892 research outputs found
Diplocephalus guidoi Frick & Isaia 2012
<i>Diplocephalus guidoi</i> Frick & Isaia, 2012 <p>LITERATURE. — Frick & Isaia (2012).</p> <p>CHOROTYPE. — ALSW.</p> <p>MACROHABITAT. — Larch woods.</p>Published as part of <i>Isaia, Marco, Paschetta, Mauro & Chiarle, Alberto, 2015, Annotated checklist of the spiders (Arachnida, Araneae) of the Site of Community Importance and Special Area of Conservation " Alpi Marittime " (NW Italy), pp. 57-114 in Zoosystema 37 (1)</i> on page 68, DOI: 10.5252/z2015n1a4, <a href="http://zenodo.org/record/4577680">http://zenodo.org/record/4577680</a>
Nemesia pedemontana Decae, Pantini & Isaia, 2015, n. sp.
<i>Nemesia pedemontana</i> n. sp. <p>(Figs 4 E, F, 7)</p> <p> <b>Type material. ITALY: Piemonte</b> Holotype: nr. NHMB.164, 1(m): Province of Torino, loc. Maglione, 45.34°N, 8.00°E, alt. 314m, mixed deciduous forest, leg. Isaia, Paschetta, 15.X. 2011, rep. MCSNB.</p> <p> Paratypes: <b>ITALY: Piemonte:</b> nr. NHMB.165, 1(f), loc. as holotype, rep. MCSNB; nr. NMHB.162, 1(m); nr. NHMB.198, 1(f), loc. Torino, Venaria Reale, Parco Regionale La Mandria, 45.13°N, 7.63°E, alt. 261m, escarpment in oak wood, leg. Isaia, Paschetta, 30.IX.2011; nr. NHMB.204, 1(f), loc. Canale, 44.83°N, 8.00°E, alt. 264m, leg. Chiarle, Franco, 30.X.2011; nr. NHMB.202 1(f), loc. Montafia, 45.02°N, 8.03°E, alt. 314m, leg. Isaia, Paschetta, 28.XI.2011. All deposited at MCSNB.</p> <p> <b>Other material examined. Piemonte:</b> Province of Alessandria, Cartosio, 44.59°N, 8.41°E, alt. 293m, leg. Isaia and Paschetta, 4(f), 10.X.2011, rep. CI; Cassinelle, 44.60°N, 8.56°E, alt. 402m, woodland, leg. Isaia, Paschetta, 1(f) 10.X.2011, rep. MCSNB. Province of Asti, Montafia, 45.02°N, 8.03°E, alt. 314m, leg. Isaia, Paschetta, 2(f), 28.XI.2011, rep. CI. Province of Cuneo, Canale, 44.83°N, 8.00°E, alt. 264m, leg. Chiarle, Franco, 3(f), 30.X.2011, rep. CI; Canale, SCI Oasi di San Nicolao, 44.83°N, 7.99°E, alt. 296m, leg. Chiarle, 2(f), 30.X.2011, rep. CI; Cortemilia, 44.61°N, 8.19°E, alt. 293m, leg. Isaia, Paschetta, 2(f), 10.X.2011, rep. CI; Roccaforte di Mondovi, 44.31°N, 7.75°E, alt. 525m, leg. Isaia, Mammola, Paschetta, 2(f), 19.VII.2012, rep. CI. Province of Torino, loc. Maglione, 45.34°N, 8.00°E, alt. 314m, leg. Isaia, Paschetta, 1(m), 6(f), 15.X.2011, rep. CI; Parco della Collina di Superga, 45.07°N, 7.76°E, alt. 497m, mixed broadleaf woodland, leg. Isaia, Paschetta, Giuliano, 3(f), 2.XI.2011, rep. CI; Parco della Rimembranza, 45.01°N, 7.71°E, alt. 560m, woodland, leg. Isaia, Rizzioli, Pantini, Chiarle, 8(f), 23.VI.2011, rep. CI & MCSNB; loc. Venaria Reale, Parco Regionale La Mandria, 45.10°N, 7.61°E, alt. 300m, woodland, leg. Isaia, Paschetta, 2(m)7(f), 26.X.2011, rep. CI. Province of Vercelli, Trino Vercellese, Parco Naturale Regionale del Bosco delle Sorti della Partecipanza, 45.22°N, 8.25°E, alt. 150m, broad leaved wood, leg. Isaia, Paschetta, 4(m)18(f)3(j), 15.X.2011, rep. CI.</p> <p> <b>Emilia-Romagna:</b> Province of Bologna, Imola, Riserva Naturale Bosco della Frattona, alt. 100m, 44.35°N, 11.66°E, woodland, leg. Fabbri, 1(m), 10.VIII–9.IX.2000, 4(m) 4(j), 9.IX–13.X.2000 (pt), rep. MCSNB. Province of Forli-Cesena, Bagno di Romagna, Parco Nazionale Foreste Casentinesi, Seghettina, alt. 570m, 43.85°N, 11.82°E, leg. Scaravelli & Bertozzi, 1(m), IX.1997 (pt), rep. MCSNB; Meldola, Riserva Naturale Bosco di Scardavilla, alt. 80–120m, 44.14°N, 12.04°E, woodland, leg. Fabbri, 2(f), 15.IV–12.V.1998, 1(m), 20.VIII– 16.IX.1998 (pt), rep. MCSNB; Premilcuore, Parco Nazionale Foreste Casentinesi, alt. 730m, 43.93°N, 11.78°E, meadow, leg. Scaravelli & Bertozzi, 3(m), 3.IX.1997 (pt), rep. MCSNB; Santa Sofia, Parco Nazionale Foreste Casentinesi, 43.89°N, 11.72°E, leg. Bertozzi, 1(f), 5.VIII.1997, 2(m), 26.IX.1997, (pt), rep. MCSNB; S.Paolo in Alpe, alt. 1020m, 43.87°N, 11.79°E, leg. Bertozzi, 1(f), 5.VIII.1997, leg. Scaravelli & Bertozzi, 2(m)1(j), 26.IX.1997 (pt), rep. MCSNB. Province of Rimini, Gemmano, Riserva Naturale Orienta di Onferno, alt. 280– 530m, 43.86°N, 12.54°E, 1(m), 24.VIII–29.IX.2000, leg. Fabbri, 8(m), 29.IX–3XI.2000, leg. Fabbri, 20(m), 21X– 22.XI.2002, leg. Bertozzi and Fabbri, 2(m), 22.XI–24.XII.2002 (pt), rep. MCSNB.</p> <p> <b>Umbria</b>: Province of Perugia, Nocera Umbra, Colle Aprico, alt. 700m, 43.11°N, 12.78°E, leg. Buttarelli, Ghilardi, Pantini, Valle, 4(m)2(j), VI–XII.1991, (pt), rep. MCSNB; San Giustino, sopra Lama, alt. 400m, 43.55°N, 12.71°E, leg. Pantini, Valle, 1(m), IX.1992 – VI.1993 (pt), rep. MCSNB.</p> <p> <b>Marche:</b> Province of Ascoli Piceno, Montemonaco, Isola San Biagio, alt. 990m, 42.90°N, 13.30°E, leg. Rismondo & Fabbri, 13(m), 27.VII–1IX.2004 (pt), rep. MCSNB; Macerata, Fiuminata, alt. 600m, 43.13°N, 12.84°E, leg. Pantini & Valle, 1(m), IX.1992 – VI.1993 (pt), rep. MCSNB.</p> <p> <b>Abruzzo:</b> Province of L’Aquila, Barisciano, Gran Sasso, San Colombo alt. 1100m, 42.33°N, 13.59°E, oak wood, leg. Marotta & Zuppa, 1(m), 5.III.2003 (pt) rep. MCSNB. Province of Teramo, Isola del Gran Sasso d’Italia, alt. 900m, 42.47°N, 13.68°E, woods leg. Marotta, 2(m)1(j), 7.X.2003, leg. Marotta and Metin, 3(m), 26.X.2002 (pt) rep. MCSNB; Monti dela Laga, Valle Casellana, alt. 800m, 42.74°N, 13.53°E, woods, leg. Marotta, 1(f), 11.VII.2003 (pt), rep. NHMR.</p> <p> <b>Molise:</b> Province of Campobasso, Casacalenda, alt. 630–760m, 41.73°N, 14.85°E, leg. Battista, 5(m) 1– 28.IX.2005, 22(m), 28.IX–15.X.2005, 77(m) 15–30.X.2005, 5(m), 30.X–15.XI.2005 (pt), rep. CT and MCSNB.