359,539 research outputs found

    Spiranthes nivea var. papillata T. C. Hsu & Suetsugu. Consequently 2023, var. nov.

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    Spiranthes nivea var. papillata T.C. Hsu & Suetsugu, var. nov. — Fig. 5. ≡ Spiranthes minutiflora Hsu (2016: 187) in Hsu & Chung (2016), nom. illeg., non Richard & Galeotti (1845: 32). Type:— TAIWAN. Ilan: Tsuifong Lake, 1800–1900 m elev., 3 June 2015, T.C. Hsu 7743 (holotype: TAIF496968!). Diagnosis:— Spiranthes nivea var. papillata is morphologically distinguishable from S. nivea var. nivea by its more densely pubescent rachis and ovaries (vs. sparsely pubescent rachis and ovaries), narrower sepals that are white tinged with pink or purple at the apex (vs. wider and entirely white sepals), and papillose labellum disc (vs. almost glabrous labellum disc). Morphological descriptions and illustrations: —See Hsu & Chung (2016: 187), as Spiranthes minutiflora. Distribution and ecology: —This variety is currently recorded around Tsuifong Lake and Taipingshan in northeastern Taiwan. It grows on sunny grasslands, semi-open roadside slopes, and cliffs around 1800–2100 m elev. Flowers were observed from May to July. Etymology:— The specific epithet is named after its papillate lip and papillate-pubescent rachis and ovaries that are diagnostic from the typical variety. Additional specimens examined:— TAIWAN. Ilan Co.: Taipingshan, 21 May 2012, T.-C. Hsu 5742 (TAIF!); same loc., 23 June 2017, T.-C. Hsu 9299 (TAIF!); Tsuifeng Lake, 22 June 2017, T.-C. Hsu 9287 (TAIF!). Taxonomic remarks: — Spiranthes nivea var. papillata shows morphological resemblance to S. nivea and S. hongkongensis, due to their shared autogamous reproductive biology. Althouth Lin (2019) considered it conspecific with S. hongkongensis, this taxon actually differs in having bracts that significantly exceed ovaries, smaller and glabrous basal labellum callosities, ovate labellum, and smaller column. These features imply a closer relationship to S. nivea, but it still differs from S. nivea var. nivea by several morphological characters mentioned in the diagnosis. Given that there are no significant differences in the labellum and column morphology, which are essential characteristics in species delimitation in S. sinensis species complex (Pace et al. 2019), we concluded that the relatively minor differences are attributed to intraspecific variation, describing it as a new variety of S. nivea.Published as part of Suetsugu, Kenji & Hsu, Tian-Chuan, 2023, Taxonomic revision of the genus Spiranthes (Orchidaceae) in Taiwan, pp. 1-10 in Phytotaxa 578 (1) on pages 6-7, DOI: 10.11646/phytotaxa.578.1.1, http://zenodo.org/record/751762

    [[alternative]]Study on genetic structure of Sibataniozephyrus kuafui Hsu & Lin and phylogenetic relationship among Sibataniozephyrus species

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    [[abstract]]Sibataniozephyrus kuafui was a recently described lycaenid butterfly ( Hsu & Lin,1994 ),which utilites Fagus hayatae as the sole larval host. The first aim of the research is to investigate the population structure of this rare butterfly in Taiwan. The second issue to be explored here is the possibility of presence of coevolutionary pattern as Sibataniozephyrus is the only lycaenid genus that is specialized on the beech trees, with each described species use a different beech species as its larval host. A seguence of 405 bp of COI gene was analyzed to investigate the genetic differentiations of S. kuafui. It turned out the population at Tongshan, Ilan is well differentiated from that of the N. Chatienshan (Fst=0.51),with the former possessing one unique haplotype and the latter two. After studying a sequence of 1068 bp of COI, tRNA and COII gene, it was found that S. kuafui shares a synapomorphy with S. fujisanus by a deletion of an amino acid code AAT in compared with the Sibataniozephyrus taxa in the Asiatic mainland. The phylogenetic pattern derived from the research did not support the coevolution model between Sibataniozephyrus lycaenids and their hosts. Alternatively, the data suggests after gaining the ability of using beech as larval host, Sibataniozephyrus horizontally shifted larval host usage, and speciated by the other evolutionary causes, independent of the diversification of the beeches.

    Spindasis syama subsp. lamuae Hsu & Liang 2020, ssp. nov.

