5,765 research outputs found

    Zebraodes lucidalis Heppner & Bae, new species

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    Zebraodes lucidalis Heppner & Bae, new species (Figs. 1–18) Type locality. Vietnam (Cuc Phuong National Park (Mac Lake), Ninh Binh Prov.). Diagnosis. This species is distinctive by its mottled, sometimes silvery forewing markings which are mostly solid along the costal margin and vertically aligned on the remainder of the forewing. The forewings and dark hindwings overall have a slight bluish iridescence in some individuals. Description. Male (Figs. 1–2). Head (Figs. 4–6): Vertex dark tan, with gray between antennal bases; frons gray, with dark gray along eye margins; labial palpus pale gray, with tan basally and mesally; antenna gray-brown. Thorax: Dark gray, with posterior spot bronze-gold; tegulae tan-gray; venter cream-white; legs gray mixed with some tan; tibiae and tarsi banded with dark brown, hindlegs mostly lustrous tan-gray. Wing expanse 24.0 mm in male (n = 9). Forewing (Figs. 1–2) dark brown to black-brown, more brownish towards dorsal margin, with overall slight bluish iridescence and violet iridescence on silvery markings, with silvery-gray markings as a solid band along costal margin about 1/4 of wing-width wide and mottled with lighter spots, remainder of forewing with numerous vertical silver irregular striae from costal band to dorsal margin, with intervening background black coloration relatively evenly spaced until the apical 1/5 where dark space is wider, and along termen; a very thin silver margin along termen; fringe dark gray; venter dark gray. Hindwing uniformly dark brown to gray, with slight bluish iridescence; venter dark gray. Abdomen: Dark gray, with bronze shine to slightly bluish towards posterior; venter gray, with light tan on posterior half of each sternite; genital tuft tan. Male genitalia (Figs. 10–11, 16–17) as described for genus. Female (Figs. 3, 7–9). Head and Thorax: Same as male, but wing expanse 27.8 mm (n = 1). Abdomen: Female genitalia (Figs. 12–13, 18) as described for genus. Specimens examined. Holotype ♂ (Fig. 2): Vietnam: Cuc Phuong National Park, Mac Lake, 155 m, Ninh Binh Prov., 9–13 April 2012, J.B. Heppner (gen. slide JBH 3495 ♂) (MGCL). Paratypes (8♂ 1♀): Vietnam: Same data as holotype (4♂, 1♀) (gen. slide JBH 3159 ♀) (MGCL); 10 Aug 2005 (1♂), Y.S. Bae, J.H. Bae, & N.V. Pham (INUC). Ninh Binh Prov.: Doi Vau Village, Cuc Phuong, 135 m, 10–16 Sep 2013 (1♂), J.B. Heppner (MGCL). Hatay-Hanoi Prov.: Ba Vi Natl. Pk., 455 m, 19–23 Jul 2010 (1♂), J.B. Heppner & Y.S. Bae (gen. slide JBH 3101 ♂) (MGCL); Ba Vi Natl. Pk., 800 m, 31 Jul 2010 (1♂), Y.S. Bae & X.V. Le (INUC). Etymology. The species name refers to the lustrous shine of the forewings. Biology. Unknown. Distribution. Recorded only from northern Vietnam. Discussion. Adults were collected mostly at Cuc Phuong National Park where two forms were taken: a lighter, more silvery form (Fig. 1) and the more common darker form (Figs. 2–3). Specimens of the dark form include both males and females, but the light form is thus far known only in one male; thus the dark form is likely the typical form of the species. The dark form has a slight violet iridescence on the silvery markings in addition to the overall bluish iridescence, whereas in the light form the violet shine is less evident. The male genitalia of the light form are shown in Fig. 16; those of the dark form are shown in Fig. 17. Because no significant differences could be detected between the genitalia of the two forms, we considered them conspecific. Although not clearly visible in Fig. 16, the male genitalia of both forms have a spatulate apex of the gnathos.Published as part of Heppner, John B. & Bae, Yang-Seop, 2017, Zebraodes, new genus, with a new species from Vietnam (Lepidoptera: Tortricidae: Tortricinae: Archipini), pp. 392-400 in Zootaxa 4236 (2) on pages 396-397, DOI: 10.11646/zootaxa.4236.2.13, http://zenodo.org/record/32180