</p> <p> <b>Toscana:</b> Province of Firenze, Marradi, Pont Valle, alt. 500m, 44.08°N, 11.60°E, leg. Usvelli,1(f), 27.IX.2002, rep. NHMR. Province of Siena, Colle di Val d’Elsa, alt. 350m, 43.46°N, 10.99°E, leg. Decae, 1(f), 27.IX.2002, rep. NHMR.</p> <p> <b>San Marino:</b> Castello della Città di San Marino, Mulini fosso di Canepa, alt. 300m, 43.93°N, 12.47°E, woodland, leg. Casali, 4(m), X.2001, leg. Fabbri, 1(f), 28.IV–25.V.2010 (pt), rep. MCSNB.</p> <p> <b>Diagnosis (Fig. 7).</b> Males differ from those of other species in the <i>N. apenninica</i> group by the long, slender lightly sigmoid curved embolus (Fig. 7 A), its dorsally regular curve (Fig. 7 B), its gradually narrowing tip and the sharp origin of the embolus, best seen in prolateral view (Fig. 7 D). Spermathecae composed of two slender, proximally diverging, centrally twisted receptacles. Proximal part widest. Distal part distinctly bipartite with proximal tube and distally lightly inflated dead end (Fig. 7 E).</p> <p> <b>Description</b>. Male holotype. Measurements: BL=11.8, CL=4.3, CW=3.4, Ca=2.5, Pa=5.3, L1=11.7, L2=10.3, L3=9.3, L4=13.7. Similar to <i>N. appenninica</i> except as noted. <i>Carapace</i>: slightly more elongate (CL/CW=1.3, CL/ Ca=1.7), color more pronounced than in <i>N. hastensis</i>, and darker, with strongly contrasting dark and lighter zones, dark parts distinctly mottled, CZ wedge shaped, warm brown, wide and truncated behind eyes and gradually narrowing towards fovea (Fig. 4 F), thoracic part with distinct central grey leaf-pattern with peripheral yellow zones, fovea somewhat irregular shaped, central groove small, clypeus, darker than in previous two species, bristles less developed than in previous two species, absent along carapace edges, pubescence fine silver grey. <i>Eyes</i>: compactly grouped on and around a steep, anteriorly light colored, process (l/w=1.3), lateral eyes larger than median eyes, ALE largest (ALE/PLE=1.2), AER slightly shorter than PER (AER/PER=0.96), light patch between ALE, POP not uniform black. <i>Chelicerae</i>: dorsal dark greenish black with lighter colored glabrous zones, distribution of bristles and structure of rastellum as in previous two species. <i>Maxillae</i>: (l/w=1.8), ventrally grayish brown further as <i>N. hastensis</i>. <i>Labium</i>: (l/w=0.6), darker than maxillae contrasting with other ventral parts, further as in previous two species. <i>Sternum</i>: (l/w=1.3) bright yellow with distinct dark edge, sigilla less visible than in <i>N. hastensis. Legs</i>: color pattern distinct from previous two species with dorsally dark colored femora and more conspicuous glabrous, longitudinal zones on patellae, segments laterally and ventrally much lighter with glabrous longitudinal zones on femora, spine patterns, pubescence, scopulae, claspers and claws as in <i>N. hastensis. Palps</i>: color and bristle pattern as legs, tibia (l/w=1.9) slightly curved upward and less inflated than in <i>N. hastensis</i>. <i>Palporgan</i>: see Fig. 7 A, D. <i>Abdomen</i>: distinctly darker and more intensely colored ventrally gray otherwise similar previous two species. <i>Spinnerets</i>: darker and more intensely colored than in <i>N. hastensis,</i> otherwise similar.</p> <p> Female paratype. Measurements: BL=17.5, CL=5.8, CW=4.3, Ca 3.5, Pa=8.5, L1=12.8, L2=11.5, L3=10.7, L4=17.1. Similar to <i>N. appenninica</i> except as noted. <i>Carapace</i>: shape as in <i>N. hastensis</i> (CL/CW=1.3, CL/Ca=1.6), color generally darker than in females of previous two species (Fig. 4 E) with dark parts more distinctly mottled than in <i>N. hastensis</i>, CZ caudally more sharply tapering than in both previously described species, fovea, bristles and pubescence as in previous two species. <i>Eyes</i>: as in <i>N. hastensis</i> (l/w=1.2), laterals larger than medians, POP connecting all eyes, ALE largest (ALE/PLE=1.3), AER slightly longer than PER (AER/PER=1.02). <i>Maxillae</i>: (l/ w=1, 9). <i>Labium</i>: (l/w=0.5). <i>Sternum</i>: (l/w=1.3). <i>Abdomen</i>: with large dorsal central light colored patch. <i>Spinnerets</i>: similar in color to ventral abdomen. <i>Spermathecae</i> (Fig. 7 E): general three-partite composition and distribution of glandular tissue.</p> <p> <b>Variation in type sample.</b> Males (n=2): BL=11.8–13.2, CL=4.3–4.9, Pa=5.3–6.5, L1=11.7–13.5, L2=10.3– 12.4, L3=9.3–11.1, L4=13.7–16.0. Ratio ranges CL/CW=1.2–1.3, CL/Ca=1.6–1.8, Eye-group l/w=1.3, ALE/ PLE=1.2, AER/PER=1.0.</p> <p>Females (n=4): BL=17.5–23.2, CL=5.8–6.9, Pa=8.5–10.6, L1=12.8–15.0, L2=11.5–13.8, L3=10.7–12.9, L4=17.1–19.5. Ratio ranges CL/CW=1.2–1.3, CL/Ca=1.6, Eye-group l/w=1.2–1.3, ALE/PLE=0.9–1.3, AER/ PER=1.0.</p> <p> <b>Etymology.</b> The species name is derived from the Latin adjective <i>pedemontanum</i> (at the foot of the mountains), from which the region of Piemonte takes its name, and where the type series was collected.</p>Published as part of <i>Decae, Arthur, Pantini, Paolo & Isaia, Marco, 2015, A new species-complex within the trapdoor spider genus Nemesia Audouin 1826 distributed in northern and central Italy, with descriptions of three new species (Araneae, Mygalomorphae, Nemesiidae), pp. 525-540 in Zootaxa 4059 (3)</i> on pages 534-536, DOI: 10.11646/zootaxa.4059.3.5, <a href="http://zenodo.org/record/242430">http://zenodo.org/record/242430</a>
I rapporti tra Cesare Cantù e Graziadio Isaia Ascoli
Si pubblicano e si commentano le lettere inedite tra Cesare Cantù e Graziadio Isaia Ascoli del fondo Cantù presso la Biblioteca Ambrosiana. Il saggio apporta nuovi elementi alla conoscenza dei rapporti scientifici tra i due corrispondenti
Histopona fioni Bolzern, Pantini & Isaia, 2013, sp. n.
