267,433 research outputs found

    Modelling patient length of stay in public hospitals in Mexico

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    This thesis is concerned with the modelling of patient length of stay in public hospitals in Mexico. Patient length of stay is the most commonly worldwide employed outcome measure for hospital resource consumption and performance monitoring. Most of the hospitals around the world use average length of stay as starting point for resource planning. However average estimates frequently gives non-accurate results due to the high variability of the length of stay data. The reason for such high variability may be attributable to the diversity in the patient population and the environment where the patient is treated.Through a systematic review of the literature on methods and models in the field of calculating and predicting patient length of stay, this research highlights the areas of opportunity and research gap from previous studies and practices, and proposes the use of finite mixture models to approximate the distribution of length of stay. Also, these models are proposed as the foundation of more sophisticated models designed to include the internal and external factors associated with LoS. In this context, the thesis proposes three different approaches to explore such factors: individual-based approach, group-based approach and multilevel group-based approach. These interrelated approaches allow a better understanding of the diversity in the patient population and enable length of stay predictions for individual patients, and for cohorts of patients within and between hospitals. In addition, this research is built and evaluated using data from all types of patients treated at two public hospitals operating in Mexico. It is the consideration of the full case-mix of these healthcare facilities that gives this research its unique nature

    Klinckowstroemia montanoi Villegas-Guzman, Perez & Reyes-Castillo 2009

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    Klinckowstroemia montanoi Villegas-Guzman, Pérez & Reyes-Castillo, 2009: 11 — Holotype female (CNAC, 006538), from Verres aff. intermedius, oak forest, 8 km NW San Miguel Coatlán, Federal highway Cuixtla-San Miguel Coatlán, Oaxaca, Mexico (16°15.064'N, 96°41.629'W, 2170 m).Published as part of LIU, DONG, YI, TIAN-CI, XU, YUN & ZHANG, ZHI-QIANG, 2013, new mite species described during 2007 to 2012 3663, pp. 1-102 in Zootaxa 3663 (1) on page 26, DOI: 10.11646/zootaxa.3663.1.1, http://zenodo.org/record/563059

    Klinckowstroemia stilla Villegas-Guzman, Reyes-Castillo & Perez 2011

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    Klinckowstroemia stilla Villegas-Guzman, Reyes-Castillo & Pérez, 2011: 28 — Holotype female (CNAC, 006739), from Proculus goryi, mountain rain forest, Ejido Benito Juárez, Reserva de la Biosfera Volcán Tacan, Chiapas, Mexico (15°05. 966'N, 92°08. 600'W, 1687 m).Published as part of LIU, DONG, YI, TIAN-CI, XU, YUN & ZHANG, ZHI-QIANG, 2013, new mite species described during 2007 to 2012 3663, pp. 1-102 in Zootaxa 3663 (1) on page 27, DOI: 10.11646/zootaxa.3663.1.1, http://zenodo.org/record/563059

    Klinckowstroemia cristinae Villegas-Guzman, Perez & Reyes-Castillo 2009

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    Klinckowstroemia cristinae Villegas-Guzman, Pérez & Reyes-Castillo, 2009: 40 — Holotype female (CNAC, 006649), from Odontotaenius zodiacus, pine-oak forest, 2 km Federal highway Crucero-Tianguistengo, Hidalgo, Mexico (20°40.413'N, 98°40.255'W, 2080 m).Published as part of LIU, DONG, YI, TIAN-CI, XU, YUN & ZHANG, ZHI-QIANG, 2013, new mite species described during 2007 to 2012 3663, pp. 1-102 in Zootaxa 3663 (1) on page 26, DOI: 10.11646/zootaxa.3663.1.1, http://zenodo.org/record/563059

    Klinckowstroemia ballesterosi Villegas-Guzman, Perez & Reyes-Castillo 2009

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    Klinckowstroemia ballesterosi Villegas-Guzman, Pérez & Reyes-Castillo, 2009: 39 — Holotype female (CNAC, 006640), from Odontotaenius zodiacus, pine-oak forest, Municipality of Soledad Atzompa, Soledad Atzompa, Veracruz, Mexico (18°44.712'N, 97°40.967'W, 2125 m).Published as part of LIU, DONG, YI, TIAN-CI, XU, YUN & ZHANG, ZHI-QIANG, 2013, new mite species described during 2007 to 2012 3663, pp. 1-102 in Zootaxa 3663 (1) on page 26, DOI: 10.11646/zootaxa.3663.1.1, http://zenodo.org/record/563059