    Tambitnotia peruviana Heppner & Bae, new species

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    Tambitnotia peruviana Heppner & Bae, new species (Figs. 1–6) Type locality. Pampa Hermosa Lodge (1220 m), [near San Ramon], Department Junín, Peru. Diagnosis. This species is very similar to T. tambita from Colombia, but it differs in having a darker gray, more expanded patch near the middle of the forewing dorsum, and with the apical strigulae in greater contrast to the ground color than in T. tambita. The male genitalia of T. peruviana have decumbent socii with a rounded distal tip in comparison to the upright, distally pointed socii of T. tambita. Plus the cucullus has a slightly different shape between the two species. Description. Wing length 17.4–19.0 mm (n = 3). Male (Fig. 1). Head (Figs. 3-4): Vertex and frons white; labial palpus white; antenna dark brown, with sparse silvery-pale white scales dorsally (pair each segment). Thorax: White, with posterior end dark gray; tegulae white; venter white; legs lustrous white except for foreleg black tarsal spines and gray foretibia; foretarsae gray-brown, with white segmental rings distally; mid-tibia pale tan, with spine-like scales on dorsal margin; mid- and hindtarsi dull pale tan, some white dorsally and black tarsal spines ventrally. Forewing gray with pale whitish basal 1/3 but gray also along the dorsal basal margin; dark brown strigulae past whitish base to apex along costal margin, with midway strigula extended darkly to end of cell, likewise for preapical strigula extending towards central apical region, and with apical strigulae on a pale yellow-white field; midwing a dark gray patch at angle from dorsal margin to center of wing, and with pale irrorations; a sinuous extension of the midwing dorsal strigula, then extended to tornus; tornus with a dark patch, distally somewhat paler and a subterminal orange area adjacent to 3 distinct black-brown spots along termen near the tornus; fringe white with scales basally gray-brown; venter gray-brown, with slight green iridescence in cell, with tan costal strigulae on dark costal margin except for white field on apex; lighter dorsal margin. Hindwing white (somewhat translucent), with distal 1/3 along termen dark brown; translucent white, with gray-brown along costa and termen. Abdomen: Lustrous white; venter cream-white; genital tufts tan with silvery-gray dorso-centrally. Male genitalia (Fig. 5) as for genus with uncus obsolete and truncated tegumen; gnathos weakly sclerotized; large, medium sclerotized socius (Fig. 5 a) as carinate ridge and posterior acute points, all slightly recurved in position fused to tegumen; valva elongated and narrowed, with quadrate and angulate saccus, and oval cucullus with setae and strong spines on margin, sacculus slightly offset from center to distal end of neck; phallus small, narrow, with angled distally, vesica with field of small thorns; vinculum reduced; juxta strongly sclerotized, triangular. Female (Fig. 2). Head and Thorax: As in male but head with vertex gray-white and labial palpus dull tan-white; forewing darker and more strongly white basally, and hindwing with the dark termen slightly wider than in male; ventrally same as male. Abdomen: Female genitalia (Fig. 6) with ovipositor short, setose; anterior apophyses about 1.2x as long as posterior apophyses; genital sternite (Fig. 6 a) quadrate with posterior convex extensions around spatulate and posteriorly truncated sterigma; a strong ridge on anterior margin of genital sternite; ostium circular in center of genital sternal plate; ductus bursae sclerotized, narrow, tubular in posterior 1/3, then widening to twice its width before end of sclerotization, with narrow ductus seminalis, which widens to small bulla seminalis; ductus bursae continues to corpus bursae about same length as from entrance of ductus seminalis to ostium, with a narrow sclerotized ridge ventrally posterior to corpus bursae; corpus bursae ovoid, as large as width of genital plate, with one large thorn-like signum (Fig. 6 b). Specimens studied. Holotype: Male, Pampa Hermosa Lodge (1220 m), nr. San Ramon [21 km N], Department Junín, Peru, 2–3 Nov 2009, J. B. Heppner (genitalia slide JBH 2827), (FSCA /McG). Paratypes (1♂, 1♀). Peru: Department Junín: Pampa Hermosa Lodge (1220 m), nr. San Ramon 21 km N, 2–6 Apr 2011 (1♂), J. B. Heppner (genitalia slide JBH 2920). Department Cusco: San Pedro (1368 m), Kosñipata Vy., [Manu Dist.], 11–15 Nov 2009 (1♀), J. B. Heppner (genitalia slide JBH 2908) (FSCA /McG). Etymology. Named after Peru. Biology. Unknown. Distribution. Known only from central Peru, Departament Junín and Cusco. Discussion. The new species is very similar to T. tambito but easily separated as noted under the diagnosis, particularly in regard to the downward directed socius, the latter being upright in T. tambito. The female paratype provides the first details of the genitalia and absence of sexual dimorphism for the genus. Because the species from Colombia and Peru have such similar males, it is likely that the females of the two species are similar, as well. Based on the row of tiny dots along the forewing termen, Tambitnotia may belong to Grapholitini, but this question requires further investigation. The type-locality in Department Junín is just north of the town of San Ramon in a small river valley at about 1220 m elevation at the base of the Reserva Forestal Pampa Hermosa and is surrounded by tropical montane forest. This area is part of a very biodiverse alto-Amazonian region of Peru long known as Chanchamayo, a long river valley from the Andean foothills into the nearest Amazonian jungle east of Lima. Numerous lepidopteran species have been described from here over the past 150 years, but few micro-moths. The same is true of the locality for one of the paratypes, the Manu District of Department Cusco.Published as part of Heppner, John B. & Bae, Yang-Seop, 2016, A new species of Tambitnotia from Peru, with the first reported female of the genus (Lepidoptera: Tortricidae: Olethreutinae), pp. 446-450 in Zootaxa 4196 (3) on pages 446-450, DOI: 10.11646/zootaxa.4196.3.10, http://zenodo.org/record/16813

