370 research outputs found
Cunaxoidinae Den Heyer 1978
Subfamily CUNAXOIDINAE Den Heyer, 1978 Type tribus: Cunaxoidini Den Heyer, 1980 bPublished as part of Heyer, Jacob den, Ueckermann, Edward A. & Khanjani, Mohammad, 2013, Iranian Cunaxidae (Acari: Prostigmata: Bdelloidea). Part III. Subfamily Cunaxoidinae, pp. 2049-2070 in Journal of Natural History (J. Nat. Hist.) (J. Nat. Hist.) 47 (31 - 32) on page 2050, DOI: 10.1080/00222933.2012.763060, http://zenodo.org/record/463191
Lupaeus Castro and Den Heyer 2009
Genus Lupaeus Castro and Den Heyer, 2009 Eupalus Ewing, 1917: 151; Berlese, 1916: 293 Pulaeus Den Heyer, 1981 b: 19; 1981: 87; El-Bishlawy and Rakha, 1983: 373. Sepasgosarian, 1984: 143; Liang, 1985: 79; Bu and Li, 1987: 21; Smiley, 1992: 300; Corpuz-Raros, 1996b: 123–131. Type species: Pulaeus martini Den Heyer, 1981Published as part of Heyer, Jacob den, Ueckermann, Edward A. & Khanjani, Mohammad, 2013, Iranian Cunaxidae (Acari: Prostigmata: Bdelloidea). Part III. Subfamily Cunaxoidinae, pp. 2049-2070 in Journal of Natural History (J. Nat. Hist.) (J. Nat. Hist.) 47 (31 - 32) on page 2061, DOI: 10.1080/00222933.2012.763060, http://zenodo.org/record/463191
Pulaeus Den Heyer
Key to the Brazilian and South African species of Pulaeus Den Heyer 1. Females with propodosomal coxal plates fused medially; median platelet with a seta anteriad to genital opening................................................................................................................................................. P. pectinatus (Ewing, 1909) - Females with propodosomal coxal plates not fused medially; without median platelet anteriad to genital opening... 2 2. Genu I with 4 solenidia plus duplex setae........................................................................ P. quadrisolenidius sp. nov. - Genu I with 2 or 3 solenidia and with or without duplex setae.................................................................................... 3 3. Genu I with 2 solenidia and with duplex setae.......................................................... P. glebulentus Den Heyer, 1980 a - Genu I with 3 solenidia and without duplex setae........................................................................................................ 4 4. Dorsal shield strongly punctated.................................................................................................. P. myrtaceus sp. nov. - Dorsal shield slightly punctated................................................................................... P. franciscae Den Heyer, 1981 bPublished as part of Castro, Tatiane Marie Martins Gomes De & Heyer, Jacob Den, 2009, A revision of the genus Pulaeus Den Heyer, with descriptions of a new genus and four new Brazilian species (Acari: Prostigmata: Cunaxidae), pp. 20-36 in Zootaxa 2141 on page 22, DOI: 10.5281/zenodo.18856
Pulaeus Den Heyer
Key to the Brazilian and South African species of Pulaeus Den Heyer 1. Females with propodosomal coxal plates fused medially; median platelet with a seta anteriad to genital opening................................................................................................................................................. P. pectinatus (Ewing, 1909) - Females with propodosomal coxal plates not fused medially; without median platelet anteriad to genital opening... 2 2. Genu I with 4 solenidia plus duplex setae........................................................................ P. quadrisolenidius sp. nov. - Genu I with 2 or 3 solenidia and with or without duplex setae.................................................................................... 3 3. Genu I with 2 solenidia and with duplex setae.......................................................... P. glebulentus Den Heyer, 1980 a - Genu I with 3 solenidia and without duplex setae........................................................................................................ 4 4. Dorsal shield strongly punctated.................................................................................................. P. myrtaceus sp. nov. - Dorsal shield slightly punctated................................................................................... P. franciscae Den Heyer, 1981 bPublished as part of Castro, Tatiane Marie Martins Gomes De & Heyer, Jacob Den, 2009, A revision of the genus Pulaeus Den Heyer, with descriptions of a new genus and four new Brazilian species (Acari: Prostigmata: Cunaxidae), pp. 20-36 in Zootaxa 2141 on page 22, DOI: 10.5281/zenodo.18856