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    Spindasis syama lamuae ssp. nov. (Figs. 9–16, 25–29) Aphnaeus syama var. leechi; Matsumura (not Swinhoe), 1919: Thousand insects of Japan (Additamenta) 3: 609. pl., 48, 4, 5. (mis-identification) Aphnaeus syama formosana; Seitz 1927 (not Matsumura): the Macrolepidoptera of the world: 937, pl. 156, fig. 156h; Hirayama (not Moore) 1933: Butterflies in Colour: pl. 20, Fig. 3; pl. 21, fig. 3. (mis-identification) Aphnaeus syama formosana ab. nakaharai Naritomi 1941: Kontyukai 91: 618, pl. 4, Fig. 4. Diagnosis. Ssp. lamuae is characteristic by distal band of central symmetry system of hindwing undersides broken posteriorly at 1A+2A (Figs. 10, 12, 14, 16), whereas this band is continuous, forming a complete V-shaped band in the other subspecies (Figs. 18, 20). The appearance of ssp. lamuae is most similar to ssp. sepulveda Fruhstorfer, 1912 of continental China (Figs. 17–20, 30–34) in wing patterns, sharing the feature of distal band of central symmetry system of forewing undersides in touch with parafocal elements. Ssp. lamuae demonstrates profound seasonal variation, with the markings on wing undersides of individuals emerged in dry/cooler months reduced and turning reddish (Figs. 12, 16). By contrast, seasonal variation is absent in ssp. sepulveda, with individuals emerged from all seasons possessing black spots and bands (Figs. 18, 20); posterior margin of costa on valva is smooth in lamuae (Fig. 27), whereas it is unevenly serrate in ssp. sepulveda (Fig. 32). Type materials. Holotype. ♂, KAOHSIUNG Co [= KAOHSIUNG CITY]: Meinong, Shuangxi, 150m, 8. II. 2006, Coll. Y. F. Hsu (NHM). Paratypes. JILONG CITY: 1♀, Longgang Trail, 2. IX. 2006, Coll. Y. F. Hsu; 1♀, same locality, 26. IX. 2006 (Y. F. Hsu). TAIPEI Co. [= NEW TAIPEI CITY]: 1♂, Shiding, Ergeshan, 28. V. 2004, reared from Maesa japonica, emgd. 10. VIII. 2004, HSU 04 F53 (J. R. Chen & C. T. Chuang); 2♂, 1♀, Danshui, Miantianshan, 2. VIII. 1987 (Y. F. Hsu). TAOYUAN Co. [= TAOYUAN CITY]: 1♂, 1♀, Fuxing, Gaoyi, VIII. 1984 (C. L. Lee). YILAN Co.: 1♂, Datong, Qilan, 5. VIII. 1988 (C. F. Li). NANTOU Co.: 3♂, Yuchi, Lianhuachi, 22. V. 1989 (C. F. Li); 1♀, Puli, 15. VIII. 1989 (C. F. Li); 1♂, Yuchi, Lianhuachi, 700m, 8. X. 2002 (Y. F. Hsu); 1♂, Renai, Huisun, ca 700m, 14. XI. 2004 (Y. F. Hsu); 1♀, same locality, 22. X. 2005 (Y. F. Hsu); 1♂, 1♀, same locality, 16. X. 2010 (Y. F. Hsu); 1♀, Renai, Nanshanxi, ca 900m, 1. IX. 2007 (Y. F. Hsu); 1♂, 1♀, Renai, Wushe, 26. VII. 2016 (J. Y. Liang) (genitalia preparation JYL446, 447). HUALIAN Co.: 1♀, Xiulin, Lushui, 11. VIII. 1988 (C. F. Li). JIAYI Co.: 1♂, Fanlu, Chukou, ca 350m, 10. X. 2005 (Y. F. Hsu); 1♀, Fanlu, Chukou, 300m, 5. IX. 2010 (Y. F. Hsu); 1♂, 1♀, Alishan, Shizhuo/Dabang, 900/ 1000m, 25. IX. 2010 (Y. F. Hsu). TAINAN Co. [= TAINAN CITY]: 1♂, Guanziling, 400m, 17. IX. 2002 (Y. F. Hsu); 1♂, Baihe, Zhentoushan, 25. X. 2002 (Y. F. Hsu); 2♂, Xinhua, 24. IX. 2010 (Y. F. Hsu); 1♂, 1♀, same locality, 29. III. 2013 (Y. F. Hsu). KAOHSIUNG Co. [= KAOHSIUNG CITY]: 1♂, Liugui, 1. II. 1983 (Y. F. Hsu); 1♂, same locality, 26. III. 1989 (D. X. Lee); 1♂, same locality, 200m, 31. XII. 2006 (Y. F. Hsu); 5♂, Liugui, Nanfengshan, 17. VI. 1989 (Y. F. Hsu); 3♀, Meinong, Shuangxi, 150m, 8. II. 2006 (Y. F. Hsu), 2♂, same locality, 22. I. 2007 (Y. F. Hsu). PINGDONG Co.: 1♀, Wutai, Wutoushan, 9. IV. 1999 (Y. F. Hsu); 1♂, same locality, 1200m, 4. IV. 2002 (Y. F. Hsu, C. C. Lu & C. L. Huang); 1♀, Wutai, 400m, 8. II. 2006 (C. C. Lu); 1♂, 2♀, Wutai, Yichangshan, 1100/ 1400m, 15/ 16. III. 2009, reared from Ardisa crenata with Crematogaster laborisa, emgd. 9. IV/ 9. V. 2009, HSU 09 C26 (Y. F. Hsu & H. C. Huang); 2♂, Sandimen, 250m, 10. II. 2007 (Y. F. Hsu); 1♂, Fangliao, Yuquan, 26. II. 2006 (Y. F. Hsu); 1♂, same locality, 19. III. 2006 (Y. F. Hsu), 1♂, Chunri, Dahanshan, 20. I. 2011, reared from Psidium guajava, emgd. 17. IV. 2011, HSU 11 A14 (J. H. Lin); 2♂, same locality, 27. IV. 2011 (J. H. Lin). Paratypes deposited in NHM, NMNS, and NTNU. Bionomics. Female butterfly oviposits on foliage (twig or leaf) of hostplant (Fig. 43, 44) in the presence of associated ants. Larvae are phytophagous but tended by Crematogaster ants on regular basis (Lin 2011). The larvae devour leaves by scratching epidermis and mesophyll. Larvae conceal themselves gregariously in shelters construct- ed by tying dry leaves while not feeding (Figs. 45). Pupation is taken place within the larval shelters (Figs. 46). Hostplants. Plants oviposited by females or utilized by immatures in the wild included Maesa japonica (Primulaceae) (04F53, oviposition), Ardisia crenata (Primulaceae) (09C26, 09J46, larvae), A. cornudentata morrisonensis (Primulaceae) (09H10, oviposition; 10H20, larvae), A. cornudentata cornudentata (Primulaceae) (09K4, 10H27, 10J52, 11D33, larvae), Mallotus japonica (Euphorbiaceae) (10G26, larva) and Smilax odortissima (Smilaceae)(10J41, larvae). Myrmecophily. This butterfly is obligatorily associated with Crematogester amia (09C26, 09H10, 10G26, 10H20, 10J41) and C. popohana (09K4, 10H27, 10J52, 11D33) in the wild, but larvae may complete development without presence of ants in laboratory (Lin 2011). Etymology. The subspecific name lamuae refers to a comic character Lam (Lamu) created by a famous manga artist Rumiko Takahashi. The patterns of wing undersides recall the graphic design of the bikini Lam wears. Remarks. Seitz (1927) states that S. syama in Taiwan is diagnosable by having the ground color of wing undersides being white, but examined specimens and illustrations of literature all have yellow or creamy yellow ground color on wing undersides.Published as part of Hsu, Yu-Feng & Liang, Jia-Yuan, 2020, On systematic status of Spindasis syama Horsfield, [1829] in Taiwan and the Philippines (Lepidoptera: Lycaenidae: Aphnaeini), pp. 485-500 in Zootaxa 4763 (4) on pages 491-494, DOI: 10.11646/zootaxa.4763.4.2, http://zenodo.org/record/376205