    Fansipaniana tamdaoensis Heppner & Bae, new species

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    Fansipaniana tamdaoensis Heppner & Bae, new species (Figs. 1–5) Type locality: Tam Dao (930 m), Vinh Phuc Province, Vietnam. Diagnosis. Diagnostic characters for this species include the lighter basal forewing area (darker in F. fansipana Razowski) and the male valva lacking the bulge midway on the valva, as well as dense lateral valval spines nearly twice as numerous along the mesal valval margin as in F. fansipana. Description. Male (Fig. 1). Head: (Figs. 2–3): Gray with scales white-tipped; same on frons and labial palpus. Antenna with segmental row of black scales each segment (basally white), with venter denuded and dull orange chitin only. Thorax: Gray (scales with white bases) and tegulae same; venter white. Legs black on fore- and midlegs, with thin white rings at tibial base joint and tarsal segments; mesally white; mesofemur white ventrally toward joint with tibia; coxae black; hindleg white at femur-tibial joint and most of femur, but tibia white-tan and tarsal segments brown-black with thin white ring for each tarsal segment; femora and tibiae dorsoventrally white. Wing expanse: 12.6–14.3 mm (n = 3). Forewing brown with tan-umber patches near base at 1 / 4 near costa and at 1 / 3 near dorsal margin; with similar tan patch at tornus and apex, and light tan triangular patch on apical 1 / 3 near costa; a dark brown bar at midwing from costa to cell and another dark brown area midway along termen; apical margin brown, slightly intersected almost midway by the extensive point of the apical 1 / 3 light tan patch; fringe dark brown except for brown at apex and light tan at tornus; venter brown at base and along dorsal 1 / 2 of wing, and darker brown on costal 1 / 2 of wing, with black-brown along costal edge and with 5 white strigulae (three evenly spaced on basal 1 / 2 and two more on apical 1 / 3, and large orange patch on apical 1 / 4 by apex emarginated with yellow basally; ventral fringe gray to black distally, with an umber fringe at tornus. Hindwing gray, slightly lighter towards basal, with a umber area on apex; fringe light gray except for umber on apex; venter pale gray-tan and red-brown at apex and extended basally along costa to 1 / 2 wing; ventral fringe silvery gray. Abdomen: Lustrous gray, lighter towards base on T 1–2, posteriorly with black-tipped scales; venter dark gray, with white-tipped scales, with basal segments S 1-3 mostly dull white; genital tuft dark gray with white-tipped scales, valvae pale tan and with large lateral scale tufts; pregenital plates enlarged (Fig. 6), with lateral margin widened and a central sclerotization extended posteriorly as a narrow line. Male genitalia (Figs. 4–5) with tegumen elongated distally to blunt end; uncus a small convex extension from tegumen, basally with slightly enlarged ends; gnathos-like sclerotization with bulbous dorsal end. Valva bent upwards from straight basal 1 / 4, the upright portion subequal in width, with strong setae directly mesally along mid-portion, with a short recurved setose dorsal process near base; sacculus elevated setose area from the disc of valva; vinculum short, without extended saccus. Anellus clothes-pin-like (upside down), with bulbous basal area, the upper arms extended around phallus; phallus short, with extended hood dorsally. Female. Unknown. Specimens studied (n = 3). Holotype, male, Vietnam, Tam Dao, 930 m, Vinh Phuc Province, 10–12 Oct 2014, J. B. Heppner (JBH photo 12594; genitalia slide JBH 3300). Deposited in FSCA / MGCL (Florida State Collection of Arthropods, McGuire Center for Lepidoptera and Biodiversity), on indefinite loan from PPRI (Hanoi, Vietnam). Paratype, male, Vietnam, same locality as holotype, 13–15 Oct 2014, J. B. Heppner (FSCA / MGCL). Additional specimen, male, Vietnam, Cuc Phuong Natl. Pk., Mac Lake, 155 m, Ninh Binh Province, 25–26 Apr 2012, J. B. Heppner (JBH photo 12867; genitalia slide JBH 3408) (FSCA / MGCL). Etymology. The species is named after Tam Dao, Vinh Phuc Province, Vietnam. Biology. Nothing is known of the life history and biology of the species. Distribution. Known only from northern Vietnam. Discussion. The new species superficially is very similar to F. fansipana, described from the Fansipan Mountains, near Sapa (1600 m elevation), near the northern border with China. As already noted, the basal forewing is more tancolored in the new species, whereas it is much darker in F. fansipana. In addition, the male genitalia are diagnostic, with the valvae evenly narrow in F. tamdaoensis but bulging mesally beyond midway in F. fansipana, and the mesal margin spines are more compact in F. fansipana than in F. tamdaoensis; in the later the spine area is much longer (nearly twice as long as in F. fansipana). A further specimen from a lowland site (Cuc Phuong Natl. Park, Ninh Binh Province, about 150 km south of Tam Dao) is not included in the type series. It is darker and smaller (wingspan 12.6 mm) than the Tam Dao specimens (wingspans 13.7–14.3 mm), but because the genitalia show no significant differences from those of F. tamdaoensis, the specimen is most likely conspecific with F. tamdaoensis. Nedoshivina (2013) reported two new records of F. fansipana from Dong Nai Province, a lowland site (121–132 m) about 1300 km south of Sapa near Hochimin (= Saigon) in southern Vietnam; however, we did not examine these specimens. Presumably, they conform in genitalia and adult characters to the Sapa population of F. fansipana as identified by Nedoshivina (2013). Unfortunately, all specimens of Fansipaniana known thus far are males, so females cannot be compared.Published as part of Heppner, John B. & Bae, Yang-Seop, 2016, A new species of Fansipaniana from Northern Vietnam (Lepidoptera: Tortricidae: Olethreutinae: Olethreutini), pp. 135-138 in Zootaxa 4097 (1) on pages 135-137, DOI: 10.11646/zootaxa.4097.1.10, http://zenodo.org/record/26693