Histopona fioni sp. n. Figures 3 –6, 9–11, 15–16, 23–24, 28. H. italica Hänggi 1990: 163, f. 21 a (m misidentified). H. italica Trotta 2005: 160, f. 193 (m misidentified). Type material. Holotype male: SWITZERLAND: Tessin: Bustorgna, Mte. S. Giorgio, 950 m, UTMED 50 32 T 496358 5083796 (lon. 8.9530 ° lat. 45.9077 °), 3, 18/IX– 3 /X/ 1989, Hänggi A. (NMB: 0 2488 a; Hänggi 1992 sub H. italica). Paratypes: SWITZERLAND: Tessin: Bustorgna, Mte. S. Giorgio, 950 m, UTMED 50 32 T 496358 5083796 (lon. 8.9530 ° lat. 45.9077 °), 33, 18/IX– 3 /X/ 1989, Hänggi A. (NMB: 20673; Hänggi 1992 sub H. italica); Paruscera, Mte. S. Giorgio, 1025 m, UTMED 50 32 T 496391 5083968 (lon. 8.9534 ° lat. 45.9092 °), 13, 28/IX/ 1988, Hänggi A. (NMB: 0 2488 c; Hänggi 1992 sub H. italica); Mte. Generoso, Pree, 1030 m, UTMED 50 32 T 500869 5082904 (lon. 9.0112 ° lat. 45.8997 °), 23, 5/IX/ 1989, Hänggi A. (NMB: 0 2488 b; Hänggi 1992 sub H. italica); V. di Scareglia, 13, 12/X/ 2005, Vicentini (NMB: 0 2488 e). ITALY: Trentino-Alto Adige: Trento: Arco, Monte Biaina, Western slope, locality Gorghi, 1200 m, 2 Ƥ 13 /VII/ 1998, Vailati D.; Concei, Val Concei, Gaverdina, 1500 m, 13 4 /X/ 1986, Vailati D.; Condino, Monte Stigolo, 1550 m, 2 Ƥ 12 /XI/ 1997, Vailati D.; Rovereto, Cengio Rosso, 450 m, 1 Ƥ 21 /XI/ 1992, Vailati D.; Storo, Val d’Ampola, 650 m, 1 Ƥ 5 /V/ 1993, Vailati D. Lombardia: Bergamo: Ardesio, Valcanale, locality Braghina, 830 m, 1 Ƥ 14 /IV– 18 /V/ 2010, Zucchelli W.; Averara, Alpe Cul, UTMED 50 32 T 548073 5099121 (lon. 9.6213 ° lat. 46.0439 °), 1990 m, alpine pasture, 23 13 /VIII– 26 /IX/2002, 1Ƥ 23 /V– 20 /VII/2003, 1Ƥ 20 /VII– 23 /VIII/2003, 1Ƥ 19 /X/ 2003 – 5 /VI/ 2004, Lodovici O., Pantini P. (Isaia et al 2007 sub H. italica); Camerata Cornello, Monte Cancervo, 1800 m, UTMED 50 32 T 547801 5084927 (lon. 9.6163 ° lat. 45.9162 °), rocky area, 13 23 /VII– 27 /VIII/2010, 13 27 /VIII– 7 /X/ 2010, Massaro M., Zucchelli W.; Camerata Cornello, Monte Venturosa, 1850 m, UTMED 50 32 T 547816 5085990 (lon. 9.6166 ° lat. 45.9258 °), rocky area, 23 10 /VIII– 9 /IX/2009, 13 23 /VII– 27 /VIII/ 2010, Massaro M., Zucchelli W.; Camerata Cornello, Monte Venturosa, 1950 m, UTMED 50 32 T 547727 5086239 (lon. 9.6155 ° lat. 45.9280 °), pasture, 33 23 /VII– 27 /VIII/2010, 13 27 / VIII– 27 /X/ 2010, Massaro M., Zucchelli W.; Camerata Cornello, Buffalora, 1100 m, UTMED 50 32 T 549373 5084292 (lon. 9.6366 ° lat. 45.9104 °), beech wood 1 Ƥ 15 /VII– 10 /VIII/ 2009, Massaro M., Zucchelli W.; Camerata Cornello, Buffalora, UTMED 50 32 T 549133 5084660 (lon. 9.6335 ° lat. 45.9137 °), 1150 m, bushy area in beech wood, 33 4 /VI– 14 /VII/2009, 13 10 /VIII– 9 /IX/ 2009, Massaro M., Zucchelli W.; Colzate, Baite Sedernello, 1200 m, 1 Ƥ 17 /VII/ 1988, Ravazzi C., Valle M. (Isaia et al 2007 sub H. italica), 13 2 /VIII/ 2001, Ferrario E., Pantini P., Pellizzoli E., Valle M.; Monasterolo del Castello, Val Torrezzo, 600 m, UTMED 50 32 T 573220 5067560 (lon. 9.9415 ° lat. 45.7577 °), wood, 1 Ƥ 6 /VII– 3 /VIII/1995, 13 19 /IX– 26 /X/1995, 1Ƥ 9 /V– 19 /VI/ 1996, Pantini P., Valle M. (Pantini 2000 sub H. italica); Oneta, slopes of Monte Alben, 2 Ƥ 13 /VI/ 1990 Valle M. (Isaia et al 2007 sub H. italica); Parzanica, Valle dei Foppi, wood, 550 m, UTMED 50 32 T 580465 5064643 (lon. 10.0341 ° lat. 45.7306 °), 23 10 /VIII– 19 /IX/1995, 2 Ƥ 9 / 5–19 /VI/ 1996, Pantini P., Valle M. (Pantini 2000 sub H. italica); Premolo, in doline, South of B. ta Camplano, 1850 m, UTMED 50 32 T 563994 5085216 (lon. 9.8252 ° lat. 45.9175 °), 13 22 / VII– 1 /X/ 2003 (Isaia et al 2007 sub H. italica); Premolo, 1850 m, UTMED 50 32 T 563999 5083688 (lon. 9.8250 ° lat. 45.9037 °), rocky area, 13 19 /VI– 22 /VII/2003, 1Ƥ 1 /X/ 2003 – 7 /VII/2004, 13 4 /VIII– 29 /IX/2004, 1Ƥ 21 /VI– 21 / VII/ 2005 (Isaia et al 2007 sub H. italica); Schilpario, road to Passo Campelli, 1750 m, UTMED 50 32 T 596371 5097461 (lon. 10.2451 ° lat. 46.0239 °), moraine 1 Ƥ 6 /VI– 26 /VI/ 2007; Serina, Valpiana, 13 IV–V/1988, 13 1988, Becci B., Pisoni R. (Isaia et al 2007 sub H. italica); Valgoglio, Val Sanguigno, 1000 m, UTMED 50 32 T 569394 5091553 (lon. 9.8957 ° lat. 45.9740 °), beech and fir mixed wood 13, 2Ƥ 11 /VI– 15 /VII/ 2009 (MSNVR), 43 15 / VII– 11 /VIII/2009, 43 11 /VIII– 15 /IX/2009, 23 6 /VII– 7 /VIII/2010, 43 7 /VIII– 15 /IX/ 2010, Massaro M., Zucchelli W.; Vigolo, Ronchi della Bratta, 850 m, UTMED 50 32 T 577754 5065587 (lon. 9.9994 ° lat. 45.7394 °), spruce wood, 1 Ƥ 18 /VII– 10 /VIII/1995, 83 10 /VIII– 19 /IX/1995, 33, 1 Ƥ 19 /IX– 26 /X/1995, 3Ƥ 26 /X/ 1995 – 20 /II/1996, 23, 7 Ƥ 20 /II– 2 /IV/1996, 73, 3 Ƥ 9 /V– 19 /VI/1996, 2Ƥ 19 /VI– 8 /VIII/ 1996, Pantini P., Valle M. (Pantini 2000 sub H. italica); Lecco: Casargo, Val Marcia, 1000 m, UTMED 50 32 T 532680 5098368 (lon. 9.4223 ° lat. 46.0381 °), wood, 83 25 /VI– 11 /IX/ 2008 Massaro M., Zucchelli W.; Casargo, Val Foppone, 1600–1750 m, UTMED 50 32 T 534471 5097454 (lon. 9.4454 ° lat. 46.0297 °), alpine pasture, 13, 1Ƥ 25 /VI– 11 /IX/2008, 13 13 /VIII– 14 /IX/ 2009, Massaro M., Zucchelli W.; Pagnona, road to Alpe Vesina, 1400–1430 m, UTMED 50 32 T 530507 5102093 (lon. 9.3944 ° lat. 46.0717 °), beech wood, 1 Ƥ 26 /III– 1 /V/1999, 23, 1 Ƥ 1 /V– 9 /VI/1999, 3Ƥ 9 /VI– 6 /VII/1999, 13, 2 Ƥ 6 / VII– 11 /VIII/1999, 13 11 /VIII– 8 /IX/ 1999, Pantini P. (Isaia et al 2007 sub H. italica); Vendrogno, Mornico, 970 m, UTMED 50 32 T 526902 5098292 (lon. 9.3476 ° lat. 46.0376 °), chestnut wood, 2 Ƥ 14 /IV– 13 /V/1999, 13 13 /V– 9 / VI/1999, 1Ƥ 9 /VI– 6 /VII/1999, 13, 2 Ƥ 6 /VII– 11 /VIII/ 1999, Pantini P. (Isaia et al 2007 sub H. italica). Other material examined. SWITZERLAND: Tessin: Bustorgna, Mte. S. Giorgio, 950 m, UTMED 50 32 T 496358 5083796 (lon. 8.9530 ° lat. 45.9077 °), 33, 5– 18 /IX/1989, 3– 30 /X/ 1989, Hänggi A. (NMB: 20674-20675; Hänggi 1992 sub H. italica); Forello, Mte. S. Giorgio, 1095 m, UTMED 50 32 T 495925 5084377 (lon. 8.9474 ° lat. 45.9129 °), 13, 05– 18 /IX/ 1989, Hänggi A. (NMB: 20679; Hänggi 1992 sub H. italica); Mte. Generoso, Pree, 1030 m, UTMED 50 32 T 500869 5082904 (lon. 9.0112 ° lat. 45.8997 °), 33, 30/VII– 12 /VIII/1988, 25/VIII– 5 /IX / 