Neoscirula Den Heyer 1977
Genus Neoscirula Den Heyer, 1977 Type species: Neoscirula theroni Den Heyer, 1977 The genus Neoscirula was erected by Den Heyer (1977) and was consequently included in the subfamily Coleoscirinae (Den Heyer 1980 a). It was transferred by Smiley (1992) to the subfamily Bonzinae Den Heyer, 1978 (Nota bene: The original spelling of the latter subfamily name is with a single i) mainly on the grounds of these mites having, according to his interpretation, a pair of geniculate hg 1 setae, a feature not mentioned in the original description of this genus. Den Heyer (1980 b) referred to this pair of setae as being “semi-geniculate” in Neoscirula sevidi. This could have led to the statement by Smiley (1992) that Neoscirula had geniculate setae hg 1, which is actually not the case for any known species of this genus. The authors of this paper are of opinion that the genus should stay in the Coleoscirinae for reasons subsequently presented. Smiley’s (1992) first step to divide the family Cunaxidae is based on the presence of a pair of geniculate setae hg 1, regarding bent setae (common in most Neoscirula spp.) also as such. In order to be precise, the meaning of the term geniculate has been determined from various sources. The specific explanation is: “...bent at a sharp or abrupt angle….”. In the Cunaxidae geniculate structures occur only in two genera, i.e. in all Bonzia spp. and in an unpublished species of Cunaxa. Den Heyer (1978) redescribed B. halacaroides Oudemans and B. sphagnicola Willmann, illustrating what he regarded as geniculate hg 1 setae. When investigating the Cunaxa spp. Den Heyer (1979 b & c) illustrated a peculiar pair of vi sensillae (fig. 1) which he also regards as geniculate. The first author of this paper is of the opinion that a new genus or subfamily should not be erected based on one species which differs from the other 20–25 species only by having geniculate sensillae vi. This means that geniculate setal structures should not be used as a first criterion for dividing a family (see Smiley 1992; p. 37). When this concept is strictly applied, only the genus Bonzia has a pair of true geniculate hg 1 setae (fig. 2). The hg 1 -setae of the genera Parabonzia and Neoscirula should be regarded as non-geniculate but bent in varying degree. Den Heyer & Castro (2008, in press) also made a change in the notation of the ventral idiosomal chaetotaxy of the Cunaxidae. The original three species included in Neoscirula were described by Den Heyer (1977) from South Africa; he added a fourth species in 1980. Shiba (1978) described one species from Malaysia while Smiley (1992) added three more species from North-America and New Zealand. Corpuz-Raros (1996) reported on seven species from the Philippines. Four species were described from China, two by Lin & Zhang (1998) and two by Lin & Zhang (2002). Mejia-Recamier & Palacios-Vargas (2007) described three species from Mexico. The present paper describes three new Brazilian species.Published as part of Heyer, Jacob Den & Castro, Tatiane Marie Martins Gomes De, 2008, Subfamilial affiliation of Neoscirula (Acari: Prostigmata: Cunaxidae) and descriptions of three new species of this genus from Brazil, pp. 51-62 in Zootaxa 1731 on pages 52-53, DOI: 10.5281/zenodo.18131
Funaxopsis Heyer & Castro, 2009, gen. nov.
Funaxopsis gen. nov. Type species: Cunaxoides passerinae Den Heyer, 1981b. Diagnosis. This genus can be distinguished from other genera by the following set of characteristics: tibia IV with 2 sts and 1 long sts; setae c2 and h2 smooth and depression for pe organ on tarsus I lacking. Etymology. The senior author just added a few more letters to the stem-“..unaxo..” and ending it on a taxonomically recognized suffixPublished as part of Heyer, Jacob Den & Castro, Tatiane M. M. G., 2009, SiGe-Space-Synthesizer-Entwicklung eines SiGe-Synthesizerdemonstrators: SISSI; Schlussbericht, pp. 1-15 in Zootaxa 2140 on page 13, DOI: 10.2314/gbv:620362286, http://zenodo.org/record/531966
Pulaeus sensu Castro and Den Heyer 1979
