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Taxonomy of Dolerus subfasciatus auct. and D. subfasciatus F. Smith with notes on the sawfly subgenus Equidolerus (Hymenoptera, Tenthredinidae)
Heidemaa, Mikk, Smith, David R., Shinohara, Akihiko (2012): Taxonomy of Dolerus subfasciatus auct. and D. subfasciatus F. Smith with notes on the sawfly subgenus Equidolerus (Hymenoptera, Tenthredinidae). Zootaxa 3525: 1-17, DOI: 10.5281/zenodo.20987
FIGURE 19 in Empria longicornis species group: taxonomic revision with notes on phylogeny and ecology (Hymenoptera, Tenthredinidae)
FIGURE 19. Phylogeny of mitochondrial sequences (complete COI and 3 partial tRNA) using MrBayes (GTR+4Γ model) and including all outgroup species (alignment length 1651 bp, non-coding and ambiguously aligned tRNA regions were excluded). Clades with posterior probabilities (PP) less than 0.90 have been collapsed. Maximum likelihood (ML) analysis (GTR+4Γ substitution model) produced nearly identical topology (few topological differences between Bayesian and ML analyses were not significantly supported, i.e. less than 0.9 PP and 0.7 BP). Bayesian and ML analyses using E. immersa and E. fletcheri as the only outgroups produced identical topologies with only slight differences in clade support values. al, E. alector; alp, E. alpina; bas, E. basalis; jap, E. japonica; lokt, E. loktini; long, E. longicornis; trid, E. tridens.Published as part of Prous, Marko, Heidemaa, Mikk & Soon, Villu, 2011, Empria longicornis species group: taxonomic revision with notes on phylogeny and ecology (Hymenoptera, Tenthredinidae), pp. 1-39 in Zootaxa 2756 on page 17, DOI: 10.5281/zenodo.20094
Empria minuta Lindqvist 1968
Empria minuta Lindqvist, 1968 Empria minuta Lindqvist, 1968: 30. Type locality: vicinity of Helsinki [original description] “Umgegend von Helsingfors” [Finland], “N. Helsinge” [from label]. Holotype Ψ, ZMH [examined]. Empria gussakovskii auct. nec Dovnar-Zapolskij, 1929. Host plants. Unknown. Distribution. Palaearctic. The verified country records are: Estonia, Finland, Mongolia, Russia (Leningrad Oblast, Yamalo-Nenets Autonomous Okrug), and possibly Sweden (the specimen UUZM _HS). Key characters. Tegulae partly or largely, posterior margin of pronotum, metatibia in basal 1 / 3, and paired patches on abdominal terga 2–6 unpigmented. Ratio of flagellum length to head breadth in females 1.8–2.1, in males 2.8 –3.0, and eye length to head length in females 1.6–1.7, in males 1.7–2.1. Number of serrulae 14–16, slightly papilliform shaped (Fig. 38). Valviceps with medium sized lobe, forming a notch between valviceps and valvura (Fig. 58); valvura to valviceps length ratio 0.63–0.72. Notes. There is no information about the locality and the collector of the female specimen from UUZM, but it has probably been collected from Sweden. The flight period of the adults might partly explain why this species has been collected rarely. The Estonian specimens were found in a Malaise trap only between 12–30 April (3 ɗ) and 30 April – 15 May 2010 (1 Ψ).Published as part of Prous, Marko, Heidemaa, Mikk & Soon, Villu, 2011, Empria longicornis species group: taxonomic revision with notes on phylogeny and ecology (Hymenoptera, Tenthredinidae), pp. 1-39 in Zootaxa 2756 on page 25, DOI: 10.5281/zenodo.20094
Empria loktini Ermolenko 1971