1989,18/IX– 7 /X/ 1989, Hänggi A. (NMB: 20676-20678; Hänggi 1992 sub H. italica), ITALY: Lombardia: Bergamo: Entratico, I Moi, 13 (paratype of Histopona italica, misidentification), 5 /IV/ 1957, Bonino. Etymology. The species is dedicated to Fion Bolzern, firstborn of AB. The species epithet is a name in apposition. Diagnosis. Males (Figures 3 –6, 15) can be separated by the absence of a patellar apophysis (present in torpida - group, except H. vignai Brignoli 1980), the distally tube-like elongated radix (absent in myops - and strinatii -group, plate-like and distally bifid in H. italica) and the distally strongly elongated conductor (broadly rounded in H. italica). Females (Figures 23 –24, 28) can be separated from other Histopona species by the glossy median indented posterior epigynal sclerite (much longer and with anterior margin only moderately indented in torpida -group) with strongly diverging margin (parallel in H. italica), the unpaired “bursa copulatrix” (completely paired copulatory ducts in myops - and strinatii -group) with anterior margin v-shaped (straight or convex in H. italica) and the narrow lateral lobes of the copulatory ducts (broad in H. italica). See also Table 1. Description. Measurements and ratios of male (n= 2, holotype male and paratype male from Pagnona): carapace 2.93–3.27 long, 2.20–2.42 wide. Head region 1.17–1.29 wide; PER 0.61–0.78 wide. Chelicerae 1.35–1.44 long, 0.54–0.58 wide. Labium as long as wide or moderately wider than long. Gnathocoxa ratio width to length: 0.510–0.571. Sternum 1.54–1.73 long, 1.27–1.46 wide. Opisthosoma 2.96–3.75 long, 1.85–2.15 wide. Ratio bulb length (laterally from cymbium base to conductor tip) to cymbium length: 0.79–0.80. Leg measurements are given in Table 2. Measurements of females (n= 2, paratypes from Pagnona and Rovereto): carapace 3.03–3.33 long, 1.95–2.24 wide. Head region 1.22–1.33 wide; PER 0.59–0.75 wide. Chelicerae 1.54 long, 0.68–0.69 wide. Labium moderately wider than long. Gnathocoxa ratio width to length: 0.62–0.64. Sternum 1.57–1.69 long, 1.25–1.40 wide. Opisthosoma 3.50–3.73 long, 2.27–2.42 wide. Epigynal plate 0.98–1.04 long, 1.04–1.10 wide; atrium 0.24– 0.26 long, 0.89–0.98 wide. Receptaculum 0.19 wide. Leg measurements are given in Table 2. Eyes: in dorsal view both eye rows straight or slightly recurved; in frontal view AER straight and PER procurved (Figures 9–10). Diameters: PME: 0.105–0.124; PLE: 0.105–0.143; AME: 0.060–0.086; ALE: 0.110– 0.124. Distances: PME–PME equal diameter of PME; PME–AME less than diameter of PME; PME–PLE less than diameter of PME; PME–ALE equal diameter of PME or slightly less; AME–AME 0.5 –1.0 times diameter of AME; AME–ALE about half diameter of AME. Clypeus height (measured under AME) about 2.5–3.5 times diameters of AME; clypeus height (measured under ALE) about 1.5–2 times diameters of ALE. Coloration: carapace with indistinct pattern only or not darkened. Sternum without coloration pattern. Opisthosoma dark grey green; cardiac mark moderately pronounced; posteriorly without pattern. Legs without colour pattern. Additional somatic characters: distal margin of labium concave. Plumose hairs present on carapace, legs and opisthosoma. Three promarginal teeth, the second one from proximal biggest; 5–6 retromarginal teeth, all equal in size (Figure 11). All trochanters notched. Tarsi I, II and IV with 7–8 dorsal trichobothria and 6–7 on tarsus III. No trichobothria on palp tarsi or cymbium. Colulus moderately divided into two separated plates, sometimes only recognizable as two hairy regions. PLS longer than all others with distal segment as long as or slightly longer than basal segment, both pale. PMS as long as ALS. ALS pale. The formulae of leg spination are listed in Table 3. Male palp (Figures 3 –6, 15– 16): RTA with a large dorsal branch, distally pointed, strongly sclerotized and moderately stepped; lateral branch forming moderately sclerotized finger-shaped appendix; ventral branch forming bulge-like moderately ventrodistally protruding stepped appendix. Tegulum broad ring-shaped, distally dividing into a filiform embolus and a tube-like apophysis (radix), proximal with a moderately serrated margin. Embolus originating (free apex) between 10 and 12 o'clock position; distal tip between 3 and 4 o’clock position. Conductor lamella-like, distally strongly elongated, laterally folded along the whole length; longer than alveolus, distally reaching over alveolus margin; terminal end forming moderately sclerotized peak. Connection of conductor and tegulum membranous, band-like. Median apophysis and tegular apophysis absent. Epigynum and vulva (Figures 23 –24, 28): rectangular epigynal plate sclerotized, often with a distinct v-shaped pattern of paler cuticula, posterior with distinct atrium region; atrium anteriorly limited by weakly sclerotized, almost straight margin of the epigynal plate; atrium posteriorly limited by a glossy sclerite (“epigynal valve”), median deeply indented with strongly diverging margins; between anterior margin and posterior sclerite atrium covered by membranous white cuticula. Copulatory openings located at anteriolateral border of atrium. Copulatory duct first unpaired (“bursa copulatrix”), anteriorly v-shaped, then dividing into paired narrow lateral lobes directing into strongly sclerotized convoluted receptacula; fertilization ducts very short. Distribution. Italy and Switzerland. Lombardian Prealps, from Lago Maggiore to Lago di Garda. Ecology. Records of H. fioni sp. n. refer to forest and open habitats such as beech or fir woods and alpine pastures at moderately high elevation, from 800 to 1600 m. The species also occur in rocky areas at an elevation of 1800–2000 m. Adults are found preferably from spring to autumn.Published as part of Bolzern, Angelo, Pantini, Paolo & Isaia, Marco, 2013, Revision of the Histopona italica group (Araneae: Agelenidae), with the description of two new species, pp. 23-41 in Zootaxa 3640 (1) on pages 28-33, DOI: 10.11646/zootaxa.3640.1.2, http://zenodo.org/record/28364
Histopona leonardoi Bolzern, Pantini & Isaia, 2013, sp. n.