    HSU Homecoming at Playing Field

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    Copy negative of the HSU Cowgirls and Cowboy Band performing on the field with other people and a car at an HSU Homecoming Ceremony in the 1950's

    Hsu, S. C.

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    Hsu, S. C.

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    Entwicklung eines offenen Lernangebots zur Vermittlung digitaler Kompetenzen für Studierende – ein Ergebnis des Projekts DigiTaKS*

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    Das im Rahmen des Projekts DigiTaKS* an der Helmut-Schmidt-Universität/Universität der Bundeswehr Hamburg (HSU/UniBw H) entwickelte adaptierbare Lehr-/Lernpaket ComDigi S* setzt einen innovativen technischen Ansatz zur Förderung digitaler Kompetenzen um. Es integriert fortschrittliche Funktionen einer modernen Learning Experience Plattform (LXP) mit dem etablierten und portablen SCORM-Standard, was eine hochgradig flexible und auf individuelle Lernbedürfnisse zugeschnittene Bildungserfahrung schafft, welche gleichzeitig maximale Kompatibilität bietet. Das hier skizzierte Teilprojekt (AP3) beleuchtet insbesondere den an DigComp 2.2 angelehnten und speziell auf die Zielgruppe der Studierenden zugeschnittenen Kompetenzrahmen und strukturiert Lernpakete nach den Prinzipien der Open Educational Resources (OER), um einen freien und umfassenden Wissensaustausch zu fördern. Weiterhin präsentiert dieser Beitrag konkrete Anwendungsfälle und Feedback von Nutzenden, um die Effektivität und Anwendbarkeit des ComDigi S*-Ansatzes zu bewerten.Vo