    Pacifigorgia sculpta Breedy & Guzman, 2004, new species

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    Pacifigorgia sculpta, new species (Figs. 1 G–H, 5 A–C) Material examined. Holotype: UCR 1497, Islote Frailes, Península de Azuero, 10–30 m, H.M. Guzman, 9 December 2001. Paratypes: MCZ 57053, Islote Frailes, 10–30 m, H.M. Guzman, 9 December 2001; STRI 389–390, Isla Jicarita, Gulf of Chiriquí, 20 m, H.M. Guzman, 8 August 2002; STRI 410–412, Isla Seca Grande, Gulf of Chiriquí, 20 m, H.M. Guzman and O. Breedy, 26 August 2002; STRI 454, Isla Roncadores, Gulf of Chiriquí, 10–20 m, H.M. Guzman and O. Breedy, 30 August 2002; STRI 476, 482, 497, Bajo Foul, Península de Azuero, 15 m, H.M. Guzman, 11 April 2003; STRI 602, Islote Frailes, 20 m, H.M. Guzman, 1 May 2003; STRI 628, Roca Catedral, 5–15 m, H.M. Guzman, 3 May 2003; STRI 650, Bajo Brincanco, Gulf of Chiriquí, 10–30 m, H.M. Guzman, 5 May 2003; STRI 718, 721–722, 729 – 731, 734, Bajo Trollope, Gulf of Panama, 10–20 m, H.M. Guzman, 6 August 2003; UCR 1037, 1042, Islote Frailes, 10–20 m, H.M. Guzman, 6 August 2003; UCR 1171, 1173, 1175, 1177, 1179, 1181, 1183, 1505, Roca Niagara, Gulf of Panama, 10–20 m, H.M. Guzman, 13 December 2001; UCR 1361 –1365, 1506, Piedra Hacha, 20–30 m, H.M. Guzman, 22 April 2002; UCR 1498, Isla Jicarita, Gulf of Chiriquí, 20–30 m, H.M. Guzman, 19 April 2002; UCR 1499, 1501, 1508, Islote Frailes, 10–30 m, H.M. Guzman, 12 December 2001. Description. Colonies wider than high, up to 120 mm in height and 200 mm in width. Most of the colonies are composed of a single fan, but some have two or three secondary fans that radiate from different parts of the main fan and grow parallel. Colour when preserved or alive is dark orange or reddish brown with lighter hues at the tips, and light ochre when dry. Colonies have a large holdfasts, and fans grow directly from this or sprout from short stems (up to 7 mm in diameter). Network is irregular. Meshes are very open (about 2–3 meshes/cm ²), mostly elongate, up to 45 mm in length, and 25 mm in width. Mesh branches are squarish in section, from 3 mm thick at their base to 1 mm at their tips. No midribs cross the fans, just some thick branches (up to 6 mm in diameter) at the base that diminish and merge with the fan. End­branchlets are long; up to 25 mm in length. Freetwigs are abundant, up to 15 mm in length; they stick out from the fans, twist and grow parallel as free branches. The polyps are retracted within dome­shaped coenenchymal mounds, which are prominent and arranged mostly in pairs along the sides of the branches. In dry specimens, the lateral distribution of the calices is more evident, and bands of coenenchyme are clear between them. The polyps are yellowish with rods arranged in strong, thick points, with some untidily arranged intermediate rods. The anthocodial rods are long, colourless or pale yellow (up to 0.18 mm in length and 0.02 mm in width). The coenenchymal sclerites are very ornamented, and are mostly large spindles (up to 0.22 mm in length, and 0.06 mm in width) with up to 8 complete whorls of tubercles, and warty ends. They are red­orange to pale yellow and bicoloured, and together with P. s e n t a, include the longest spindles found in the genus. Capstans are less abundant in the slide samples; they are scarcely ornate, with only short tubercles. Holotype. The holotype (Fig. 1 G) is a single fan, 100 mm in height and 135 mm in width. Part of the holdfast was left behind when the specimen was collected. The preserved colony is reddish brown colony. Mesh branches are thick, about 2 mm in