Histopona leonardoi sp. n. Figures 7 –8, 12, 17 –20, 25–26, 29 H. italica Brignoli, 1977: 35, f. 14–15, (m misidentified). Type material. Holotype male: ITALY: Piemonte, Cuneo: Acceglio, springs of Maira River, 1680 m, UTMED 50 32 T 335892 4926442 (lon. 6.9366 ° lat. 44.4726 °), sparse larch wood, 3 4 /VI/ 2009, Rosso M. Paratypes: ITALY: Val d'Aosta: Aosta: Ayas, Champoluc, 1700 m, UTMED 50 32 T 401771 5076342 (lon. 7.7352 ° lat. 45.8336 °), sparse larch wood, 13 31 /VIII/2007, 13 15 /VII/ 2009, Franco I. (CI); Gressoney-St. Jean, 2100 m, UTMED 50 32 T 409050 5067251 (lon. 7.8306 ° lat. 45.7528 °), alpine praires 13 7 /IX/ 2007, Negro M. (CI, Negro et al. 2009 sub H. italica); Gressoney-La-Trinité, 1700 m, UTMED 50 32 T 407631 5079494 (lon. 7.8100° lat. 45.8628 °), sparse larch wood, 13, 4Ƥ 30 /VI/ 2006, Negro M. (NMB: 20853, Negro et al. 2009 sub H. italica); Gressoney-La-Trinité, Gabiet, 2458 m, UTMED 50 32 T 410779 5079371 (lon. 7.8506 ° lat. 45.8621 °), alpine praires, 2 Ƥ 20 /VIII/ 2008, Negro M. (CI, Negro et al. 2010 sub H. italica). Piemonte: Moncerchio di Vallanzengo, Val Sessera, UTMED 50 32 T 428560 5058143 (lon. 8.0827 ° lat. 45.6731 °), beech wood, 3 m, 2 Ƥ 2 /V/2009, 63 5 / IX/ 2009, 583, 1 Ƥ 2 /IX/ 2009 Franco I., Negro M.; 2 Ƥ 2 /V/ 2009, Franco I.; Cuneo: Acceglio, springs of Maira River, 1680 m, UTMED 50 32 T 335892 4926442 (lon. 6.9366 ° lat. 44.4726 °), sparse larch wood, 1 Ƥ 4 /VI/ 2009, Rosso M. (CI); Entracque, Natural Park of Alpi Marittime, 1100m, UTMED 50 32 T 376018 4893859 (lon. 7.4487 ° lat. 44.1874 °), beech wood close to Busset stream, 83, 5 Ƥ 29 /VI– 9 /VIII/ 2007, Wolf-Schwenninger, 2 Ƥ 21 /IX/ 2008, Isaia M., Paschetta M. (CI); Terme di Valdieri, Natural Park of Alpi Marittime, Vallone del Valasco, UTMED 50 32 T 358295 4895033 (lon. 7.2267 ° lat. 44.1947 °), alpine pasture with sparse larch wood, 73, 1Ƥ 11 / VII– 27 /VIII/ 2009, Isaia M., Paschetta M.; Natural Park of Alpi Marittime, Pian della Casa, 1473 m, UTMED 50 32 T 361775 4896277 (lon. 7.2699 ° lat. 44.2066 °), alpine pasture, 13 11 /VII/ 2008, Isaia M., Paschetta M. (CI, Paschetta et al., 2012 sub H. italica); Natural Park of Alpi Marittime, Piano del Valasco, UTMED 50 32 T 358300 4895039 (lon. 7.2267 ° lat. 44.1948 °), alpine pasture with sparse larch wood, 83, 2Ƥ 27 /VIII– 23 /IX/ 2009, Isaia M., Paschetta M. (MSNVR); Terme di Valdieri, Natural Park of Alpi Marittime, 1368 m, UTMED 50 32 T 362064 4896332 (lon. 7.2735 ° lat. 44.2071 °), beech wood, 1 Ƥ 29 /VI/ 2009, Isaia M., Paschetta M. (CI); Vernante, Natural Park of Alpi Marittime, Palanfrè, 1370 m, UTMED 50 32 T 379547 4894524 (lon. 7.4926 ° lat. 44.1939 °), beech wood, 1 Ƥ 10 /IX/2008, 2Ƥ 22 /VI/ 2009, Isaia M., Paschetta M., 2 Ƥ 2 /VII/ 2010, Isaia M. (CI); Torino: Ribordone, Santuario Prascundù, 1400 m, 1 Ƥ 28 /IX/ 2004, Giachino P.M.; cave “Tuna del Diau, 1621 Pi/TO”, 1080 m, UTMED 50 32 T 350672 4979013 (lon. 7.1070 ° lat. 44.9488 °), 13 5 /X/ 2002, Lana E. (CI, Isaia et al., 2011 sub H. italica); Vistrorio, 1 Ƥ V– VIII/ 1993, Giachino P.M.; Verbania-Cusio-Ossola: Varzo, cave “Grotta di San Carlo”, 1 Ƥ 4 /VI/ 1978, Casale A. (MSNVR, Brignoli, 1979 sub H. italica). Liguria: Genova: Mezzanego, Ghiaiette, UTMED 50 32 T 537328 4918352 (lon. 9.4688 ° lat. 44.4174 °), beech wood 850 m, 2 Ƥ 31 /X/ 2009 – 25 /V/2010, 13 25 /V– 18 /VIII/ 2010 Lodovici O., Pantini P., Valle M.; Mezzanego, Forest of Monte Zatta c/o ex Colonia Devoto, 1050 m, UTMED 50 32 T 535942 4917017 (lon. 9.4513 ° lat. 44.4055 °), beech wood, 2 Ƥ 31 /X/ 2009 – 25 /V/ 2010, Lodovici O., Pantini P., Valle M., 1 Ƥ 25 /V/2010, 53, 7 Ƥ 25 /V– 18 /VIII/ 2010 Lodovici O., Pantini P.; Propata, north slope of Monte Cremado, 1640 m, 1 Ƥ 5 /VI– 12 /VII/ 1988, Cartasegna F., Pesce D. (CG); Torriglia, Passo del Colletto, 1280 m, 1 Ƥ 21 /V– 1 /VII/ 1999, Pesce D. (CG); Torriglia, SE slope of Monte Duso, 1380 m, 1 Ƥ 21 /V– 1 / VII/ 1999, Cartasegna F. (CG); La Spezia: Varese Ligure, Passo Cento Croci, UTMED 50 32 T 549768 4918763 (lon. 9.6251 ° lat. 44.4204 °), 1000 m, 43, 1Ƥ IV– VIII/ 1991, Cerbino R., Valle M., 23 VI–IX/ 1992, Pantini P., Valle M.; Savona: Sassello, Rio del Nido, 1000 m, beech wood, 43, 18/VII– 10 /X/ 2001; Sassello, Monte Beigua, 1000 m, 1 Ƥ 17 /VII/ 2001. Lombardia: Pavia: Santa Margherita di Staffora, Hotel Colletta, 1380 m, beech wood, 113 31 /VII– 19 /IX/2001, 43, 7 Ƥ 19 /IX/ 2001 – 20 /III/2002, 1Ƥ 26 /IV– 27 /VI/ 2002, Pantini P. Emilia Romagna: Parma: Bedonia, Passo di Montevacà, 800 m, UTMED 50 32 T 548856 4931317 (lon. 9.6149 ° lat. 44.5335 °), 13 IX/ 1991 – V/ 1992, Buttarelli G., Cerbino R., Pantini P., Valle M.; Piacenza: Bobbio, Passo Penice, 1100 m, UTMED 50 32 T 525987 4960430 (lon. 9.3285 ° lat. 44.7967 °), wood, 6 Ƥ 20 /V– 20 /VI/ 2001 (3 Ƥ NMB: 20536), 1 Ƥ 20 /VI– 31 /VII/2001, 13 31 /VII– 19 /IX/2001, 23, 4 Ƥ 19 /IX/ 2001 – 20 /III/2002, 1Ƥ 20 /III– 26 /IV/ 2002 Pantini P.; Bobbio, road to Monte Penice, 1400 m, UTMED 50 32 T 525246 4959349 (lon. 9.3191 ° lat. 44.7870 °), wood, 133, 1Ƥ 31 /VII– 19 /IX/ 2001, road margin 1400 m, 53, 1Ƥ 19 /IX/ 2001 – 20 /III/2002, 1Ƥ 26 /IV– 27 /VI/ 2002 Pantini P. Other material examined. SWITZERLAND: Tessin: Centovalli, Lionza, 930 m, UTMED 50 32 T 470518 5112649 (lon. 8.6181 ° lat. 46.1667 °), 13, 2Ƥ, 6 /VI/1989, 5/VII/1989, 11– 25 /VIII/ 1989, Hänggi A. (NMB: 0 2488 d, 20671-20672; Hänggi 1992, sub H. italica); Val Careccio, 23, 1Ƥ, 29 /IV– 19 /IX/ 1988, Pronini, P. (NMB: 20669- 20670; Pronini 1989 sub H. italica). ITALY: Piemonte: Biella: Oropa, 13 24 /VIII/ 1972, Vigna Taglianti A. (MSNVR, paratype of H. italica, misidentification); Crissolo, Monviso, 1300 m, 1 Ƥ VII/ 1967, Osella G. (MSNVR, paratype of H. italica, misidentification). Etymology. The species is dedicated to Leonardo Pantini, firstborn of PP. The species epithet is a name in apposition. Diagnosis. Males (Figures 7 –8, 17– 20) can be separated by the absence of a patellar apophysis (present in torpida group, except H. vignai Brignoli 1980), the distally spoon-like elongated radix (absent in myops - and strinatii group, plate-like and distally bifid in H. italica, tube-like in H. fioni sp. n.) and the distally broadly rounded conductor (strongly elongated in H. fioni sp. n.). Females (Figures 25 –26, 29) can be separated from other Histopona species by the glossy median indented posterior epigynal sclerite (much longer and with anterior margin only moderately indented in torpida group) with moderately diverging margin (parallel in H. italica, strongly diverging in H. fioni sp. n.), the unpaired “bursa copulatrix” (completely paired copulatory ducts in myops - and strinatii group) with anterior margin concave (straight or convex in H. italica, v-shaped in H. fioni sp. n.) and the narrow lateral lobes of the copulatory ducts (broad in H. italica). See also Table 1. Description. Measurements and ratios of male (n= 2, holotype and paratype from Entracque): carapace 2.25– 2.86 long, 1.54–2.05 wide. Head region 0.80–1.10 wide; PER 0.45–0.62 wide. Chelicerae 1.02–1.34 long, 0.46– 0.56 wide. Labium as long as wide. Gnathocoxa ratio width to length: 0.508–0.543. Sternum 1.23–1.51 long, 1.05– 1.25 wide. Opisthosoma 