    Spiranthes hongkongensis S. Y. Hu & Baretto 1976

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    Spiranthes hongkongensis S.Y. Hu & Baretto (1976: 2) — Fig. 3. Type:— HONG KONG: Tai Po, the garden of Gloria Barretto, 4 April 1975, S. Y. Hu 13658 (holotype: K000942682!; isotypes:A00104456, image!, CUHK). Synonym:— Spiranthes suishaensis auct. non (Hayata 1916: 86) Schlechter (1919: 161): Lin (2019: 284). Morphological descriptions and illustrations: —See Hu & Baretto (1976: 2; f. 2), Hsu & Chung (2014: 406; f.1; 2016: 186), Surveswaran et al. (2017: 125; f. 4), and Lin (2019: 265; f. 116; pl. 13), as Spiranthes suishaensis. Distribution and ecology: — Spiranthes hongkongensis is currently known in Hong Kong, Taiwan, China (Guangdong and Hainan Provinces), Japan, and Borneo. In Taiwan, this species is occasionally found on lowland hills around the Taipei Basin where it grows on very humid sunny grassland accompanied with some wetland plants such as Eriocaulon spp. and Utricularia spp. It was also found growing as a weed in some greenhouses in southern Taiwan, and these populations are presumed as unintentionally introduced along with horticultural plants. Flowers were observed from April to May. Additional specimens examined: — TAIWAN. New Taipei City: Chepingliao, 27 May 2009, T.-C. Hsu 2214 (TAIF!); same loc., 20 April 2014, T.-C. Hsu 7079 (TAIF!); same loc., 8 May 2015, T.-C. Hsu 7674 (TAIF!); same loc., 22 April 2017, J.-Z. Lin 1 (TAIF!). Pingtung Co.: Kaoshu, 26 April 2018, T.-C. Hsu 10536 (TAIF!). CHINA. Hainan Prov.: Mt. Diaoluo National Forest Park, 26 February 2012, T.-C. Hsu 5440 (TAIF!). HONG KONG: 20 April 1975, S. Y. Hu 13673 (KYO!). JAPAN. Okinawa Pref.: Ishigaki Island, Ishigaki City, in campis humidis litoris ad Nagura, 4 April 1937, Takamine s.n. (KPM-NA0304131!). Taxonomic remarks: — Spiranthes hongkongensis can be distinguished from the allogamous taxa such as S. australis and S. sinensis by its modified rostellum and stigma morphology associated with its autogamous breeding system. Spiranthes hongkongensis is most similar to S. nivea, another autogamous species with white flowers. However, it differs from S. nivea, in having papillate (vs. nearly glabrous) labellum disc, larger papillate (vs. smaller glabrous) basal labellum callosities, and densely glandular pubescent (vs. sparsely pubescent) rachis, ovaries, and sepals.Published as part of Suetsugu, Kenji & Hsu, Tian-Chuan, 2023, Taxonomic revision of the genus Spiranthes (Orchidaceae) in Taiwan, pp. 1-10 in Phytotaxa 578 (1) on page 5, DOI: 10.11646/phytotaxa.578.1.1, http://zenodo.org/record/751762

    Wroughtonia pterolophiae Chou et Hsu 1998

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    Wroughtonia pterolophiae Chou et Hsu, 1998 Wroughtonia pterolophiae Chou et Hsu, 1998: 299 -300. Comparative diagnosis. Wroughtonia pterolophiae Chou et Hsu is similar to W. spinator (Lepeletier) and differs by having the second tergite punctate (smooth in W. spinator); apical half of hind femur and hind tibia dark brown (mainly yellowish brown) and the dorsal carinae of first tergite weakly developed and remaining far removed from each other (fig. 77 in Chou et Hsu, 1998; strongly developed, lamelliform basally and converging to each other (fig. 44 in van Achterberg, 1987)). Host. Parasitoid of Pterolophia annulata (Chevrolat) (Coleoptera: Cerambycidae). Distribution. China (Taiwan).Published as part of Yan, Cheng-Jin, Achterberg, Cornelis Van, He, Jun-Hua & Chen, Xue-Xin, 2017, Review of the tribe Helconini Foerster s. s. from China, with the description of 18 new species, pp. 401-457 in Zootaxa 4291 (3) on page 442, DOI: 10.11646/zootaxa.4291.3.1, http://zenodo.org/record/82752
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