diameter. Numerous free twigs radiate from the fan as free branchlets. End­branchlets reach 12 mm. Coenenchymal sclerites are red­orange, pale yellow and bicoloured. They are mostly large spindles (up to 0.18 mm in length, and 0.06 mm in width) with 4–8 complete whorls of delicately sculpted tubercles, and with elongated warty ends, blunt, or acute (Fig. 5 A). There are also small, pale yellow capstans (up to 0.05 mm in length and 0.03 mm in width), and larger ones (up to 0.08 mm in length by 0.04 mm in width) with short, moderately warty tubercles (Fig. 5 B). Anthocodial sclerites are pale yellow. They are thin, long rods (up to 0.17 mm in length and 0.02 mm in width) with dentate margins and have acute, small warts, concentrated at the ends (Fig. 5 C). Remarks. This species is very similar to P. s e n t a, however, P. senta attains a larger size, the mesh branches are thinner, the meshwork finer (up to 23 mm long), and the colony has a more delicate appearance in comparison to the more robust P. sculpta. Dry specimens of P. s e n t a are brittle and the sclerites fall off easily, which is not the case in dry specimens of P. sculpta. Sclerites in both species are the largest recorded for the genus. Spindles in P. s e n t a and in P. sculpta reach the same size (up to 0.22 mm in length, and 0.06 mm in width), however, in P. s e n t a, the spindles have more whorls of tubercles (up to 10) than in P. sculpta (up to 8); thus sclerites of the latter have larger spaces between the whorls (and very warty tubercles). Capstans of both species are of similar shapes, but smaller sizes are reported for P. s e n t a (up to 0.06 mm in length) (Breedy & Guzman 2003 b). The colour of coenenchymal sclerites is definitely different. In all of the specimens of P. sculpta examined, two layers of differently coloured sclerites are clearly defined: reddish­orange sclerites in the inner coenenchyme and pale yellow on the surface. In P. s e n t a, on the other hand, all sclerites are of the same colour; brownish pink to colourless. Anthocodial rods are also different, being shorter (up to 0.14 mm in length) and less spiny in P. s e n t a. We have found P. s c u l p t a at several localities in the Gulf of Chiriquí, and also from two sites in the Gulf of Panama, down to 30 m in depth. Breedy & Guzman (2003 b) pointed out that Stiasny (1943) dealt with a species from Isla del Rey, Gulf of Panama, sent to him by Hickson, which agrees with P. senta. Therefore, it was expected that P. s e n t a would be found to occur in Panama. Pacifigorgia senta has been collected from deeper waters, down to 40 m in Costa Rica. In recent collections made by dredging 35–60 m in depth, in Panamá, specimens of P. senta were indeed found, thus the occurrence of P. s e n t a is herein reported and confirmed. Curiously, both P. senta and P. sculpta, were collected together in the same dredge, what indicates that they may occur together. Habitat. Found from 10–40 m in depth, on vertical basaltic walls, living together with large P. e x i m i a colonies and many other octocorals. Though abundant in some places, this species is never the dominant species. Etymology. An adjective (L), sculptus = carved, in allusion to the ornamentation of the spindles. Distribution. Found widely distributed along Gulf of Panama, Gulf of Chiriquí, and Península de Azuero.Published as part of Breedy, Odalisca & Guzman, Hector M., 2004, New species of the gorgoniian genus Pacifigorgia (Coelenterata: Octocorallia: Gorgoniidae) from Pacific Panama, pp. 1-15 in Zootaxa 541 on pages 12-14, DOI: 10.5281/zenodo.15770