    Prophaecasia malawiana Heppner & Bae 2017, new species

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    Prophaecasia malawiana Heppner & Bae, new species (Figs. 1–4) Type locality: Malawi (Senga Bay, Salima District). Diagnosis. Superficially, Prophaecasia malawiana can be distinguished by the pale tan scaling of the head scaling (olive-green in P. anthion, olive gray in P. usambarae), and the bicolored gray-green and dark pink forewing (olive-green and light pink in P. anthion, green and pink and white in P. lindae). Also, the forewing white vertical stria present in P. anthion and P. lindae is absent in P. malawiana. In contrast to P. usambarae, the pink mark at the apex of the forewing is lacking in P. malawiana, and the inner margin of the pink distal forewing patch is curved inward in P. malawiana, whereas it is straight in P. usambarae. In the male genitalia the longer and wider caulis arms and reduced base of the caulis of P. malawiana differ from that found in congeners, and in P. usambarae the valvae are recurved ventrally (dorsally in P. malawiana), whereas in P. caemelionopa the valvae are strongly convex toward the apex. Description. Male (Fig. 1). Head (Figs. 2–3): Vertex pale tan-white; frons pale tan-white; labial palpus pale tanwhite; antenna bronze-brown, with scape light tan-white. Thorax: Grayish pale olive-green, with anterior margin dark brown; tegulae with some dark brown scales; venter white; legs pale tan-white, with dark gray dorsally and black-brown bands on tarsi, hindlegs tan-white except for small black-brown spot on distal two tarsi. Wing expanse 10.8–11.3 mm male (n = 2). Forewing (Fig. 1) gray in basal half, with irregular mottling of pale gray, interrupted striae and spots, with fine, faint, interrupted black striae from costa to dorsal margin; costa with small black strigulae from base to apex, with short white strigulae between black strigulae; distal half of forewing deep pink with narrow black line along basal margin, arched from dorsal margin to near apex, with a slender pink line directed towards wing base near costal edge of pink region; entire pink area covered with irregular black spots and dashes, some merged into a subterminal black line from apex to tornus; apical half of wing dark gray along costa except for one pink line extended from pink area to near apex; termen with fine line of pink, with several black spots on upper termen; fringe pink with some black scales on termen and tornus; venter dark gray, with pale gray-tan on cubital section. Hindwing dark gray-brown; fringe dark gray with a basal line of pale tan, but mixed with pink from apex to tornus; venter dark gray. Abdomen: Dorsum lustrous dark gray; venter light tan; genital tuft light tan; pregenital plates (Fig. 4b) tree trunk-shaped (or spearhead-like) on sternite, with a flanged base at anterior end, acute and attenuated posteriorly. Male genitalia (Fig. 4) with tegumen somewhat quadrate, truncated dorsally, lacking distinct uncus; gnathos a thin rod-like structure curved dorsally; socius (Fig. 4a) with broad, short arm, heavily setose on distal half, with very long setae; valva with straight sacculus, followed by an 45° angle dorsally to narrow and somewhat acute apex, densely setose, with strong, dense setae on saccular angle and a separate wide setal tuft near base; juxta (Fig. 4a) triangular at base and as Y-shaped short narrow flanged caulis with wide dorsal arms (twice as long as rod-like base) of juxta-caulis complex; anellus a circular short collar fused to caulis arms; phallus short and tubular, with indistinct cornuti; vinculum undeveloped, rounded. Female unknown. Specimens studied. Holotype male: Malawi: Safari Beach Lodge, Senga Bay, Salima Dist., 13–16 Feb 2009, J. B. Heppner (gen. slide JBH 3497). Paratype (1♂): Malawi: Makuzi Lodge, Chinteche (480 m), Nkata Bay Dist., 23–24 Feb 2014 (1♂), J.B. Heppner & R.J. Murphy. Etymology. The species is named after the country of Malawi. Biology. Unknown. Distribution. Known only from Malawi. Discussion. Prophaecasia malawiana appears to be most closely related to P. anthion from Borneo and P. lindae from Cambodia. Both of the latter species have male genitalia very similar to the new species from Malawi, other than the small caulis of P. malawiana. Another species possibly related to P. malawiana is from Nigeria, Anthozela prodiga Razwoski & Wojtusiak (2012), with male genitalia similar to Prophaecasia, and it is likely that this species also belongs in the genus Prophaecasia. The wing maculation of P. malawiana also is remarkably like that of A. prodiga, but the genitalia differ considerably, whereas the male genitalia of P. malawiana are very similar to those of P. anthion and P. lindae. Razowski & Wojtusiak (2012) have placed other Africa species in the genus Anthozela that are very similar in maculation to P. malawiana and probably belong to other genera. Anthozela prodiga possibly belongs in Prophaecasia; Anthozela postuma Razowski & Wojtusiak probably belongs in Loboschiza; and Anthozela usambarae Razowski & Wojtusiak (2014), is transferred to Prophaecasia. All species of Prophaecasia are known from very few specimens, so additional material is required before any convincing conclusions can be drawn.Published as part of Heppner, John B. & Bae, Yang-Seop, 2018, Prophaecasia species in Malawi (Lepidoptera: Tortricidae: Olethreutinae: Olethreutini), pp. 133-136 in Zootaxa 4388 (1) on pages 133-136, DOI: 10.11646/zootaxa.4388.1.10, http://zenodo.org/record/118779

