Fig. 5 a): Head glabrous and pigmented. Labrum with a row containing 19–23 labral setae, and above a row with 6 supralabral setae. Mandibles slightly pigmented, with 2 external teeth, 4 internal teeth and 10 rows of pectinate lamellae. Eyes with 36–43 ocelli arranged in 5 rows. Antennae pigmented and densely setose. First antennomere small, second and fourth similar, third longer, fifth and sixth little smaller than second and fourth, being the sixth larger. Presence of basiconic sensilla in latter edge of fifth and sixth antennomere. Gnatochilarium typical of the genus. Trunk: Body pigmented. Prozone dark and metazone reddish yellow. Lateral region of each ring with transverse striae. Telson, anal shield and anal valve pigmented. First male pair of legs (Fig. 5 b): Coxae (Cx) larger; densely setose; square-shaped. Prefemur (Prf) with shorter oral process parallel (P) to the coxae. P with bristles arranged on base and base width equal to the distal region. Gonopod (Fig. 5 c, d): Gonopod short, stout and sclerified. Coxae reduced; glabrous and adhered to basal region of gonopod. Basal section (Bs) with width little larger than half of length; basiconic bristles arranged in rows along the entire the base of gonopod. Shoulder (Sh) evident and rounded. Distal section (Ds) slightly longer than wide and little smaller than half of length of Bs. Solenomere (S) trianguliform; elongated and little wide; trunk elongated and glabrous. Distal region squamous and slightly bifurcated, with an acute external tip and rounded internal with a seminal spine (Sp). Internal branch (Ib) with small base and a thin distal region, starting right below the Sh line on Bs. Ib projecting in order to interlace the S and exceeding his length. Longer bristles arranged on edge of Ib; presence of a swollen apical projection Notes on the natural history and habitat. The karst region where populations of P. robsoni were found is considered an area of environmental, historical and economic importance. Furthermore, this region stands out in terms of harboring the greatest concentration of limestone caves known in Brazil at the present moment (over 1,200 recorded caves). However, since the 1960 s the area has undergone intense changes due to mining activities, which remove the limestone to produce cement and lime. Such activities are causing irreversible impacts on the landscape. In addition, it also highlights the intense changes suffered by the vegetation of the area (especially due to the expansion of agricultural activities), which has resulted in a rather fragmented landscape, where remaining vegetation is associated with only the tops of outcrops (inappropriate areas for human use). There are alterations in drainages near many caves, that in association with the loss of the original vegetation, may be leading to an intense change in import of organic resources to many caves (primarily dependent on imported organic resources from the external environment). The extraction of limestone has also often led to direct changes in the caves, ranging from partial destruction of conduits to the complete destruction of caves (Zampaulo 2010). This species is well distributed in caves in the region, although populations of every cave are often reduced. Exception is made for Brega cave (Fig. 15 i), where numerous individuals were found associated with the guano of hematophagous bats (Desmodus rotundus) and rodent feces (Fig. 6).Published as part of Iniesta, Luiz Felipe Moretti & Ferreira, Rodrigo Lopes, 2014, New species of Pseudonannolene Silvestri, 1895 from Brazilian limestone caves with comments on the potential distribution of the genus in South America (Spirostreptida: Pseudonannolenidae), pp. 361-397 in Zootaxa 3846 (3) on pages 365-369, DOI: 10.11646/zootaxa.3846.3.3, http://zenodo.org/record/25010
Retrato de "Iniesta" con mano recargada sobre pedestal
No full textI. O. Anverso: "Abril-14". Reverso: "ORIGINAL Iniesta Gral Culiacan Sinaloa 352"
Taula rodona 1 de la 'Jornada La Promoció de la Salut i el Treball en Xarxa'
No full textLa taula il·lustra tres experiències de treball en xarxa vinculades a nivell internacional : Cristina Iniesta, coordinadora de la Xarxa Catalana d’Hospitals i Serveis Promotors de la Salut; Carmen Gallardo, coordinadora de la Red Española de Universidades Saludables; Enric Llorca, president de la Red Española de Ciudades Saludables; totes elles xarxes implantades al nostre territori1451.mp4
1451.mp
Ruanda 1994, el genocidi. Les interpretacions de la qüestió ètnica
No full textTreballs Finals de Grau en Història, Facultat de Geografia i Història, Universitat de Barcelona, Curs: 2012-2013, Tutor: Ferran Iniesta VernetAquest treball és una introducció a la interpretació teòrica de la qüestió ètnica al
genocidi ruandès de 1994. S'exposen les diverses corrents teòriques sobre l'etnicitat a través de l'anàlisi històric del conflicte i de la reconciliació social després de la pau
Folcloredesmus thomasi Bouzan & Iniesta & Brescovit 2021, n. sp.
