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    Pseudoplandriina HANLEY 2002

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    SUBTRIBE PSEUDOPLANDRIINA HANLEY, 2002 A <p>(FIGS 26,27)</p> <p>Pseudoplandriina Hanley, 2002a:301. Hanley, in press b.</p> <p> <i>Type-genus:</i> <i>Pseudoplandria</i> Fenyes, 1921.</p> <p> <i>Diagnosis:</i> In addition to the characters of the tribe, this subtribe is characterized by a 5-5-5 tarsal formula; a very long ligula, length to width ratio greater than 2.0; mesosternum with a distinct carina. Males typically have a more or less broad carina on the medioapical region of the elytra.</p> <p> <i>Comments:</i> This subtribe consists of one genus, <i>Pseudoplandria</i>, with 71 described species primarily distributed throughout the Oriental Region (one species is described from Japan). Most species are known only from type specimens. The habits and food preferences of members of this lineage are completely unknown.</p>Published as part of <i>Hanley, Rodney S., 2003, Generic revision of the staphylinid beetle tribe Hoplandriini (Insecta: Coleoptera: Staphylinidae: Aleocharinae), pp. 83-140 in Zoological Journal of the Linnean Society 138 (1)</i> on page 130, DOI: 10.1046/j.1096-3642.2003.00060.x, <a href="http://zenodo.org/record/5437792">http://zenodo.org/record/5437792</a&gt

    Platandriina HANLEY 2002

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    SUBTRIBE PLATANDRIINA HANLEY, 2002 A <p>(FIGS 16–24)</p> <p>Platandriina Hanley, 2002a:301. Hanley, in press b.</p> <p> <i>Type-genus:</i> <i>Platandria</i> Casey, 1893.</p> <p> <i>Diagnosis:</i> In addition to the characters of the tribe, this subtribe is characterized by 4-5-5 or 4-4-5 tarsal formula; narrowly separated mesocoxae; ligula short, with a relatively deep fork. Secondary sexual characteristics often present in males as small carinae or denticles on abdominal terga IV and VIII, and rarely VII (never the elytra).</p> <p> <i>Comments:</i> This subtribe currently consists of seven genera and 26 described species distributed in the Afrotropical, Nearctic, Neotropical, and Oriental regions, with one species of <i>Omoplandria</i> described from Japan. Based on limited label data from few genera, I hypothesize that members of this lineage occur in association with flowers where they are most likely feeding on nectar or pollen.</p>Published as part of <i>Hanley, Rodney S., 2003, Generic revision of the staphylinid beetle tribe Hoplandriini (Insecta: Coleoptera: Staphylinidae: Aleocharinae), pp. 83-140 in Zoological Journal of the Linnean Society 138 (1)</i> on page 112, DOI: 10.1046/j.1096-3642.2003.00060.x, <a href="http://zenodo.org/record/5437792">http://zenodo.org/record/5437792</a&gt