    Anthozela cypriflammella Heppner & Bae 2018, new species

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    Anthozela cypriflammella Heppner & Bae, new species (Figs. 1–8) Type locality: Cuc Phuong Natl. Park, Mac Lake (alt. 155 m), Ninh Binh Province, Vietnam. Diagnosis. Anthozela cypriflammella is superficially similar to other species of Anthozela, even those form Africa, but can be distinguished by the strong carinate ridge on the sacculus of the male genitalia and the absence of strong posterior spines on the 8th sternal plate. The female genitalia lack an accessory bursa but have a wide colliculum (usually narrower in other congeners) and a subtriangular, funnel-shape ostium. Description. Male (Fig. 1). Head (Figs. 2–3): Vertex yellow-tan, mixed with some dark brown; frons same; antenna dark brown with few yellow-tan scales; scape yellow-tan, with dark brown streak dorsally; labial palpus light yellow, with apical segment light brown, mesally same. Thorax: Light yellow mixed over dark brown, more yellow posteriorly; tegulae same; venter tan-white, with light yellow-tan at neck; legs white basally and femurs, with tibia tan and with two angled dark brown rings; midtibia with spine-like scales on dorsal margin; hindtibia lustrous gray; tarsal segments alternating gray and pale tan. Wingspan 10.5–10.8 mm in males (n = 2), 10.8 mm in female (n = 1). Forewing with yellow scales scattered over black-brown on basal half of forewing, with indistinct black-brown costal strigulae, and with a cross-wing vertical fascia of silver-gray, straight mesally but very irregular distally, nearly to costal margin but interrupted by thin line of burnt-orange and a fine silver line angled from costal black-brown strigula towards apex; apical half of costa with several black-brown strigulae angled towards tornus, each separated by burnt orange, and on costal margin by light yellow; apical half of forewing otherwise reddishburnt-orange with several irregular lines and spots of silver-gray, subtermen and tornus with similar coloration; fringe black-brown, with a near-apical and near-tornal patch of light orange. Venter gray-brown with pale white on cell and cubitus and dorsal margin, with tan and dark brown strigulae on costal margin as in dorsal surface. Hindwing dark brown-gray with slightly paler area near midwing. Venter similar to dorsum but duller. Abdomen: Brown-gray; venter pale tan-white, with brown anterior line on each sternite; genital tuft brown-gray; pregenital plates as simple posteriorly concave tergal plate (Fig. 7b), intersegmental membrane connected to male genitalia very long (subequal to 8th segment), with dorsal setal patch on each side. Male genitalia (Fig. 7) with tegumen elongated as strong reversed V-shape, with strongly convex uncus; socius laterally as curved concave plate with strong row of stout, comb-like spines directed mesally, of uniform length (Fig. 7a); valva elongated, setose, with slightly bulbous distal end, and saccular margin with a strong dorsally-directed carinate ridge about 1/6 of valval length from base; vinculum simple, V-shaped; juxta a narrow band with lateral flattened quadratic basal plate, elongated dorsally to narrow arms to anellus, latter with a longer emarginated flattened lateral flange-like plate; caulis dorsally with groove-like invaginated apex with wedge-shaped lateral point around phallus; phallus short and slender, slightly bulged medially, vesica with patch of about 30 cornuti as thumbtack-like spines (points directed outwardly); Female (Fig. 2). Head and Thorax: Essentially as in male, but head with somewhat paler yellow-white scaling, particularly on labial palpus; hindwing with midwing pale area somewhat more contrasting than in male. Abdomen: Female genitalia (Fig. 8) elongated, with short ovipositor, papilla anales setose and flattened; sterigma a simple semicircular ring at posterior margin to simple, subtriangular funnel-shaped ostium bursae (Fig. 8a) equal in width to ductus bursae; colliculum somewhat indistinct and of similar width as ductus bursae; length of ductus bursae subequal to that of corpus bursae, with sinuous entry into corpus bursa as the ductus and bursae gradually widen to full bursal width; ductus seminalis emergent at posterior end of corpus bursa; corpus bursae large, ovate, with simple scobinations, and two large, elongated, thorn-like signa on lateral wall of bursa (Fig. 8b); secondary bursa absent. Holotype: ♂, Vietnam, Ninh Binh Province, Cuc Phuong National Park., Mac Lake, 155m, 4–7 May 2009, J. B. Heppner (genitalia slide JBH 3491), deposited in MGCL on indefinite loan from PPRI (Hanoi, Vietnam). Paratypes (1 ♂, 1 ♀): 1 ♂, same data as holotype (MGCL); 1 ♀, Ba Be Natl. Park, 240 m, Bac Kan Province, 21–26 Jun 2008, J. B. Heppner (genitalia slide JBH 3323) (MGCL). Biology. The biology of the new species is unknown. Larval hosts have been reported for three species of Anthozela: A. anonidii Ghesquière, which has been recorded from the fruit of Anonidium manii (Annonaceae) (Ghesquière 1940); A. chrysoxantha Meyrick, which has been recorded from Pavetta sp. (Rubiacacae) (Meyrick 1936); and A. hemidoxa (Meyrick), the pepper shoot borer or top shoot borer, which has been reared on shoots of pepper vine, Piper nigrum (Piperaceae), in India (Fletcher 1921, Devasahayam & Abdulla Koya 1993, 1994) and Indonesia (Kalshoven 1950). Horak (2006) notes that larvae of A. hemidoxa have been found on Piper kadsura (Piperaceae) in the Ryukyu Islands, but comments that there may be more than one species of Anthozela involved. Distribution. The new species is known only from northern Vietnam. Etymology. The specific epithet refers to the red and yellow forewing coloration, latinized for "beautiful little flame-colored" moth.Published as part of Heppner, John B. & Bae, Yang-Seop, 2018, A new species of Anthozela from Vietnam, with a list of species in the genus (Lepidoptera: Tortricidae: Olethreutinae: Enarmoniini), pp. 394-400 in Zootaxa 4450 (3) on pages 395-398, DOI: 10.11646/zootaxa.4450.3.6, http://zenodo.org/record/144486