Empria loktini Ermolenko, 1971 Empria loktini Ermolenko, 1971: 22 –23. Type locality: Novoaleksandrovsk [Sakhalin Oblast, Russia]. Holotype Ψ, (SIZ) [examined]. Host plants. Unknown, but the adults have been collected from Filipendula kamtschatica (see Ermolenko 1971) and at least some of the Japanese specimens have been collected from the localities where this plant species is common (A. Shinohara, personal communication). Distribution. East - Palaearctic. The verified country records are: Japan (Hokkaido) and Russia (Sakhalin Oblast). Key characters. Posterior margin of pronotum, metatibia in basal 1 / 3, and paired patches on abdominal terga 2–3 or 2–4 (5) unpigmented; tegulae unpigmented (females) or black (males). Ratio of flagellum length to head breadth in females 1.8–1.9, in males 2.3–2.4, and eye length to head length in females 1.2–1.7, in males 1.5 –2.0. Number of serrulae 13–14 (15), triangular in shape (Fig. 39). Penis valve with relatively large apical tooth and apical part of valvular duct going clearly further from dorsal rim of valvura; valviceps with medium sized lobe, forming a notch between valviceps and valvura (Fig. 55). Notes. The paratypes 1 Ψ 2 ɗ (1 ɗ labelled as allotype) were also studied.Published as part of Prous, Marko, Heidemaa, Mikk & Soon, Villu, 2011, Empria longicornis species group: taxonomic revision with notes on phylogeny and ecology (Hymenoptera, Tenthredinidae), pp. 1-39 in Zootaxa 2756 on page 24, DOI: 10.5281/zenodo.20094
Empria alpina Benson 1938
Empria alpina Benson, 1938 Empria alpina Benson, 1938: 190 –191. Type locality: Scotland, Perthshire, Breadalbane Mountains, Crags above Lochan à Lairige [UK]. Holotype Ψ, BMNH [not examined, see Notes]. Host plants. Dryas octopetala (Veli Vikberg, personal communication). Distribution. Holarctic (no Nearctic specimens were studied). The verified country records are: Finland, Germany, Great Britain, Italy, and Switzerland. Key characters. Paired patches on abdominal terga 2–6 (7) unpigmented; tegulae, posterior margin of pronotum, and metatibia mostly black. Ratio of flagellum length to head breadth in females 1.6–2.1, in males 2.7–3.5, and eye length to head length 1.3–1.5. Number of serrulae 14, conspicuously papilliform (Fig. 36). Valviceps with a long basal lobe, forming very deep notch between valviceps and valvura (Fig. 53). Notes. Taxonomic identity is evident from the original description (e.g. fig 7 a in Benson, 1938), but confirmed also by studying three paratypes (1 Ψ and 1 ɗ with same collecting data as holotype).Published as part of Prous, Marko, Heidemaa, Mikk & Soon, Villu, 2011, Empria longicornis species group: taxonomic revision with notes on phylogeny and ecology (Hymenoptera, Tenthredinidae), pp. 1-39 in Zootaxa 2756 on page 22, DOI: 10.5281/zenodo.20094
FIGURES 30–31 in Studies toward a World Catalog of Symphyta (Hymenoptera) 2254
FIGURES 30–31. Head in frontal view, photos by A. Taeger: 30, Tenthredo sauteri (Rohwer, 1916); 31, T. sporadipunctata Malaise, 1945.Published as part of Blank, Stephan M., Taeger, Andreas, Liston, Andrew D., Smith, David R., Rasnitsyn, Alexandr P., Shinohara, Akihiko, Heidemaa, Mikk & Viitasaari, Matti, 2009, Studies toward a World Catalog of Symphyta (Hymenoptera) 2254, pp. 1-96 in Zootaxa 2254 (1) on page 68, DOI: 10.11646/zootaxa.2254.1.1, http://zenodo.org/record/531851
FIGURE 11. Tenthredo collaris Donovan, 1808 in Studies toward a World Catalog of Symphyta (Hymenoptera) 2254
FIGURE 11. Tenthredo collaris Donovan, 1808 [= Dolerus haematodes (Schrank)], fig. 1 from plate 441 in Donovan (1808).Published as part of Blank, Stephan M., Taeger, Andreas, Liston, Andrew D., Smith, David R., Rasnitsyn, Alexandr P., Shinohara, Akihiko, Heidemaa, Mikk & Viitasaari, Matti, 2009, Studies toward a World Catalog of Symphyta (Hymenoptera) 2254, pp. 1-96 in Zootaxa 2254 (1) on page 23, DOI: 10.11646/zootaxa.2254.1.1, http://zenodo.org/record/531851
FIGURE 11. Tenthredo collaris Donovan, 1808 in Studies toward a World Catalog of Symphyta (Hymenoptera) 2254