No full text<i>Folcloredesmus thomasi</i> n. sp. <p>Figures 26, 27C–D, 28</p> <p>urn:lsid:zoobank.org:act: 26862968-716B-47D3-9090-416B8C209ABA</p> <p> <b>Type material.</b> Male holotype from Fazenda São Nicolau (09° 50’ 25”S, 58° 14’ 53”W), Cotriguaçu, Mato Grosso, Brazil, 02.XI.2014, M. Karam-Gemael coll., deposited in IBSP 7788. Paratypes: four males and two females from Fazenda São Nicolau, Cotriguaçu, Mato Grosso, Brazil, 11.XII.2009, L. D. Batirola coll., deposited in IBSP 5452 and IBSP 5453.</p> <p> <b>Additional material.</b> BRAZIL: <i>Mato Grosso</i>: Cotriguaçu, Fazenda São Nicolau (09° 50’ 25”S, 58° 14’ 53”W), 2♂, X.2017 (CZUFMT 849); Cotriguaçu, Fazenda São Nicolau (09° 50’ 25”S, 58° 14’ 53”W), 10♂, 02.XI.2014, M. Karam-Gemael coll. (CZUFMT 833).</p> <p> <b>Diagnosis.</b> The same characteristics of the genus.</p> <p> <b>Description.</b> Male (Holotype, IBSP 7788). Coloration (long-preserved in 70% ethanol): Head and antennae ochre with the seventh antennomere broken. Body ocher with the edges of the paranota whitish (Fig. 26A–C), however in recently collected specimens the body’s coloration is reddish brown with the edge of the paranota yellowish (Figs 1A–B). Legs ochre whitish. Telson ocher. Total length: 55.69. Total width: 7.46. Collum 2.66 length, 7.65 width. Antennomere lengths (1>7): 0.62; 1.62; 1.49; 1.54; 1.55; 1.64; 0.26. Podomeres lengths (1>7): 0.99; 1.42; 2.33; 1.29; 1.52; 1.85; 0.47. Gonopod aperture 1.54 length, 3.01 width. Telson 1.20 length. Gonopod: 2.37 length, 2.37 width. Gonocoxae: 1.13 length, 1.26 width. A small spiniform process present (Fig. 26D). Telopodite: 2.30 length, 0.87 width. Prefemoral region 1/3 the size of telopodite. Prefemoral process (Figs 26D–F) long, narrow and falciform, starting from an angle of 135°, ascending parallel to the solenomere and partially covering it in ventral view; in ectal view, with indentations in the apical portion. Cingulum medially positioned (Fig. 26F). Solenomere long and thin with apex acute and slightly curved (Fig. 26F).</p> <p>Female (Paratype, IBSP 5453). Body as in male. Total length: 67.74. Total width: 8.90. Collum 3.40 length, 8.27 width. Epigyne short and triangular with irregular edges. Vulvae (Figs 27C–D): 1.21 length, 0.75 width. External valve: 0.89 length, 0.40 width. Internal valve: 0.88 length, 0.37 width. Operculum: 0.30 length, 0.35 width.</p> <p> <b>Distribution.</b> Known only from the type locality (Fig. 28).</p> <p> <b>Etymology.</b> The species epithet is taken as a noun in apposition honoring a very important person in the life of the first author, who has a passion for stories of Brazilian folklore. Noun in apposition.</p>Published as part of <i>Bouzan, Rodrigo Salvador, Iniesta, Luiz Felipe M. & Brescovit, Antonio Domingos, 2021, Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae), pp. 201-256 in Zootaxa 4970 (2)</i> on pages 246-247, DOI: 10.11646/zootaxa.4970.2.1, <a href="http://zenodo.org/record/4761544">http://zenodo.org/record/4761544</a>
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