    Dichrorampha sapodilla Heppner. B 1981

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    Dichrorampha sapodilla Heppner, 1981 Hemimene sp., Kimball, 1965 Dichrorampha sapodilla was described by Heppner (1981) along with another new species of Dichrorampha Guenée from Florida based on a specimen in the Baranowski collection at the USNM. D. sapodilla feeds on sapodilla, while D. manilkara Heppner, 1981 feeds on wild dilly (Manilkara bahamensis (Baker) Lam. and Meusse). It is a congener of D. azteca Walsingham, 1914 and D. odorata Brown and Zachariades, 2007. Description. Egg. Translucent yellow in newly laid eggs and translucent golden-yellow before they hatch. Larva. Transparent yellowish-white with head dark-orange from first to fourth instar; fifth instar body not transparent; dark yellowish-white; head dark-orange. Mature larva length 2.9–3.2 mm (Fig. 1A). Pre-pupa. The same as the last instar, but thoracic segments wider and body paler. Pupa. Yellow except for head, which is brown; some spines present dorsally on abdomen. Adult. Not sexually dimorphic. Wing expanse: 3.6–4.7 mm. Wings. Dorsal surface: forewings narrow and long; gray with dark band angled toward apical area from middle of wing to basal third of inner margin; six to seven matte-black strigiform markings in costal area continue diagonally toward tornus, surrounded by orangish-yellow; costal margin with white borders; four apical black strigiform markings surrounded by orangish-yellow through tornus distad acute curve at radius; tornus without orangish-yellow; a large matte-dark spot with orangish-yellow on distal area of spot; speculum with five small dark spots in orangish-yellow area; matte-gray in fringe; hindwings dark-gray; fringe pale-gray. Ventral surface: Both wings shiny bronze. Head. Antennae: Filiform, distally brown and dark-gray, ⅓ body length. Vertex: Grayish-brown. Mouthparts: Palps small pale brownish-gray. Body. Thorax and abdomen: Brownish-gray above and pale-gray below. Legs: Brownish-gray (Fig. 2A). Distribution. Dichrorampha sapodilla is native to the USA. It has only been found it in south Florida (Heppner 1981; Myers et al. 2008). Hosts. Manilkara zapota (L.) van Royen recorded by Heppner (1981) and Myers et al. (2008). Natural history. The adults have been observed flying year-round in Florida (Heppner, 1981). Females lay eggs on the petals and sepals of mature flowers. Upon hatching, the neonate larva bores into the flowers and eat the petals and the base of the pistil. The duration of larval development ranges from 12–16 days. Pupation takes place inside of the sepals where the larva makes a silken cocoon utilizing its excrement. The duration of the pupal stage is close to 15–17 days. Adult moths usually feed on nectar from flowers and also drink water; however, the feeding habits of adult moths of the sapodilla pod borer have not been studied. This species has been observed only in sapodilla (Heppner, 1981). Damage. The larvae feed on mature flowers only, mainly on the petals and the base of pistil, damag- ing several flowers until they complete larval development and begin pupation. The petals in damaged flowers turn reddish brown, dry and die. This moth can cause significant losses. Management. Currently just one parasitoid, an unidentified species of Bracon, has been found associated with larvae of the sapodilla pod borer. There are other natural enemies (e.g. Gasteracantha cancriformis (Linnaeus)). Potential insecticides for the sapodilla pod borer moth have not been thoroughly investigated (Myers et al. 2008).Published as part of Martinez, Jose I., Crane, Jonathan H., Wasielewski, Jeff, Miller, Jacqueline Y. & Carrillo, Daniel, 2019, Lepidoptera pests of sapodilla (Manilkara zapota (L.) van Royen) in south Florida, with some comments on life history and natural control, pp. 1-26 in Insecta Mundi 739 (739) on pages 14-15, DOI: 10.5281/zenodo.367659

    Torodora squariella Park & Heppner 2023, sp. nov.