1.98–2.46 long, 1.00– 1.35 wide. Ratio bulb length (laterally from cymbium base to conductor tip) to cymbium length: 0.670–0.749. Leg measurements are given in Table 2. Measurements of females (n= 2, Paratype females from Acceglio and Entracque): carapace 2.04–2.28 long, 1.38–1.63 wide. Head region 0.81–1.01 wide; PER 0.48–0.54 wide. Chelicerae 0.87–1.04 long, 0.45–0.49 wide. Labium as long as wide. Gnathocoxa ratio width to length: 0.56. Sternum 1.20–1.25 long, 1.00– 1.08 wide. Opisthosoma 2.01–2.69 long, 1.35–1.81 wide. Epigynal plate 0.70–0.72 long, 0.76–0.78 wide; atrium 0.16–0.18 long, 0.67–0.70 wide. Leg measurements are given in Table 2. Eyes: in dorsal view both eye rows straight or slightly recurved; in frontal view AER straight or slightly procurved, PER procurved. Diameters: PME: 0.103–0.128; PLE: 0.096–0.129; AME: 0.059–0.082; ALE; 0.109– 0.130. Distances: PME–PME about half diameter of PME or slightly less; PME–AME about half diameter of PME or slightly less; PME–PLE about half diameter of PME; PME–ALE about half diameter of PME or slightly less; AME–AME about half diameter of AME or slightly less; AME–ALE less than half diameter of AME. Clypeus height (measured under AME) about 3 diameters of AME or slightly more; clypeus height (measured under ALE) about twice diameter of ALE or slightly less. Coloration: Carapace with narrow, continuous dark margin; two longitudinal symmetric darkened rows of triangular dots present on carapace; narrow darkened band median at head region present. Sternum without pattern. Opisthosoma dark grey green; cardiac mark moderately pronounced; posteriorly with indistinct pattern of chevrons. Legs without color pattern. Additional somatic characters: distal margin of labium weakly concave. Plumose hairs present on carapace, legs and opisthosoma. Three promarginal teeth, the second one from proximal biggest; 5–7 retromarginal teeth, all equal in size. All trochanters notched. All tarsi with 6–7 dorsal trichobothria. No trichobothria on palp tarsi or cymbium. Pale colulus, sometimes moderately darkened, divided into two plates. PLS longer than all others with distal segment as long as basal segment, both moderately darkened. PMS as long as ALS. ALS moderately darkened. The formulae of leg spination are listed in Table 3. Male palp (Figures 7 –8, 12, 17–20): RTA with a big dorsal branch, distally pointed, strongly sclerotized and moderately stepped; lateral branch forming moderately sclerotized finger-shaped appendix; ventral branch forming bulge-like moderately ventrodistally protruding stepped appendix, lateral with 2–3 small stepped bands. Tegulum broadly ring-shaped, distally dividing into a filiform embolus and an elongated, distally spoon-like apophysis (radix), terminally often with a transparent portion. Embolus originating (free apex) at 11 o'clock position; distal tip between 3 and 4 o’clock position. Conductor lamella-like, distally broadly rounded and moderately elongated, laterally folded along the whole length; shorter than alveolus, distally not reaching over alveolus margin; terminal end forming moderately sclerotized peak. Connection of conductor and tegulum membranous, band-like. Median apophysis and tegular apophysis absent. Epigynum and vulva (Figures 25 –26, 29): rectangular epigynal plate sclerotized, posterior with distinct atrium; atrium anteriorly limited by strongly sclerotized, m-shaped margin of the epigynal plate with a posteriorly tapered median region; atrium posteriorly limited by a glossy sclerite (“epigynal valve”), median deeply indented with diverging margins; between anterior margin and posterior sclerite atrium covered by membranous white cuticula. Copulatory openings located at anteriolateral border of atrium. Copulatory duct first unpaired (“bursa copulatrix”), then dividing into narrow paired lateral lobes directing into strongly sclerotized convoluted receptacula; fertilization ducts very short. Distribution. Italy and Switzerland (Tessin). All along the Western Alps and the Northern Apennine. Ecology. Records of H. leonardoi sp. n. mostly refer to forest habitats (beech woods at an elevation of 1000– 1500 m). The species also occurs at higher elevation in alpine pastures (maximum elevation reached at 2458 m Gabiet, Aosta Valley). In a few cases H. leonardoi sp. n. occurred in caves. Adults are preferably found from spring to autumn. Anterior limitation of atrium M-shaped margin of the Almost straight margin M-shaped margin of the epigynal plate with a posteriad of the epigynal plate epigynal plate with a tapered median region posteriad tapered median vulva region and Median posterior margins sclerite of glossy Almost parallel Strongly divergent Divergent Epigynum Shape Figures copulatory of 27 anterior – duct 29) (arrows part of in Straight or moderately convex Concave, v-shaped Concave Lateral lobes of copulatory Very broad, distinct Narrow, band-like Narrow, band-like ducts Histopona. italica Brignoli 1977 Paratype male from Apecchio fe pa ti mt ta total Palp 1.15 0.49 0.40 - 1.50 3.54 I 2.28 0.93 2.01 2.14 1.61 8.97 II 2.06 0.89 1.55 1.83 1.27 7.60 III 2.04 0.84 1.58 2.04 1.14 7.64 IV 2.68 1.01 2.26 2.94 1.43 10.32 Paratype female from Apecchio Palp 0.95 0.46 0.65 - 1.11 3.17 I 2.03 0.88 1.66 1.72 1.41 7.70 II 1.86 0.85 1.36 1.51 1.08 6.66 III 1.85 0.75 1.34 1.81 1.08 6.83 IV 2.30 0.88 2.01 2.60 1.40 9.19 H. fioni sp. n. Holotype male and paratype male from Pagnona (n= 2) Palp 1.23–1.40 0.49–0.58 0.43–0.48 - 1.54–1.79 3.69–4.25 I 2.61–2.79 1.00– 1.03 2.36–2.52 2.42–2.61 1.82–1.97 10.21–10.92 II 2.45–2.72 0.97–1.06 1.97–2.12 2.27–2.42 1.64–1.82 9.3–10.14 III 2.42 0.94 1.85 2.45 1.52 9.18 IV 3.06–3.33 0.97 –1.00 2.67–2.85 3.42–3.64 1.85–1.97 11.97–12.79 Paratype females from Pagnona and Rovereto (n= 2) Palp 1.18–1.23 0.49–0.60 0.51–0.85 - 1.05–1.26 3.23–3.94 I 2.64–2.75 1.09–1.15 2.30–2.50 2.24–2.50 1.67–1.85 9.94–10.75 II 2.45–2.60 1.00– 1.06 1.94–2.10 2.15–2.25 1.58–1.60 9.12–9.61 III 2.36–2.42 0.94–0.96 1.82–1.96 2.31–2.36 1.30–1.31 8.73–9.01 IV 2.97–3.08 1.04–1.06 2.64–2.77 3.21–3.35 1.67–1.73 11.53–11.99 H. leonardoi sp. n. Holotype male and paratype male from Entracque (n= 2) Palp 0.89–1.19 0.38–0.47 0.33–0.38 - 1.01–1.63 2.61–3.67 I 2.07–2.33 0.76–0.91 1.81–2.12 1.92–2.15 1.34–1.63 7.90–9.14 II 1.95–2.25 0.78–0.86 1.51–1.77 1.78–1.99 1.26–1.52 7.28–8.39 III 1.86–2.06 0.69–0.93 1.40–1.69 1.91–2.25 1.11–1.24 6.97–8.17 IV 2.42–2.73 0.78–0.92 2.09–2.42 2.71–3.15 1.44–1.58 9.44–10.8 Paratype females from Acceglio and Entracque (n= 2) Palp 0.78–0.84 0.32–0.37 0.50–0.52 - 0.88–0.91 2.48–2.64 I 1.70–1.88 0.70–0.83 1.38–1.61 1.40–1.64 0.92–1.28 6.10–7.24 II 1.52–1.78 0.68–0.73 1.10–1.23 1.32–1.48 0.94–1.06 5.56–6.28 III 1.50–1.72 0.66–0.73 1.08–1.29 1.38–1.68 0.76–0.90 5.38–6.32 IV 1.94–2.25 0.74–0.79 1.62–1.92 2.04–2.45 1.02–1.19 7.36–8.60 According to the identification key provided by Deeleman-Reinhold (1983), Histopona italica forms a singlespecies group within the genus. The two new species described in this work increase the membership of the italica group, which is defined for females by the presence of a glossy, median deeply indented posterior epigynal