    Ligulata HANLEY & ASHE 1998

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    GENUS LIGULATA HANLEY & ASHE, 1998 (FIGS 19,20) Ligulata Hanley & Ashe, 1998:184. Hanley, 2002a:301. Hanley, in press b. Type species: Ligulata hansoni Hanley & Ashe (1998) by monotypy. Diagnosis: This genus is distinguishable from the other genera of Platandriina by the following combination of characters: body average in size, 2.4–3.4 mm; eyes average in size, 0.4–0.6 times length of head; apex of galea densely covered with rows of very long hairs, giving appearance of flowing hairs; labial palps with three distinct articles; mesosternum without a medial carina; mesocoxal cavities widely separated, completely separated by meso- and metasternal processes; empodial bristle of hind legs distinctly longer than tarsal claws; apices of abdominal terga with typical macrosetae, not large, heavy, black setae; abdominal tergum IX with a distinct crescentshaped setal pattern; males without secondary sexual features. Description: In agreement with Platandria description, except for the following characters. [1] Lengths of adults 2.4–3.4 mm. [2] Body (Fig. 19A) robust, elongate in dorsal outline, more or less tapering from broad base of pronotum and elytra towards apex of abdomen, [3] surface more or less glossy, evenly pubescent with [4] faintly reticulate microsculpturing. Head: (Fig. 19A) [10] Distinctly broader than long. [11] Eyes moderate in size, 0.4–0.6 times length of head. [13] Infraorbital carina strongly developed and complete. Antenna with [15] articles 1–3 elongate, 2– 8 times longer than wide; [16] segment 4 slightly elongate, 1.2 times longer than wide; [17] articles 5–10 slightly elongate to quadrate; [18] coeloconical sensilla absent. Mouthparts: Labrum (Fig. 19B) with epipharyngeal area with [21] numerous, moderately large pores present between longitudinal sensory field and lateral sclerotized areas (Fig. 19C). Mandible (Fig. 19D,E) [22] asymmetrical with right with a distinct median tooth, left without median tooth; [23] apex more or less acute and curved adorally; [24] condylar molar patch moderate in size, width less than 1/5 of basal mandibular width, [25] composed of very small denticles, [26] densely arranged in irregular transverse rows; [27] subcondylar molar patch moderate in size, width about 1/4 of basal mandibular width; dorsobasal ‘velvety patch’ [28] very large, width about 2/3 of width of mandibular base, [29] modified into 4 transverse rows of moderately large teeth. Maxilla (Fig. 19F) with [31] lacinia about as long as galea, [32] lacinia acute apically, [33] teeth on adoral margin short to relatively long, about 2–15 times longer than wide, with [36] numerous setae and 2 large spinose setae on dorsal surface; [37] galea broad, width subequal to lacinea at widest area, and obliquely truncate apically, [38] membranous in apical 1/2–1/3, [39] densely covered with rows of very long, fine hairs. Labium (Fig. 19G) with ligula [45] elongate, as long or longer than labial palpi 1 + 2, [46] broadened apically with [47] rounded apex, [48] with numerous long hairs irregularly arranged; [49] two long medial setae of prementum present, longer than ligula; [53] median pseudopore field narrow and linear, with numerous pseudopores; [55] hypoglossal lobes (Fig. 19H) very long, typically reaching to midpoint of ligula, [56] with long, about 3.0–5.5 times longer than width of lobe, comb-like internally curved setae, [57] along entire length of adoral surface. Labial palpi (Fig. 19G) [58] elongate, overall length 5.5–7.0 times longer than greatest width, [60] with distal pore field composed of two small, weakly defined pores; [61] segment 1 about 3.7 times longer than segment 2, [62] segment 3 about 2.9 times longer than segment 2. Mentum with [67] many sensory pores more or less uniformly distributed in middle 2/3, primarily near midline. Thorax: Pronotum (Fig. 19A) [68] transverse, [69] about 1.7 times wider than long, [70] strongly convex, about as wide at base as base of elytra. Setae [72] directed primarily posteriorly. Elytra [74] moderately broad, slightly wider apically than basally; [75] apicolateral angles slightly sinuate; [76] elytra together about 1.9–2.2 times as wide as long; microsetae [78] uniformly distributed. Mesocoxal cavities (Fig. 20A) [81] widely separated by meso- and metasternal processes, by about 1/4 length of coxal cavities. Mesosternal process [82] longer than metasternal process, extended to basal 5/6 of coxal cavities; [83] meso- and metasternal processes in contact, isthmus absent; [84] mesosternal process slightly rounded to flattened. Metasternum [85] shorter than width of mesocoxae. Legs with [93] empodial bristle longer than tarsal claws; [94] segment 1 of hind tarsus about 1.3 times length of segment 2. Abdomen: (Fig. 19A,B) [97] Narrowly fusiform, tapering apically to broadly pointed apex. Secondary sexual characteristics: [102,103,104,105, 106,107,108,109] Absent. Aedeagus: (Fig. 20C) Bulb of median lobe [110] more or less elongate, length slightly shorter to subequal to length of tube, [111] with very small ventral projection; [113] parameres (Fig. 20D) with apical lobe of paramerite relatively short and narrow, not extended beyond velum; [115] paramerite anterior margin flattened to slightly concave; condylite [117] generally shorter in length to apex of paramerite. Spermatheca: (Fig. 20E) [118] L-shaped; neck, [120] bent at about 90∞ angle; and tube, [121] membranous, loosely curved. Habitat: Unknown. All specimens of this genus were collected using Malaise traps at 1600 m elevation near Zurqui de Moravia, Costa Rica. Comments: As mentioned in Hanley & Ashe (1998), specimens of Ligulata superficially resemble those of Platandria. However they differ from this latter genus in structure of the maxilla, the long setose ligula and the broadly separated mesocoxae (see Génier & Klimaszewski, 1986 for a discussion of characters of Platandria). Ligulata is composed of one species, L. hansoni Hanley and Ashe. Ligulata appears to be closely related to the genus Bessoglossa with which it shares the following possible synapomorphies: ligula long with numerous long setae at apex; and hypoglossal lobes long, comb-like with long, internally curved setae. Unfortunately, this hypothesis of relationship remains tenuous because I have not been able to study specimens of Bessoglossa. The genus was originally described by Pace (1986) from one specimen of the type species, Bessoglossa peruviana Pace. The specimen was noted in the original publication to have been deposited in the personal collection of Dr Herbert Franz. Unfortunately, repeated attempts to borrow this specimen ended in failure.Published as part of Hanley, Rodney S., 2003, Generic revision of the staphylinid beetle tribe Hoplandriini (Insecta: Coleoptera: Staphylinidae: Aleocharinae), pp. 83-140 in Zoological Journal of the Linnean Society 138 (1) on pages 119-122, DOI: 10.1046/j.1096-3642.2003.00060.x, http://zenodo.org/record/543779