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    Torodora squariella Park & Heppner, sp. nov. LSID: urn:lsid:zoobank.org:act: D40B94A4-4593-41BC-88FB-494A195D43ED (Fig. 14) Type specimen. Holotype: Female, N. Sumatra, 7 km SE Prapat, 30 Aug. 1992, ca. 1,400 m, J.B. Heppner & E.W. Diehl, gen slide no. JBH-4169. Diagnosis. This new species is superficially similar to the Oriental species. T. macrosigna Gozmány, 1973, but larger (viz. 15–18 mm in T. macrosigna). The forewing is less elongated and abdominal sternite VIII is deeply emarginated into a U-shape. The female genitalia are characterized by a large signum, more or less elliptical. Description. Female (Fig. 14A). Wingspan 19.5 mm. Head: frons pale gray-tan; scales over scape to frons dull white.; vertex mustard brown. Antenna slightly longer than forewing; scape white, with small pale brown spot dorsally; flagellum creamy white ventrally, dark brown dorsally in basal 2/3, then with dark brown annulations beyond. Labial palpi broken away (only base of segment 1 remains and it is white). Thorax: tegula and thorax mustard brown. Hind tibia with mustard brown rough scales dorsally, with white apex; tarsi yellowish white dorsally, with mustard brown band apically on each segment. Forewing ground color mustard brown evenly; costa arched around basal 1/4, then nearly straight, with very small, yellowish-white, wedgeshaped patch at 3/4, post median line weakly presented from costal patch to near basal 4/5 of inner margin; apex obtuse; termen slightly concave before middle; fringe concolorous. Hindwing ground color same as that of forewing. Abdomen: mustard brown on dorsal surface, excepting dorsal surface of segment VIII. Female genitalia (Figs 14B–D): Apophyses anteriores very short, 1/3 the length of apophyses posteriores. Abdominal sternite VIII deeply emarginated at middle, about half length of sternite VIII (Fig. 14C). Antrum large, membraneous, Cup-shaped, with flat caudal margin; broader distally and narrowed toward conjunction with ductus bursae, about 1/3 the length of ductus bursae. Ductus bursae broad; ductus seminalis arising from near distal end. Corpus bursae semi-ovate, as long as ductus bursae; signum large, more or less elliptical, slightly concave on caudal margin, densely covered with minute conical spines. Male unknown. Distribution. Indonesia (N. Sumatra). Etymology. The specific name is derived from the Latin, square (= quadrate), referring to the quadrate signum of the female genitalia.Published as part of Park, Kyu-Tek & Heppner, John B., 2023, Review of the genera Thubana Walker and Torodora Meyrick (Lepidoptera: Lecithoceridae) from Sumatra, Indonesia, pp. 434-456 in Zootaxa 5256 (5) on pages 454-455, DOI: 10.11646/zootaxa.5256.5.2, http://zenodo.org/record/775883

    Ukamenia babeana Heppner & Bae 2020, new species

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    Ukamenia babeana Heppner & Bae, new species (Fig. 1 -4) Type locality: Vietnam (Ba Be Natl. Park, Bac Kan Province). Diagnosis. This species is distinctive in the female genitalia, with a wide, massively sclerotized, tubular, sclerotized protrusion from the ostium in the female (compared to a narrow protrusion in U. sapporensis). The black-brown forewings with numerous blue-metallic spots are otherwise distinctive for both species of Ukamenia, as also the dull tan-brown head and thorax. Description. Wing expanse: 15.4 mm female (n = 1). Female (Fig. 1). Head (Fig. 2‒3): Vertex and frons brown with scales tan-tipped; labial palpi similar but with bands of brown on second and distal segments; antenna and scape tan, with scattered dark brown scales dorsally, venter light tan. Thorax: Brown with scales tan-tipped, and with blue spots on patagia, blue medial line and at posterior ends of tegulae; venter white-tan with golden iridescence. Legs with coxae tan and golden; tibiae and femorae dark brown, mesally golden (ventrally pale tan); tarsi dark brown, with segmental ends margined with white-tan. Forewing (Fig. 1) overall FIG. 4. Ukamenia babeana n. sp., female genitalia, with details. A. Sterigma and ostium. B. signum at posterior end of bursa (gen. slide JBH 3131). black-brown, with numerous blue-iridescent spots, scattered and subequidistant on basal half of forewing, with the distal margin of this blue-spotted field being angeled toward tornus, and with the distal 1/3 of forewing with the blue posts more densely arranged in a rough subterminal band and leaving the apical 1/4 with only a few blus spots on a dark orange field; a terminal fine black line; fringe silvery gray; venter gray (with copper-golden iridescence), with pale white on dorsal margin. Hindwing dark gray-brown, paler on basal half; fringe brown, distally dull white; venter gray, with copper-golden iridescence. Abdomen: Lustrous gray-brown; venter silver-white and gradually to dark gray on posterior segments; genital tufts dark gray. Female genitalia (Fig. 4) with ovipositor relatively short (length subequal to last abdominal segment); pa- pilla anales nearly flattened, setose with ragged emarginations; sterigma circular with weak basal band around the ostium and with a greatly extended and heavily sclerotized tubular projection (as long as ovipositor); ductus bursae posteriorly as narrow colliculum-like sclerotized tube, then gradually broadening towards the bursa as a single coil before entering bursa; ductus seminalis emergent from posterior area of widened ductus bursae just anterior to sclerotized tubular sec- tion and narrow until ovate bulla seminalis; corpus bursae ovate and overall scobinate, with single ventral circular and centrally 4-pointed, star-like signum and surrounded by denser scobinations. Male unknown. Holotype. ♀, Ba Be Natl. Pk. (elev. 255 m), Bac Kan Prov., Vietnam, 20‒23 Sep 2013, J. B. Heppner (gen. slide JBH 3131; photo 11830) (deposited at McGuire Center for Lepidoptera and Biodiversity, MGCL). Etymology. The species is named after Ba Be National Park, Bac Kan Province, Vietnam. Biology. Unknown. Ukamenia sapporensis is known to feed on Castanea and Quercus (Fagaceae), Hamamelis (Hamamelidaceae), and Vaccinium (Ericaceae), thus the hostplant for the new species may also be among these plant families. Distribution. Known only from northern Vietnam. Whereas U. sapporensis occurs from northern China to the Russian Far East and Korea to Japan, the new species may range beyond northern Vietnam since the border with China (Yunnan) is nearby. Discussion. The new species is a surprising addition to the genus Ukamenia, being so distant from the known range of U. sapporensis, as well as not having been discovered previously in Vietnam after many years of intensive collecting of micro-moths. Since the male of the new species remains unknown, the male genitalia of U. sapporensis (Fig. 5) are shown herein for future comparison for what may be similar morphology of male U. babeana. Likewise, the female genital characters are shown for U. sapporensis (Fig. 6) for comparisons: note the colliculum-like (narrow and with an external protrusion much longer in the new species), and the similar star-shaped signa (4-pointed in the new species, 5- or 6-pointed in U. sapporensis). We have two additional new species of blue-spotted olethreutines from northern Vietnam, both also known thus far from only single females, yet notwithstanding their similar wing maculation, these clearly represent species of the tribe Grapholitini based on their genitalia and other morphological features.Published as part of Heppner, John B. & Bae, Yang-Seop, 2020, A new species of Ukamenia from Vietnam (Lepidoptera: Tortricidae: Olethreutinae Gatesclarkeanini), pp. 280-284 in Zootaxa 4743 (2) on pages 281-283, DOI: 10.11646/zootaxa.4743.2.11, http://zenodo.org/record/368773