Genus Pulaeus Den Heyer, 1979 Eupalus Ewing, 1909: 402; Berlese, 1916: 293; Thor and Willmann, 1941: 183. Cunaxoides Baker and Hoffmann, 1948: 245–246; Shiba, 1978: 123–126. Pulaeus Den Heyer, 1979: 18–33; El-Bishlawi and Raka, 1983: 373–375; Bu and Li, 1991: 70–72; Barilo, 1991: 132–134. Neocunaxoides Chaudhri et al., 1979: 176–177; Corpuz-Raros, 1996a: 133–135; Corpuz-Raros, 1996b: 125–129, 131–135; Chaudhri, 1977: 50. Type species Eupalus pectinatus Ewing, 1909. The genus Pulaeus was erected by Den Heyer (1979). It has since been revised and divided by Castro and Den Heyer (2009); thereby erecting the genus Lupaeus. The latter authors provided information on species belonging to these genera; also an historical review and a literature review are provided in their papers. Chaudhri (1977) was the first to describe a species with considerable longer chelicerae, palps and hypostome as usually occurring in this genus, namely Neocunaxoides krama (= Pulaeus sensu Castro and Den Heyer). A further two descriptions followed: Pulaeus longignathos Bu and Li, 1987 and Pulaeus payatapalpus Corpuz-Raros,1996. Sergeyenko (2011) synonymizes both Pulaeus longignathos Bu and Li,1987 and Pulaeus chongqingensis Bu and Li, 1987 as they are, respectively, male and female of Pulaeus krama (Chaudhri, 1977); Sergeyenko (2011) comprehensively redescribed this species from Crimean material. The Iranian male has the dimensions and leg chaetotaxy as provided below.Published as part of Heyer, Jacob den, Ueckermann, Edward A. & Khanjani, Mohammad, 2013, Iranian Cunaxidae (Acari: Prostigmata: Bdelloidea). Part III. Subfamily Cunaxoidinae, pp. 2049-2070 in Journal of Natural History (J. Nat. Hist.) (J. Nat. Hist.) 47 (31 - 32) on page 2056, DOI: 10.1080/00222933.2012.763060, http://zenodo.org/record/463191
Pulaeus Den Heyer 1980
Genus Pulaeus Den Heyer, 1980 a Eupalus — Ewing, 1909: 402; Berlese, 1916: 293; Thor & Willmann, 1941: 183 Cunaxoides— Baker & Hoffmann, 1948: 245–246 Pulaeus— Den Heyer, 1980 a: 19; El-Bishlawy & Rakha, 1983: 373–375; Bu & Li, 1991: 70–72; Barilo, 1991: 132–134; Corpus-Raros, 1996: 119–138; Lin et al, 2001: 145–153 Neocunaxoides— Chaudhri, 1977: 50; Chaudhri et al, 1979: 176–177; Tseng, 1980: 267–272 Type species: Eupalus pectinatus Ewing, 1909. Historical-taxonomic review. Species belonging to this genus are (re-) described and/or figured by the following: Ewing (1909) describes a new species from Illinois, USA, viz. Eupalus pectinatus. Berlese (1916) describes E. sternalis as a new species from Palermo (Panormitano), Italy but also states that this form differs from E. pectinatus, which he received from Ewing, only in palp length. The co-author in 1975 studied specimens of these species in the Berlese Collection in Florence and concluded that they are conspecific. Berlese describes in the same paper a new species, viz. E. subterraneus, from a nest of field mice at Terrara, Italy. Thor & Willmann (1941) provide redescriptions of all above species. In their work on the Cunaxidae Baker & Hoffmann (1948) provide redescriptions of all above-mentioned species. Muma (1960) gives a short description and drawings of a new species from Florida, USA, viz. C. pectinellus. We regard this species to be conspecific with P. pectinatus mentioned in this paper. Chaudhri et al (1979) redescribed Neocunaxoides krama Chaudhri, 1977; this species very strongly resembles P. longignathus Bu & Li, 1987 and is regarded by us as a new combination. We are also of opinion that the following species remain in this genus: Cunaxoides americanus Baker & Hoffmann, 1948; P. ardeola, Barilo, 1991; P. cebuensis Corpuz-Raros, 2007; P. chongqingensis Bu & Li, 1987; P. longignathus Bu & Li, 1987; P. m u s c i Liang, 1985; P. rimandoi Corpuz- Raros, 1996; P. s a m a re n s i s Corpuz-Raros, 2007 and P. w h a r t o n i (Baker & Hoffmann, 1948). Redescription. Mites of this genus are rather well sclerotized and characterised by the presence of fairly well-demarcated idiosomal shields, of a depression for peg organ on tarsus I proximally positioned; genital setae (g 1 —g 4) in approximately straigth row; palp tibiotarsus with a single pointed process. In his redescription of one of the species, viz. Pulaeus glebulentus Den Heyer, Smiley (1992) mentioned the following numbers of setae on basifemora I—IV: 4 sts— 5 sts— 4 sts— 2 sts; he informed that the redescription presented by him corresponds to the original description of the species. Although, the numbers of the setae he mentioned for the basifemora I—IV do not correspond to those presented in the original description of that species (4 sts— 6 sts— 3 sts— 2 sts) being characteristic of the genus, the taxonomical position of P. glebulentus remains unchanged. Etymology: The co-author, when he erected the genus Pulaeus Den Heyer, 1980 a, did not explain the name of this genus. The name is an anagram of the name Eupalus, as given by Koch (1838), a name restricted to the subfamily Cunaxoidinae, as is the genus name Lupaeus, as applied in the paper.Published as part of Castro, Tatiane Marie Martins Gomes De & Heyer, Jacob Den, 2009, A revision of the genus Pulaeus Den Heyer, with descriptions of a new genus and four new Brazilian species (Acari: Prostigmata: Cunaxidae), pp. 20-36 in Zootaxa 2141 on pages 21-22, DOI: 10.5281/zenodo.18856