    Succinctonotum Gordon and Hanley 2017, new genus

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    <i>Succinctonotum</i> Gordon and Hanley, new genus <p> <b>Description.</b> Cephaloscymnini with body form short, widely oval, narrowed from apex of pronotum to apex of abdomen. Frons narrow, widened from vertex to apex, widest point nearly as wide as eye; apex of frons not extended beyond antennal insertion; male clypeus and anterior portion of frons pale yellow or yellowish white, not densely pubescent (Fig. 308); female head without maculation. Apical maxillary palpomere slender, narrowed from middle to apex as in <i>Neaporia</i>. Pronotum short, about two times as long as scutellum, with anterior margin deeply excavated for reception of head, with oblique surface groove as in <i>Neaporia,</i> strongly projected forward laterally to midpoint of eye (Fig. 308). Prothoracic hypomeron without fossa. Epipleuron narrow, flat. Male pro–, meso– and metasterna mostly flat except weak depression between pro– and mesosterna; prosternum short, wide, without lateral carina, apex without modified setae. Male metasternum without pit medially adjacent to metepisternum. Tarsal claw without basal angulation. Apex of male 5th ventrite arcuate.</p> <p> <b>Remarks.</b> This genus is thus far known from two specimens. It is recognized by the short body; pronotum only slightly longer than scutellum medially; narrow, elongate head; and flat, unmodified prosternal process. The male is described below and female genitalia of the other specimen are also described, but the female is not designated as a type because it cannot be determined with certainty that these specimens are the same species. <i>Succinctonotum</i> is similar to <i>Neaporia</i>, sharing the same pronotal groove and lack of prosternal carinae.</p> <p> <b>Etymology.</b> The genus name refers to the short pronotum of the type species; gender feminine.</p>Published as part of <i>Gordon, Robert D. & Hanley, Guy A., 2017, South American Coccinellidae (Coleoptera), Part XVII: systematic revision of Western Hemisphere Cephaloscymnini (Coccinellinae) with description of a cryptic new genus and species of Coccidulini (Coccinellinae), pp. 1-158 in Insecta Mundi 2017 (601)</i> on page 50, DOI: <a href="http://zenodo.org/record/5170031">10.5281/zenodo.5170031</a&gt