    Klinckowstroemia melissae Villegas-Guzman, Reyes-Castillo & Perez 2011

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    Klinckowstroemia melissae Villegas-Guzman, Reyes-Castillo & Pérez, 2011: 25 — Holotype female (CNAC, 006729), from undescribed Pseudacanthus sp., mountain rain forest, 7 km. E San Pedro-San Pablo Ayutla, Ixtlán de Juárez, Oaxaca, Mexico (17 35. 293'N, 96 27. 162'W, 2459 m).Published as part of LIU, DONG, YI, TIAN-CI, XU, YUN & ZHANG, ZHI-QIANG, 2013, new mite species described during 2007 to 2012 3663, pp. 1-102 in Zootaxa 3663 (1) on page 26, DOI: 10.11646/zootaxa.3663.1.1, http://zenodo.org/record/563059

    Klinckowstroemia cristinae Villegas-Guzman, Pérez & Reyes-Castillo, 2009, sp. nov.

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    Klinckowstroemia cristinae sp. nov. Material examined. Holotype. Ƥ (CNAC 006649), Hidalgo, Mexico, 2 km Federal highway Crucero- Tianguistengo (20 ° 40.413 ' N, 98 ° 40.255 ' W), 2,080 m, 23 September 2006, pine-oak forest, from Odontotaenius zodiacus, O. Francke, A. Valdez and H. Montaño coll. Paratypes. 3 Ƥ (CNAC 006650- CNAC 006652), 2 33 (CNAC 006653, CNAC 006654), same data as holotype, from two specimens of Odontotaenius zodiacus; 1 Ƥ (CNAC 006655), Hidalgo, Mexico, 2 km Federal highway Crucero- Tianguistengo (20 ° 40.413 ' N, 98 ° 40.255 ' W), 2,080 m, 23 September 2006, pine-oak forest, from Oileus rimator, O. Francke, A. Valdez and H. Montaño coll. Holotype, three paratype females and two paratype males deposited in the CNAC, one female paratype deposited in NMNH. Female (n = 5). Body oval. Idiosoma, L = 928 (928–1005), W = 672 (672–710) (Fig. 21 A). Ve n tr al idiosoma. Hyaline hood extending to level of coxa I, seta a 1 long and slightly serrated (Fig. 23 A), L = 34 (32–38). Tritosternum base wider than long. Tetartosternum shield (Fig. 22 D), L = 43 (40–43), anterior W = 115 (115–124); shagreened, anterior margin slightly concave, medial notch small, triangular. Sternal seta st 1 moderately long, slightly serrated (Fig. 22 E), L = 20 (20–25), near centre of shield; lyriform sternal pore stp 1 close to posterior margin of shield. Sternal shield, L = 81 (81–93) shagreened. Pore stp 2 near anterior margin of shield, behind and below it is sternal seta st 2, medium length and simple, L = 19 (12–19). Setae st 3 long, slightly serrated, L = 28 (25–37), st 4 is medium length, slightly serrated, L = 16 (16–25). Setae st 3 and st 4 towards sides of shield, close to posterior margin (Fig. 22 C). Sternogynial shield, L = 62, W = 171 (164– 186) triangular, posterior margin pointed at apex, surface shagreened, stp 3 near anterior margin (Fig. 21 A). Sternogynial apodeme long and thin, extending to posterior level of latigynial and mesogynial condyles. Latigynial shields (Fig. 22 C), L = 124 (124–140), W= 78 (74–78) each with an oval pore and three or four simple setae, usually four; medial margins of shields straight and short, shield are shagreened; latigynial apodeme long and thin. Mesogynial shield, L = 74 (71–78), W = 99 (93–102), triangular, wider posteriorly and anteriorly (Fig. 22 C), rounded at apex, surface shagreened, mesogynial condyles below level of shield apex. Ventral shield, L = 245 (245–264), posterior W = 531 (512–557), triangular, truncated posteriorly (Fig. 22 F), surface shagreened, reticulated like a honey-comb (Fig. 22 G), with 12–14 pairs of small setae, usually 14. One pore underneath posterior margin of coxa IV. Metapodal-peritremal-exopodal shield reticulated, with a pore, with two setae. Anal shield, L = 161 (155–177), anterior W = 448 (435–480), with six pairs of simple setae plus longer antero anal seta, L = 29 (29–33), shield shagreened, with two pairs of pores, one near anterior margin and the other below the anus. Gnathosoma. Capitular setae (sc) medium and simple, L = 16 (12–17). Hypostomal setae hyp 1 longest and simple, L = 66 (65–71). Seta hyp 2 long and barbed, 40 (40–45), hyp 3 moderately long and simple, L = 19 (16–19). Chelicera with barbed seta, movable digit with four teeth, two large and two small, fixed digit with five teeth, all similar in size, movable finger divided at the tip. Palpal setae are