Cunaxoidinae Den Heyer 1978
Key to the subfamily Cunaxoidinae Den Heyer, 1978 1 Five setae on palp segment II; tarsi III and IV with long setae ending in terminal bulb- like knob (very small); seta formula for telofemora 4– 4 or 3– or 2 or 1–3 or 2 or 1; usually, but not always, six setae on palp tibiotarsus; trichobothrium absent or present........................................................................................ Cunaxoidini Den Heyer, 1980....2 - Six setae on palp segment II; tarsi III (except tsl) and IV with only simple setae; seta formula for telofemora 5–5–4–3 sts; usually, but not always, five setae on palptibiotarsi; trichobothrium always present......Pulaeini Den Heyer, 1980................................................................................................................................................................. 14 2 Femora I and II not divided; setae f2 present; trichobothrium absent....................... Denheyernaxoides Smiley, 1992 - All leg femora divided; setae f2 absent; trichobothrium present or absent.................................................................. 3 3 Dorsum with vague and very poorly demarcated dorsal regions; palp tibiotarsus with subterminal pointed process and small teeth (=serrated edge) on ventral ridge; cheliceral setae absent................................................................... 4 - Dorsum with well demarcated and sclerotised dorsal plates; palp tibiotarsus without serrated edge; pointed process placed more proximally; cheliceral setae present; dorsal setae usually stout and setose....................................................................................................................................................................................... Scutopalus Den Heyer, 1979b 4 Trichobothrium on tibiae IV lacking; depression for pe organ on tarsus I either present or lacking........................... 5 - Trichobothrium on tibiae IV present; depression for pe organ on distal portion of tarsus I..................................................................................................................................................................... Cunaxoides Baker & Hoffmann, 1948 5 Tibiae III with 1 bsl and 3, 4 or 5 sts.; tibiae IV with 2 or 4 sts................................................................................... 6 - Tibiae III with 1 lts, 4 sts; tibiae IV with 4sts, 1 lsts............................................................... Dunaxeus gen. nov......8 6 Tibiae III with 1 bsl and 3, 4 or 5sts; tibiae IV with 2 sts, 1 lts........................................... Funaxopsis gen. nov......12 - Tibiae III with 1 bsl and 5sts......................................................................................................................................... 7 7 Tibiae IV with 4sts, 1 lsts; telofemur IV with 3 sts..................................... Qunaxella triasetosa gen. nov et sp. nov. - Tibiae IV with only 4 sts.; lacks depression and pe organ.................................................... Bunaxella gen. nov......10 8 Telofemur III with 4-4-3-1 sts; basifemur with 1 sts...................................................................................................9 - Telofemur III with 4-4-3-2 sts; basifemur IV with 2 sts............................................................ D. duosetosus sp. nov. 9 Telofemur IV with 2 sts........................................................................... D. capensis (Den Heyer, 1981b) comb.nov. - Telofemur IV with 3 sts; basifemora IV with 1 sts................................... D. elongatus (Den Heyer, 1981b comb.nov 10 Basifemora I—IV with 3—3—2—0 sts; telofemora I—IV with 4— 4—3—1 sts; dorsal setae fan-shaped, except for smooth f2................................................................................................... B. quini (Den Heyer, 1981b) comb.nov - Basifemora I—IV with 4—4—3—1 sts; telofemora I—IV with 4—4—3—2 sts; dorsal setae plumose; setae h2 may be smooth................................................................................................................................................................... 