    Asterophilia Hanley 1989

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    Asterophilia Hanley, 1989 culcitae Britayev & Fauchald, 2005: Nha Trang Bay, Vietnam. ZMMSU N4512, ZMMSU N4513-4518Published as part of Salazar-Vallejo, Sergio I., Carrera-Parra, Luis F., Muir, Alexander I., León-González, Jesús Angel De, Piotrowski, Christina & Sato, Masanori, 2014, Polychaete species (Annelida) described from the Philippine and China Seas, pp. 1-68 in Zootaxa 3842 (1) on page 31, DOI: 10.11646/zootaxa.3842.1.1, http://zenodo.org/record/492848

    Verrucapelma Hanley & Burke 1991

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    Verrucapelma Hanley & Burke, 1991 conifera Barnich, Fiege & Sun, 2004: Hainan Island, China. IOCAS type not located, SMF13422 –13424Published as part of Salazar-Vallejo, Sergio I., Carrera-Parra, Luis F., Muir, Alexander I., León-González, Jesús Angel De, Piotrowski, Christina & Sato, Masanori, 2014, Polychaete species (Annelida) described from the Philippine and China Seas, pp. 1-68 in Zootaxa 3842 (1) on page 34, DOI: 10.11646/zootaxa.3842.1.1, http://zenodo.org/record/492848

    The impacts of elicitation context on stated preferences for agricultural landscapes

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    Funded by UK Research Council, ESRC, and NERC.Statements of willingness to pay (WTP) have been shown to be dependent upon the framing of the hypothetical market. In this paper we investigate the effects of variations in the timing and location of choice experiment questions concerned with conservation of a UK national park, as research involving measurement of psychological well-being suggests potential differences for the same individual dependent upon when and where preferences are elicited. We apply the choice experiment technique to the valuation of changes in upland agricultural and semi-natural landscapes in the Peak District National Park in the UK, to investigate whether timing and location of elicitation (context) affects the value associated with changes in ecosystem services under different management regimes. Four treatments are employed - using the same sample of individuals answering the same choice scenarios - to measure WTP ex-ante (off site), in situ (on site), and ex-post at two different time intervals (off site). We show that our on-site (in situ) treatment generates very different estimates of preferences than any of the off-site treatments. That stated preferences associated with environmental goods are so context dependent may have implications for the use of stated preferences in policy analysis in terms of identifying how environmental policy is funded and the divergence in value attributed to sampling different populations.Peer reviewe

    Ponaria Gordon and Hanley 2017, new genus

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    <i>Ponaria</i> Gordon and Hanley, new genus <p> <b>Type species.</b> <i>Neaporia caerulea</i> Gorham, by present designation.</p> <p> <b>Description.</b> Coccidulini with body form oval. Antenna with 9 articles. Frons narrow, equal in width from vertex to clypeus, inner margins of eyes parallel, narrower than eye, frons short, apex of frons extended beyond antennal insertion by less than width of basal antennal article; frontal extension onto eye long, extended nearly completely across eye; male clypeus and frons immaculate, densely pubescent (Fig. 715); female head without dense pubescence. Pronotum long, with anterior margin moderately excavated for reception of head, weakly projected forward laterally to about apical 4/5 of eye, pronotal surface without surface groove. Epipleuron wide, flat. Male pro–, meso– and metasterna flat, not medially depressed; prosternum weakly expanded to conceal mouthparts, prosternal process with apex truncate, process narrow, short, with coarse, sparse punctures, lateral carina present on each side adjacent to procoxa; male prosternal process without modified setae. Male metasternum without pit medially adjacent to metepisternum. Tarsal claw with slight basal angulation. Apex of male 5th venter barely perceptibly truncate. Female genitalia with long, wide infundulum; spermathecal capsule with large, round ramus, narrowed from ramus to nearly acute apex of cornu.</p> <p> <b>Remarks.</b> <i>Ponaria</i> bears a strong, superficial resemblance to members of Cephaloscymnini with whom it is most likely to be confused. Similarities include the large, elongate eye; narrow frons densely pubescent in male; and antenna with 9 articles. Characters that remove it from Cephaloscymnini are lateral extension of frons nearly completely dividing eye; gena not expanded anterior to eye to form shelf for reception of antenna; and female genitalia with ramus of spermathecal capsule large, round, remainder of capsule slender, tapered to almost acute apex of cornu. Mimicry is possible here given the strong similarity of this genus to Cephaloscymnini genera.</p> <p>We place this genus in Coccidulini, where it would belong in the formerly recognized Scymini. It is distinguished from other known Coccidulini by a combination of large, elongate eye; and narrow frons with lateral extension almost completely dividing eye.</p> <p> <b>Etymology.</b> The genus name is an arbitrary rearrangement of several letters in <i>Neaporia</i> Gorham; gender feminine.</p>Published as part of <i>Gordon, Robert D. & Hanley, Guy A., 2017, South American Coccinellidae (Coleoptera), Part XVII: systematic revision of Western Hemisphere Cephaloscymnini (Coccinellinae) with description of a cryptic new genus and species of Coccidulini (Coccinellinae), pp. 1-158 in Insecta Mundi 2017 (601)</i> on pages 107-108, DOI: <a href="http://zenodo.org/record/5170031">10.5281/zenodo.5170031</a&gt