    Ambulatory assisted living fallers at greatest risk for head injury

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    OBJECTIVES: To determine the relationship between head injuries sustained during each fall with various known high risk health and demographic factors predictive of falls. DESIGN: Prospective cohort study conducted over 1 year SETTING: Assisted living and skilled nursing units of a Continuing Care Retirement Community located in the northeastern United States. PARTICIPANTS: Sixty nine OAs who fell. MEASUREMENTS: Age, gender, diagnosis, high risk medication, functional, cognitive, ambulation/elimination status, mode of locomotion, fall related symptoms and the position of the fall, were analyzed using General Estimating Equations among elderly fallers with and without head injury. RESULTS: A total of 173 falls (average of 2.9 times) were observed for 62 patients who had complete injury data. Injuries were recorded in 40.5% of falls, with 41.4% being head injuries. Head injuries were more likely to be hematomas than lacerations (66.7% vs. 14.7%) and among assisted living residents (p=0.04). Head injured patients were more likely to be walking at the time of the fall (69% vs. 36.1%) and less likely to have bowel incontinence (3.5% vs. 28.5%; p=0.04). None of the high risk diagnosis or medications associated with falls risk increased risk for head injury. CONCLUSION: Those at greatest risk for head injury were ambulatory assisted living residents. None of the known clinical conditions predictive of risk to fall were predictive of head injury. For head injury prevention to be successful we need a closer examination of resident’s mobility, shoe-wear, health behavior with respect to ability to use assistive devices, and floor surface landing area. Future health policy implications include measures to ensure standard of care practices for head injured patients are in place.This is the peer reviewed version of the following article: Gray-Miceli, D. L., Ratcliffe, S. J. and Thomasson, A. (2013), Ambulatory Assisted Living Fallers at Greatest Risk for Head Injury. Journal of the American Geriatrics Society, 61: 1817–1819, which has been published in final form at https://dx.doi.org/10.1111/jgs.12467. This article may be used for non-commercial purposes in accordance with Wiley Terms and Conditions for Self-Archiving.Peer reviewe

    Teaching Strategies for Atypical Presentation of Illness in Older Adults

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    Atypical presentation of illness is one of those phenomena where “seeing is believing”. Expert geriatric nurses and clinicians know all to well the early signs and symptoms of this frequent masquerader of bacterial infections, pain, acute myocardial infarction, heart failure or other serious medical ailments in older adults. Students however, as novices to clinical practice, require interactive learning approaches to reflect on the client’s illness presentations, help with developing the necessary skills to analyze and synthesize clinically relevant data, and to witness resolution of an atypical presentation when found and treated. We discuss various learner-centered, interactive approaches to teach students how to recognize an atypical presentation of illness using a real-life clinical case. Outlined are teaching strategies for faculty, drawn on visual, auditory, reading and kinesthetic modes of student learning. Use of the senses to teach nurses about care of patient’s is not entirely new or innovative, as reflected on by Florence Nightingale’s (1846) earliest writings of the "rules of nursing".Peer reviewe
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