generally simple, but al 1 (av 1) and al 2 (av 2) on trochanter are branched and serrate respectively. Femur setae pv 1 serrated, al 2, ad 1 and ad 2 barbed. Genu setae al 1 and al 2 are barbed, pl 1 and ad 1 are serrated. Anterior margin of palp trochanter with a blunt process (b) and process (s) (Fig. 22 H). Legs. Most leg setae simple, except: Leg I: trochanter ad 1, al 1 and pv 1 serrated; femur pv 1, pv 2 and pv 3 serrated; genu al 1, av 1 and pv 1 serrated; tibia ad 1, ad 2, ad 3, pd 1 and pd 3 serrated. LegII: femur av 1 and pv 1 serrated; genu av 1 and pv 1 serrated. Leg III: trochanter pv 1 slightly serrated; femur av 1 serrated; genu av 1 and pv 1 serrated; tibia ad 3 serrated. Leg IV: trochanter pl 1 slightly serrated; femur av 1 serrated; genu av 1 and pv 1 serrated; tibia ad 3 barbed. Seta pv 1 is serrated and long, 26 (26–30). Male (n = 3) Body oval, similar to female. Idiosoma, L = 966–1011, W = 698–717 (Fig. 21 B). Ve n tr al idiosoma. Tetartosternum (Tst), L = 47–53, anterior W = 124–127, shagreened (Fig. 23 D), anterior margin concave as in female. Tetartosternal notch small, V-shaped and divides anterior margin in two parts, shield with a pore near posterior margin. Sternal seta st 1, L = 22–25, slightly serrated. Sternal shield shagreened (Fig. 23 B), L = 143–149. Setae st 2, st 3 and st 4, microseta simple; shield with two pores, one near anterior margin of the shield, one below coxa IV. Genital opening oval, L = 62, W= 87–90. Ventral shield, L = 372, posterior W= 544–557; shagreened and reticulated as in female, 34 setae (Fig. 21 B); with two pores, one near anterior margin of coxa IV and one below of coxa IV. Anal shield shagreened, L = 171–177, anterior W = 454; with six pairs of simple setae plus longer antero anal aa seta, L = 31–34, with two pores, one near anterior margin and one below anus. Metapodal-peritremal-exapodal shield reticulated, with a pore, without setae. Gnathosoma. Hypostomal setae (Fig. 23 C), hyp 1 long and simple, L = 65–71, hyp 2 serrated and long, L = 43–47, hyp 3 simple and medium length, L = 19. Etymology. This specific name cristinae is named in honour of Mrs. Cristina Olvera for her hospitality and kindness in hosting the senior author during this research. Remarks. This species can be distinguished from all others because the anterior margin of the tetartosternum shield is concave in both sexes, and sternal seta st 3 is the longest and is slightly serrated. The mesogynial shield is long and wider posteriorly; the latigynial shield is long with the medial margins reduced, all the shields are shagreened, except the ventral shield, which is reticulated. The male has st 2, st 3 and st 4 microsetae and simple; the ventral shield is reticulated and shagreened, and the other shields are only shagreened. The tetartosternal notch is small and V-shaped, and the ventral shield has 34 pairs of simple setae. Klinckowstoemia cristinae is similar to K. schusteri because they have three to four simple setae on the latigynial shield, hyp 2 and hyp 3 are serrated and long and moderatly long respectively, in both species. However, the new species is smaller (idiosoma 953.6 versus 1374), has sternal seta st 3 (29.2) long and slightly serrated, and has shagreened sternogenital shields. In comparison, K. schusteri has several shields reticulated (sternal, sternogynial, mesogynial and latigynial) and st 3 (8) small and simple. Both species have large latigynial shields but that of K. cristinae has reduced medial margins (19.5) compared with K. schusteri. The passalids carrying this species were found in decaying trunks in the same locality at Hidalgo, Mexico, on two hosts, Odontotaenius zodiacus and Oileus rimator. Mites were found in the alcohol, but on O. zodiacus we found a male of K. cristinae on coxa I. On O. rimator we found two other species of klinckowstroemiids, K. scotti and K. cristinae.Published as part of Villegas-Guzman, Gabriel A., Pérez, Tila M. & Reyes-Castillo, Pedro, 2009, New species of the genus Klinckowstroemia Baker & Wharton from Mexico (Acari: Mesostigmata: Trigynaspida: Klinckowstroemiidae), pp. 1-46 in Zootaxa 2248 on pages 40-42, DOI: 10.5281/zenodo.19071