    Prodilis kristy Gordon and Hanley 2017, new species

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    22. <i>Prodilis kristy</i> Gordon and Hanley, new species <p> <b>Description. Male</b> holotype. Length 2.4 mm, width 1.9 mm; body oval, elytron with side rounded, wider than pronotal base, widest at middle of elytra. Dorsal surface shiny. Color blue (Fig. 410); head blue with anterior ½ yellow, base of yellow area tridentate, pubescent; antenna, legs yellow; epipleuron brownish red; mouthparts yellow except apical maxillary palpomere brown; abdomen medially brown with lateral, apical 1/4 yellow. Head punctures large, separated by less than a diameter; pronotal punctures smaller than on head, separated by les than a diameter; elytral punctures larger than on head, separated by a diameter or less; prosternal punctures sparse, nearly absent; mesosternal punctures large, separated by less than a diameter; metasternal punctures small, nearly absent except present on lateral, anterior 1/4; abdomen with punctures on ventrites 1–2 small, nearly absent, punctures on remaining ventrites smaller, separated by about a diameter. Head with frons widened from vertex to clypeus, as wide as 1.4 times eye measured at vertex; eye canthus short; apical maxillary palpomere short, widened from base to apex. Pronotum widest at middle, reflexed lateral margin wide, equal in width from base to apex. Epipleuron flat, wide, slightly descending externally, as wide as pronotal hypomeron, without depressions. Prosternum wider than long, about as long as mesosternum, base weakly arcuate, lateral carina slender, extended to base of procoxa. Postcoxal line on ventrite 1 long, angulate, extended to apex of ventrite (Fig, 441). Apex of ventrite 5 with wide, short emargination. Basal lobe of genitalia shorter than paramere, slender, widened from base to emarginate apex, each on each side of emargination widened, round; paramere slender basally, widened from base to rounded apex, without marginal serrations (Fig. 443, 444); sipho robust (Fig. 445).</p> <p> <b>Female.</b> Head blue, not pubescent (Fig. 442). Female genitalia with spermathecal capsule long slender, slightly narrowed at middle, cornu apically rounded (Fig. 446).</p> <p> <b>Variation</b>. None observed.</p> <p> <b>Type material.</b> Holotype male; BRAZIL: Am, (Amazonas), Reserve Ducki, 26km NE Manaus, Hurtado, L.C. G., <i>Micropholis guyanensis</i> 15.x1995, Tree No. 50a, Tray NO. 5, BMNH(E) 2003–84. (BMNH) Paratypes 5, same data as holotype except Tree No. 129 Tray No. 10 <i>Eschweilera pseudodecolorans</i> 25.vi.1996, Tree No. 166 Tray No. 5 <i>Licania micrantha</i> 27.vi.1996, Tree No. 166 Tray No. 8 <i>Licania micrantha</i> 27.vi.1996, Tree No. 147 Tray No. <i>Corythophora alta</i> 22,vi,1996, Tree No. 155, Tray No. 7 <i>Micropholis guyanensis</i> 07.ii.1996. (BMNH).</p> <p> <b>Remarks.</b> This species is characterized in males by head maculation and genital structure.</p>Published as part of <i>Gordon, Robert D. & Hanley, Guy A., 2017, South American Coccinellidae (Coleoptera), Part XVII: systematic revision of Western Hemisphere Cephaloscymnini (Coccinellinae) with description of a cryptic new genus and species of Coccidulini (Coccinellinae), pp. 1-158 in Insecta Mundi 2017 (601)</i> on pages 72-73, DOI: <a href="http://zenodo.org/record/5170031">10.5281/zenodo.5170031</a&gt