FIGURE 11. Tenthredo collaris Donovan, 1808 [= Dolerus haematodes (Schrank)], fig. 1 from plate 441 in Donovan (1808).Published as part of Blank, Stephan M., Taeger, Andreas, Liston, Andrew D., Smith, David R., Rasnitsyn, Alexandr P., Shinohara, Akihiko, Heidemaa, Mikk & Viitasaari, Matti, 2009, Studies toward a World Catalog of Symphyta (Hymenoptera) 2254, pp. 1-96 in Zootaxa 2254 (1) on page 23, DOI: 10.11646/zootaxa.2254.1.1, http://zenodo.org/record/531851
Empria basalis Lindqvist 1968
Empria basalis Lindqvist, 1968 Empria basalis Lindqvist, 1968: 26 –27. Type locality: Kallvik [from label], vicinity of Helsinki ”Umgegend von Helsingfors” [original description]. Holotype Ψ, ZMH [examined]. Host plants. Geum rivale and possibly G. urbanum. Distribution. West-Palaearctic. The verified country records are: Czech Republic, Denmark, Estonia, Finland, France, Germany, Hungary, Russia (Leningrad Oblast and Ulyanovsk Oblast), Slovakia, Sweden, Ukraine. Key characters. Posterior margin of pronotum and paired patches on abdominal terga 2–5 or 2–6 unpigmented; tegulae and metatibia in basal 1 / 3 black or partly unpigmented. Ratio of flagellum length to head breadth in females mostly 1.7 –2.0, in males 2.1–2.7, and eye length to head length mostly 1.6–2.2. Number of serrulae 15– 16 (rarely 14 or 17), from which 4–5 basal ones without denticles; triangular in shape (Fig. 44). Valviceps with medium sized lobe, forming a notch between valviceps and valvura (Fig. 52). Notes. The paratype male (labelled as allotype) was also studied.Published as part of Prous, Marko, Heidemaa, Mikk & Soon, Villu, 2011, Empria longicornis species group: taxonomic revision with notes on phylogeny and ecology (Hymenoptera, Tenthredinidae), pp. 1-39 in Zootaxa 2756 on page 22, DOI: 10.5281/zenodo.20094
Empria mongolica Konow 1895
Empria mongolica (Konow, 1895) Poecilosoma mongolica Konow, 1895: 75. Type locality: northern Mongolia “ Mongolia bor[ealis]”. Lectotype (here designated) Ψ: “N. Mongolei Leder 92 ” [white, printed]; “ Poecilosoma Mongolica Knw. Mongol. bor.” [white, framed, handwritten]; “ Typus ” [red, printed]; “ Holotypus ” (Sic!) [red, printed]; “Coll. Konow” [white, printed]; “Conde revid. 1937 ” [white, handwritten]; “GBIF-GISHym 3782 ” [white, printed]; “ LECTOTYPUS Poecilosoma mongolica KONOW, 1895 Ψ M.Heidemaa & M.Prous des.” [red, printed]; “ Empria mongolica (Konow, 1895) det. M.Prous 2008 ” [white, printed]; SDEI (see Notes). Host plants. Dasiphora fruticosa (see Verzhutskii 1981). Distribution. East - Palaearctic. The verified country records are: Mongolia, Russia (Kamchatka Krai, Irkutsk Oblast: Verzhutskii 1981). The record from arctic Norway (Kiaer 1898) is incorrect due to original misidentification of E. fletcheri specimens (determined by E. Lindqvist in 1948) as E. mongolica. Key characters. Posterior margin of pronotum, small basal part or basal 1 / 3 of metatibia, and paired patches on abdominal terga 2–6 (7) unpigmented; tegulae largely unpigmented (females) or black (males). Ratio of flagellum length to head breadth in females 1.7 –2.0, in males 2.7–3.2, and eye length to head length in females 1.5–1.7, in males 1.7–2.2. Number of serrulae 14–15, conspicuously papilliform shaped (Fig. 37). Valviceps with a long basal lobe, forming relatively deep notch between valviceps and valvura (Fig. 54). Notes. Oehlke & Wudowenz (1984) assumed that the only female of Poecilosoma mongolica Konow, 1895 found in Konow’s collection is the holotype but neither holotype was designated nor explicitly stated in the original description that there was only one specimen (thus a syntype).Published as part of Prous, Marko, Heidemaa, Mikk & Soon, Villu, 2011, Empria longicornis species group: taxonomic revision with notes on phylogeny and ecology (Hymenoptera, Tenthredinidae), pp. 1-39 in Zootaxa 2756 on page 25, DOI: 10.5281/zenodo.20094
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