    Klinckowstroemia bifurcata Villegas-Guzman, Pérez & Reyes-Castillo, 2009, sp. nov.

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    Klinckowstroemia bifurcata sp. nov. Material examined. Holotype. Ƥ (CNAC 006514), Hidalgo, Mexico, 2 km Federal Highway Crucero- Tianguistengo (20 ° 40.413 ' N, 98 ° 40.255 ' W), 2,080 m, 23 September 2006, pine-oak forest, from Odontotaenius zodiacus (Truqui), O. Francke, A. Valdez and H. Montaño coll., mites found in the alcohol. Paratypes. 4 Ƥ (CNAC 006515- CNAC 006518), 1 3 (CNAC 006519), same data as holotype, from 3 specimens of Odontotaenius zodiacus; 8 Ƥ (CNAC 006520- CNAC 006527), 2 33 (CNAC 006528, CNAC 006529), Hidalgo, Mexico, 2 km Mojonera-Tizapan, municipality Zacualtipan (20 ° 38.107 ' N, 98 ° 35.981 ' W), 2,019 m, 22 September 2006, from 4 specimens of Odontotaenius zodiacus, O. Francke, A. Valdez and H. Montaño coll. Mites were on coxa I, mesepisternum and loose in the alcohol. Holotype, 9 paratype females and 3 paratype males deposited in the CNAC, one female paratype each deposited in NMNH, UGA and UMMZ. Female (n = 13). Body oval. Idiosoma: L = 966 (922–1,088), W = 723 (659–762) (Fig. 1 A). Ve n tr al idiosoma. Hyaline hood extending to level of coxa I, seta a 1 large and simple (Fig. 2 G), L = 34 (30–37). Tritosternum base wider than long. Tetartosternum (Tst) L = 40 (40–44), anterior W= 124 (112–127) (Fig. 2 E); near the posterior margin is a lyriform sternal pore stp 1. Tetartosternal notch square, shagreened and with a line near the anterior margin. Sternal seta st 1, L = 18 (16–22), bifurcated (Fig. 2 C). Sternal shield (Fig. 2 B) L = 90 (78–91), stp 2 in centre of shield, below it st 2, L = 11 (9–19), simple. Setae st 3, L = 31 (19– 40) and st 4, L = 16 (9–22) near posterior margin of shield, both large and simple, close together (Fig. 2 B). Shield shagreened, without reticulations. Sternogynial shield, L = 68 (62–96); W = 192 (167–201) triangular (Fig. 2 F), shagreened, with stp 3 closer to anterior margin. Sternogynial apodeme short. Latigynial shields, L = 130 (124–146), W = 76 (75–84) each with an oval pore and four setae (Fig. 2 F), medial margin of shield straight, extremity blunt. Mesogynial shield, L = 76 (68–84), W = 106 (96–112), triangular, wider anteriorly and slightly pointed at apex, surface shagreened. Mesogynial condyles extending almost to level of shield apex. Ventral shield, L = 269 (248–288), posterior W = 576 (518–608), shield triangular, truncated posteriorly, surface shagreened, with 12 pairs of small setae (Fig. 1 A). One pore at level of posterior margin of coxa IV. Anal shield, L = 192 (167–218); anterior W = 518 (467–576), with six pairs of simple setae plus longer antero anal seta, L = 25 (17–31), two pairs of pores, one near the anterior margin, the other below the anus. Gnathosoma. Capitular seta (sc) short and serrate, 12 (9–16). Hypostomal seta hyp 1 long and simple, 62 (56–71). Seta hyp 2 long and serrate, L = 40 (34–47). Seta hyp 3 short and simple, L = 7 (7–19) (Fig. 2 A). Chelicera with barbed seta, movable digit with four teeth, fixed digit with five teeth. Palp setae are simple, av 1 and av 2 on trochanter branched and serrate respectively; dorsal setae of femur pl 1 and pl 2 serrate and feathery respectively (Fig. 2 A). Anterior margin of palp trochanter with blunt process (b) (Fig. 2 H). Legs. Most setae on leg I serrate, on trochanter ad 1, al 1 and pv 1, genu al 1 and av 1. Serrate