10 11 Setae h2 plumose.................................................................................... B. oribensis (Den Heyer, 1981b) comb. nov. - Setae h2 smooth............................................................................... B. zebedielensis (Den Heyer, 1981b) comb. nov. 12 Basifemora I—IV with 3—3—3—1 sts; setae sci smooth.............................. F.visci (Den Heyer, 1981b) comb.nov. - Basifemora I—IV with 2—2—2—0; setae sci finely setose..................................................................................... 13 13 Telofemora I—IV with 4—3—1—1 sts; seta h1 smooth..................... F. passerinae (Den Heyer, 1981b) comb. nov. - Telofemora I—IV with 4—4—3—1 sts; seta h1 finely setose.............. F. vaneedeni (Den Heyer, 1981b) comb. nov. 14 Setae f2 present.......................................................................................................................................................... 15 - Setae f2 absent; setal formula for basifemora I—IV 3—5—2—0 sts; males with 2 proximal solenidia on tarsus II............................................................................................................................................ Neocunaxoides Smiley, 1975 15 Setal formula for basifemora I–IV 4—6—3—2 sts; males with two proximal solenidia on tarsus II; depression for pe organ on proximal half of tarsus of tarsus I; tibiae II and III with non-striated blunt solenidion; palptibiotarsus with only a single ventral pointed process................................................................................... Pulaeus Den Heyer, 1979a - Setal formula for basifemora I—IV 4—6—3—1 sts; males with one or two proximal solenidia on tarsus II; Depression for pe organ distally (subapically) on tarsus I; tibiae II and III with transversely striated blunt solenidia; palp tibiotarsus with 2 pointed processes, one ventral and one median........................ Lupaeus Castro & Den Heyer, 2009Published as part of Heyer, Jacob Den & Castro, Tatiane M. M. G., 2009, SiGe-Space-Synthesizer-Entwicklung eines SiGe-Synthesizerdemonstrators: SISSI; Schlussbericht, pp. 1-15 in Zootaxa 2140 on pages 3-4, DOI: 10.2314/gbv:620362286, http://zenodo.org/record/531966
Riscus Den Heyer 2006
Genus RISCUS Den Heyer, 2006 Riscus Den Heyer, 2006: 195 –201 Type species Riscus thailandensis Den Heyer, 2006 Genus description. With the acquisition of material from South America the genus characteristics, provided by Den Heyer (2006), have to be revised. They are: very poorly sclerotised, soft-bodied mites; no plates. Setae sci, being the longest idiosomal st-seta and almost reaching the nearest sensillum vi, occur anteriad to a line between sensillae sce. Sensillae are finely and densely pilose. Setae f 2 absent. Lyrifissures im laterad to imaginary line e 1 - f 1. Setae hg 4 occur almost in line with hg 3 forming a transverse row. Hysterogastral setae 3 or 4 pairs. Genital papillae 2 or 3 pairs and small. Genital valves with 4 or 5 pairs of g-setae in female, 4 in tritonymph and 3 in the deutonymph. One pair of paragenital setae flank the genital valves. Paracoxal setae (pcs) on edge of coxae IV or on integument. No para-anal setae. Lyrifissures ih close to anus. The hypognathum possesses a pair of peg-like structures (proprioceptors) laterally to the base of the chelicerae. Palpal genu with 3 or 4 setal structures which may be straight or curved. Palptibiotarsus with a tubercle of varying stoutness. Solenidion formula (all types, i.e. asl, bsl, dtsl, tsl) for genua I-IV: 5 or 4 – 2 – 1 – 1; tibiae I-IV: 2 – 1 – 1 - 0; tarsi I-IV: 7 – 3 – 1 - 0. Femora I and II not divided, III and IV clearly divided. Tarsal lobes very small. Sts- formula for basifemora 3 – 3 – 2 – 1 sts, telofemora 4 – 4 – 4 – 4 sts. All legs are shorter than the body with legs IV being the longer. The known representatives of this genus inhabit an array of plants. Riscus thailandensis occur on Mangifera indica L.; R. austroamericanus sp. nov. was found on Coffea arabica L., Solanum nigrum L., Euterpe sp. and Pisidium guajava. No information exists on their colour and boilogy.Published as part of Heyer, Jacob Den & Castro, Tatiane Marie Martins Gomes De, 2012, New Neotropical cunaxine species (Acari: Prostigmata: Cunaxidae), pp. 22-42 in Zootaxa 3265 on page 36, DOI: 10.5281/zenodo.28074
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