sclerite and by the unpaired copulatory ducts, and for males by the absence of a patellar apophysis and by the shape of the embolus, originating basal to the protruding radix. During the examination of the material presented here, large differences in the size of the male palp could be observed, even between specimens from the same locality (e.g. 23 from Liguria, La Spezia: Varese Ligure, Passo Cento Croci). Within the examined specimens of H. leonardoi sp. n., two males appeared to be distinctly larger, the palps showing minor morphological differences (Figures 17–18). Likewise, Bolzern et al. (accepted) and Kraus (1955) observed high intraspecific variation in different species of agelenids, like Malthonica picta (Simon 1870) and Tegenaria femoralis Simon 1873. The absence of valid morphological characters (the only significant difference being the size) and the lack of concrete evidence (no different female forms found and syntopy of both male forms - same locality and same time of the year) provide convincing support to consider the larger males as larger forms of the same species. Records of species belonging to the Histopona italica group are known from large parts of Italy (from Calabria to Trentino, along the entire Appenine range, through the Western Alps up to the Lombardian Prealps) (Figure 30). In some cases, specimens of H. italica and H. leonardoi sp. n. were collected at the same locality. Accordingly, the known distribution of H. italica overlaps that of H. leonardoi sp. n. in the district of Maritime Alps and Northern Apennines, the first extending southwards along the Apennines and the latter northwards, along the Alps. It is likely that H. leonardoi sp. n. and H. italica may also occur in French Maritime Alps, but no direct evidence support the presence of this species in France. Records of H. fioni sp. n. are only known from the Lombardian Prealps (Lombardia and southern Trentino in Italy and Tessin in Switzerland). Apparently, no overlap occurs between H. fioni sp. n. and H. leonardoi sp. n., being separated by Lake Maggiore, at the border between Piemonte and Lombardia (Tessin Valley). The same separation is known to occur for Coelotes pickardi tirolensis (Kulczyn'ski 1906) and C. pickardi pickardi O. P.- Cambridge 1873 (Isaia & Pantini 2009) and for Troglohyphantes lucifuga Simon 1884 and T. sciakyi Pesarini 1989 (Isaia & Pantini 2010).Published as part of Bolzern, Angelo, Pantini, Paolo & Isaia, Marco, 2013, Revision of the Histopona italica group (Araneae: Agelenidae), with the description of two new species, pp. 23-41 in Zootaxa 3640 (1) on pages 33-38, DOI: 10.11646/zootaxa.3640.1.2, http://zenodo.org/record/28364
Troglohyphantes konradi Brignoli 1975
<i>Troglohyphantes konradi</i> Brignoli, 1975 <p> LITERATURE. — Brignoli (1975, 1985); Bologna & Vigna Taglianti (1982); Pesarini (2001); Arnò & Lana (2005); Isaia <i>et al.</i> (2007, 2010c, 2011); Isaia & Pantini (2010).</p> <p>CHOROTYPE. — ALSW.</p> <p>MACROHABITAT. — Caves.</p>Published as part of <i>Isaia, Marco, Paschetta, Mauro & Chiarle, Alberto, 2015, Annotated checklist of the spiders (Arachnida, Araneae) of the Site of Community Importance and Special Area of Conservation " Alpi Marittime " (NW Italy), pp. 57-114 in Zoosystema 37 (1)</i> on page 73, DOI: 10.5252/z2015n1a4, <a href="http://zenodo.org/record/4577680">http://zenodo.org/record/4577680</a>
Piniphantes agnellus Milano & Mammola & Rollard & Leccia & Isaia 2019, n. comb.
Piniphantes agnellus (Maurer & Thaler, 1988) n. comb. (Figs 6, 7, 8) Lepthyphantes agnellus Maurer & Thaler, 1988: 338, figs 17-19. MATERIAL. — France. Provence-Alpes-Côte d’Azur, Alpes-de-Haute- Provence, Barcelonnette, pathway to Chapeau de Gendarme, talus caves, 2047 m, 02.VII.2017, Isaia & Mammola leg., 9 ♀, coll. MI. OTHER MATERIAL. — Italy. Piemonte, Province of Cuneo, Valdieri, Galleria di Valscura, alpine scree, 2100 m, 12.VII.2009, Isaia leg., 1 ♂ (sub L. agnellus in Isaia et al. 2015; coll. MI); Piemonte, Province of Cuneo, Valdieri, Lago Soprano della Sella, alpine prairies with rocky debris, 2300 m, 01.VIII.2011, Chiarle leg., 1 ♀ (sub L. agnellus in Isaia et al. 2015; coll. MI); Piemonte, Province of Torino, Cesana, Champlas Janvier, grasslands, 2200 m, VII.2011, Chamberlain leg., 1 ♂, coll. MI; Piemonte, Province of Cuneo, Vinadio, Colle della Lombarda, rocky lands, 2600 m, 06. VI.2016, Isaia leg., 1 ♀, coll. MI; Piemonte, Province of Cuneo, Terme di Valdieri, pathway to Fremamorta, alpine prairies with rocky debris, 2200 m, 11. VI.2016, Isaia leg., 3 ♀, coll. MI; Piemonte, Province of Cuneo, S. Anna di Vinadio, Fortini Laghi Lausfer, alpine prairies with rocky debris, 2300 m, 20.VII.2016, Isaia, Mammola & Milano leg., 13 ♀, coll. MI; Liguria, Province of Imperia, Pigna, Monte Grai, abandoned mine, 1903 m, 12.VIII.2016, Isaia & Beikes leg., 1 ♂, 4 ♀, coll. MI; Piemonte, Province of Cuneo, Ormea, Monte Mongioie, alpine prairies with rocky debris, 1950-2100 m, 03.X.1972, Thaler leg., 5 ♀ (Maurer & Thaler 1988) (not examined). France. Provence-Alpes-Côte d’Azur, Alpes-Maritimes, Belvédère, Pas de l’Arpette, rocky debris, 2400 m, 01.VIII.1986, Maurer leg., 1 ♀ (Maurer & Thaler 1988) (not examined); Provence-Alpes-Côte d’Azur, Alpes-Maritimes, Tende, Vallée des Merveilles, next to Lac Mouton, alpine scree with vegetation, 2190 m, 10.VIII.2004, Hervé & Gargominy leg., 2♀, ARM002, MNHN; Provence-Alpes- Côte d’Azur, Alpes-Maritimes, Belvédère, next to Lac Autier, alpine scree, 2400 m, 12.VIII.2004, Hervé leg., 1♀, ARM016, MNHN; Provence-Alpes-Côte d’Azur, Alpes-Maritimes, Tende, La Minière de Vallauria, waterfall scree, 1508 m, 13. VI.2005, Hervé leg., 2 ♀, ARM061, MNHN; Provence-Alpes-Côte d’Azur, Alpes-de-Haute- Provence, Allos, Sommets des Garrets, western slope, alpine scree, 2670 m, 08.IX.2005, Hervé leg., 2 ♀ 1♂, ARM139, MNHN. CHOROTYPE. — ALSW. MACROHABITAT. — Caves, rocky lands. NOTE. — The collection of this rare species in the frame of this work give us the opportunity to clarify its taxonomic position and to describe the so far unknown male, collected in the frame of previous researches. We hereby provide an exhaustive taxonomical note and information about its current known distribution and provide new diagnostic drawings for male and female (Figs 6, 7, 8). TYPE MATERIAL. — Holotype, female. Leg. Maurer 06.VIII.1986, Museum of Natural History of Genève (not examined). TYPE LOCALITY. — Boulder fields in the vicinity of Lac de l’Agnel (Tende, France), Alpes-Maritimes, 2530 m, in rock field. DESCRIPTION OF THE MALE (Specimen from Valdieri, Galleria Valscura): overall size and leg length small. Prosoma 0.49 long, 0.40 wide, light-yellowish. Thoracic region yellowish with grey shades. Cephalic region not elevated with a few bristles interspersed among the eyes. Clypeus 0.04 long, slightly indented under the eyes, then convex, with one bristle just below the head region. Eyes normally developed, with pigment and black margins. AME smallest. PLE, PME and ALE almost equal in diameter. ALE and PLE contiguous. PLE–PME distance: 0.011, ALE–AME distance: 0.012, PME–PME distance: 0.013. Eye diameters: AME: 0.015, PME: 0.020, ALE: 