    Prodilis kristi Gordon and Hanley 2017, new species

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    36. <i>Prodilis kristi</i> Gordon and Hanley, new species <p> <b>Description. Male</b> holotype. Length 2.5 mm, width 1.7 mm; body oval, elytron with side straight, wider than pronotal base, widest at middle of elytra. Dorsal surface entirely shiny without trace of microsculpture. Color variable; head with base of frons and vertex black, remainder of head yellow (Fig. 521); pronotum medially black with lateral 1/3 yellow; elytron blue (Fig. 519); antenna, mouthparts, legs reddish yellow; ventral surface including epipleuron dark reddish brown; abdomen with basal ventrite brown, ventrite 2 yellow with median 2/3 brown, remainder of abdomen brownish yellow. Head punctures small, separated by a diameter or less; pronotal punctures larger than on head, separated by less than a diameter; elytral punctures larger than on pronotum, separated by less than a diameter; prosternal, mesosternal punctures large, separated by 1 to 2 times a diameter; metasternal punctures as large as on mesosternum separated by less than a diameter on basal and lateral borders, small, nearly absent medially; abdomen with punctures on ventrites 1–3 large, separated by a diameter or less, punctures on remaining ventrites smaller, separated by about a diameter. Head with frons weakly widened from vertex to clypeus, 1.4 times as wide as eye measured at vertex; eye canthus short; apical maxillary palpomere short, widened from base to apex. Pronotum widest at middle, reflexed lateral margin narrow, equal in width from base to apex. Epipleuron flat, slightly widened in basal ½, as wide as pronotal hypomeron, with feeble depressions for reception of femoral apices. Prosternum longer than wide, longer than mesosternum, anteriorly produced, base broadly, weakly emarginate medially, lateral carina wide, extended anterior to apex of coxa. Postcoxal line on ventrite 1 long, slightly angulate, extended 2/3 distance to ventrite apex (Fig. 520). Apex of ventrite 5 arcuate. Genitalia with basal lobe longer than paramere, wide, widest medially, lateral margin curved from base to abruptly curved apex, apex slightly emarginate; paramere weakly curved, slender, equal in width to rounded apex, without marginal serrations (Fig. 522, 523); sipho long, slender, unmodified, broken in image (Fig. 524).</p> <p> <b>Female.</b> Unknown.</p> <p> <b>Variation</b>. None observed.</p> <p> <b>Type material.</b> Holotype male; BRAZIL: CORCOVADO, Rio Guanabara, X. I.1960, Seabra e Alvarenga. (DZUP).</p> <p> <b>Other specimen</b>. A single male BMNH specimen labeled ”2290/Fry RioJano./Fry Coll. 1905.100. is probably also this species. It differs from the holotype by male genitalia with basal lobe slightly wider with lateral margins more arcuate. In all other regards, including densely punctured dorsal surface and prosternal base medially emarginate, it matches the holotype of <i>P</i>. <i>kristi</i>.</p> <p> <b>Remarks.</b> <i>Prodilis kristi</i> is distinguished by its large size, blue elytron, lateral pronotal margin widely yellow, densely punctured dorsal surface, and prosternal base medially emarginate. This species and <i>P</i>. <i>sandy</i> may be conspecific, see remarks under the latter species.</p>Published as part of <i>Gordon, Robert D. & Hanley, Guy A., 2017, South American Coccinellidae (Coleoptera), Part XVII: systematic revision of Western Hemisphere Cephaloscymnini (Coccinellinae) with description of a cryptic new genus and species of Coccidulini (Coccinellinae), pp. 1-158 in Insecta Mundi 2017 (601)</i> on pages 82-83, DOI: <a href="http://zenodo.org/record/5170031">10.5281/zenodo.5170031</a&gt
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