0.021, PLE: 0.210. Sternum heart-shaped, yellowish with blackish shades. Chelicerae 0.10 long, light brownish, with 18-20 lateral stridulatory ridges and armed with four contiguous posterior teeth grouped close to the base of the fang (the distal bigger) and three anterior teeth, equally distributed along the cheliceral margin, the median bigger. Legs uniformly light yellowish. Leg I: femur 0.56, other articles missing; leg II: femur 0.39, patella 0.09, tibia‰.65, metatarsus‰.42, tarsus‰.36, TLL 1.91; leg III: femur 1.84, patella 0.11, other articles missing; leg IV: femur 0.48, patella 0.12, other articles missing. Abdomen 0.54 long, 0.38 wide; light-brownish, darker than the prosoma. Palp (Fig. 6): femur 0.12, patella 0.04, tibia 0.03, cymbium 0.12. Cymbium faintly convex, roughly rectangular when seen from above, ending proximally with a straight border, perpendicular to the main axis (Fig. 6 B). Paracymbium U-shaped in lateral view, bearing some hairs on the proximal part, apical part gradually narrowed anteriorly (Figs 6 A, 7 D). Distal suprategular apophysis directed upwards, with a sharp end (Fig. 7 C). Proximal part of the embolus with elongated projection bearing numerous finger-like protrusions (Fig. 7 A). Embolus sickle shaped, thumb well-developed. Embolus proper bifid (Fig. 7 A). Lamella characteristica duck-head shaped with an upper sclerotized horizontal branch and a lower one, smaller and less sclerotized (Fig. 7 B). SPINATION (BASED ON ALL MALES EXAMINED) Femur I with one prolateral spine; femur II, III and IV with no spines. Patella I-IV with one dorsal spine. Tibia I with two dorsal, one prolateral, and one retrolateral spines; tibia II with one dorsal, and one prolateral and one retrolateral spines; tibia III and IV with one dorsal and one retrolateral spines. Metatarsus I–IV with one dorsal spine. Position of TmI: 0.19. Trichobothrium on Mt IV absent. ECOLOGY AND DISTRIBUTION Specimens of P. agnellus n. comb. have been collected primarily in talus caves and rocky areas at medium-high altitudes, between 1900 and 2600 m a.s.l. Mammola et al. (2018) consider the species as a troglophile elements. The distribution of the species is centred on the Alpine districts of Maritime Alps and Ligurian Alps. However, the record of one male in Champlas Janvier (Cottian Alps) let envisage a wider distribution, extending north. TAXONOMICAL REMARKS In the original description, Maurer & Thaler (1988) assigned the newly described species to the genus Lepthyphantes Menge, 1866. In lack of males, the diagnosis – and presumably the genus assignation – was based on the morphology of the epigyne, bearing some characteristic lateral extensions at the base of the proscape. The occurrence of males of a possible undescribed species together with females of former Lephtyphantes agnellus at two sites (Galleria Valscura, Valdieri, Maritime Alps and Monte Grai, Triora, Ligurian Alps) allowed to pair males and females. Moreover, the match was confirmed by morphological characters shared by males and females, such as chaetotaxy, cheliceral teeth, stridulatory ridges, abdominal pattern and ocular pattern. Some years after the description of L. agnellus, Saaristo & Tanasevitch (1993, 1996) reclassified the genus Lepthyphantes using a typological approach, examining the morphology of the genital organs, especially males. As a result, most of the European Lepthyphantes species were transferred or assigned to new genera. On the other hand, given the lack of males, L. agnellus was not transferred to any of the newly created genera. Our finding of the unknown male now allows the placement of this species within the genus Piniphantes Saaristo & Tanasevitch, 1996. According to the original description, the genus Piniphantes includes small Linyphiids, having in males an elongated projection at the proximal part of the embolus bearing numerous finger-like protrusion (Saaristo & Tanasevitch 1996). Such character is particularly remarkable in our case (Fig. 7 A). Moreover, other details given in the genus description (Saaristo & Tanasevitch 1996) match our case, such as chaetotaxy and lack of abdominal pattern. The species is then assigned to the genus Piniphantes, with representatives in the area of Tian Shian Mountains (Central Asia) (5 species: P. cinereus (Tanasevitch, 1986), P. macer (Tanasevitch, 1986), P. plumatus (Tanasevitch, 1986), P. uzbekistanicus (Tanasevitch, 1983), P.zonsteini (Tanasevitch, 1989)), Himalaya (one species, P. himalayensis (Tanasevitch, 1987)), Centro-Asiatic-European region (one species, P.pinicola (Simon, 1884)) and Corsica (one species, P. cirratus (Thaler, 1986)). According to the morphology of the male genitalia, P. agnellus n. comb. is similar to P. cirratus, for which the only holotype male is known, preventing any comparison of the female.Published as part of Milano, Filippo, Mammola, Stefano, Rollard, Christine, Leccia, Marie-France & Isaia, Marco, 2019, An inventory of the spider species of Barcelonnette (France), with taxonomic notes on Piniphantes agnellus n. comb. (Araneae, Linyphiidae), pp. 29-58 in Zoosystema 41 (4) on pages 39-42, DOI: 10.5252/zoosystema2019v41a4, http://zenodo.org/record/371848
Cybaeus vignai Brignoli 1977
<i>Cybaeus vignai</i> Brignoli, 1977 <p>LITERATURE. — Isaia & Chiarle (2015).</p> <p>CHOROTYPE. — ALSW.</p> <p>MACROHABITAT. — Broadleaved woods, riparian habitats.</p>Published as part of <i>Isaia, Marco, Paschetta, Mauro & Chiarle, Alberto, 2015, Annotated checklist of the spiders (Arachnida, Araneae) of the Site of Community Importance and Special Area of Conservation " Alpi Marittime " (NW Italy), pp. 57-114 in Zoosystema 37 (1)</i> on page 90, DOI: 10.5252/z2015n1a4, <a href="http://zenodo.org/record/4577680">http://zenodo.org/record/4577680</a>
Dysdera cribrata Simon 1882
<i>Dysdera cribrata</i> Simon, 1882 <p>LITERATURE. — Isaia & Chiarle (2015).</p> <p>CHOROTYPE. — ALSW.</p> <p>MACROHABITAT. — Alpine prairies, riparian habitats.</p>Published as part of <i>Isaia, Marco, Paschetta, Mauro & Chiarle, Alberto, 2015, Annotated checklist of the spiders (Arachnida, Araneae) of the Site of Community Importance and Special Area of Conservation " Alpi Marittime " (NW Italy), pp. 57-114 in Zoosystema 37 (1)</i> on page 62, DOI: 10.5252/z2015n1a4, <a href="http://zenodo.org/record/4577680">http://zenodo.org/record/4577680</a>
Diaea dorsata
<i>Diaea dorsata</i> (Fabricius, 1777) <p>LITERATURE. — Isaia 2005; [Rossi & Bosio 2012].</p> <p>CHOROTYPE. — SIE.</p> <p>MACROHABITAT. — Not specified.</p> <p> <i>Ebrechtella tricuspidata</i> (Fabricius, 1775) (Fig. 15)</p> <p>LITERATURE. — Rossi & Arnò 1995; [Isaia 2005]; [Rossi & Bosio 2012].</p> <p>CHOROTYPE. — PAL.</p> <p>MACROHABITAT. — Shrublands, broadleaved forests.</p>Published as part of <i>Paschetta, Mauro, Christille, Claretta, Marguerettaz, Fabio & Isaia, Marco, 2016, Regional catalogue of the spiders (Arachnida, Araneae) of Aosta Valley (NW Italy), pp. 49-125 in Zoosystema 38 (1)</i> on pages 107-108, DOI: 10.5252/z2016n1a3, <a href="http://zenodo.org/record/4578074">http://zenodo.org/record/4578074</a>
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