setae on leg II: femur av 1 and genu av 1. Serrate setae on leg III: femur av 1, genu pv 1, tibia ad 3 and al 3; on leg IV, genu pv 1, tibia ad 3 and al 3. Other setae simple, except pv 1 on trochanter II–IV lightly barbed (Fig. 2 D). Male (n = 3). Body oval, similar to female. Idiosoma L = 986–1024, W = 691–736 (Fig. 1 B). Ve n tr al idiosoma. Tetartosternum L = 50–53, posterior W = 102–112, tetartosternal notch shallow (Fig. 4 A), shagreened, with one line near anterior margin as in female. Sternal seta st 1, L = 19–24, simple. The other sternal setae are microsetae. Sternal shield shagreened, L = 152–155 (Fig. 4 B). Genital opening oval, L = 59–66, W = 87–94. Ventral shield with 32 setae (Fig. 1 B), L = 363–378, posterior W = 608, shagreened. Anal shield L = 179–217, anterior W = 512–544; with six pairs of simple setae plus longer antero anal seta, L = 20–30. Gnathosoma. Hypostomal seta hyp 1 is long and simple, L = 65–71; seta hyp 2 is serrate and long, L = 43– 50, while seta hyp 3 is simple and shorter, L = 16–17. Etymology. This name is from Latin bifurcata (= bifurcate) referring to the shape of st 1. Remarks. This species can be distinguished from all others because sternal seta st 1 is bifurcate; and the tetartosternum, sternal and genital shields are shagreened. Sternal setae st 3 and st 4 are slightly serrated, seta st 3 longest. The mesogynial shield is wider on the posterior margin and the latigynial shield has four small simple setae and a pore. Some setae on the palp and leg I are serrated and feathery. The male has sternal seta st 1 simple and the tetartosternum is similar to that of the female. Klinckowstroemia bifurcata is similar to K. reyesi and K. truncata because these three species have the same number of setae on the latigynial, ventral and anal shields. Klinckowstroemia bifurcata (981) is slightly bigger than K. truncata (820) but is slightly smaller than K. reyesi (1009). Klinckowstroemia bifurcata has four setae on the latigynial shield and sternal seta 3 and 4 are serrate as in K. truncata. Sternal seta 3 is longest as in K. reyesi. Klinckowswtroemia bifurcata has the medial margin of the latigynial shield straight and its extremity blunt, while that of K. reyesi and K. truncata is longer and completely straight. The specimens were found in decaying trunks at two different localities in the state of Hidalgo, Mexico, but on the same host (Odontotaenius zodiacus). In these localities we collected three and four passalids carrying K. bifurcata. The mites were usually found in the alcohol, only in one case they were located on the passalid’s body, on coxa I and mesoepisternum. Two or more mite species were found on one beetle: one male of K. victoriae, seven (4 Ƥ, 3 33) of K. bifurcata; and another passalid was found with three species, K. bifurcata; K. hunteri and K. valdezi. This association is known as synhospitality (Eichler 1966).Published as part of Villegas-Guzman, Gabriel A., Pérez, Tila M. & Reyes-Castillo, Pedro, 2009, New species of the genus Klinckowstroemia Baker & Wharton from Mexico (Acari: Mesostigmata: Trigynaspida: Klinckowstroemiidae), pp. 1-46 in Zootaxa 2248 on pages 7-8, DOI: 10.5281/zenodo.19071

    Klinckowstroemia rectimarginata Villegas-Guzman, Pérez & Reyes-Castillo, 2009, sp. nov.

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    Klinckowstroemia rectimarginata sp. nov. Material examined. Holotype. Ƥ (CNAC 006597), Veracruz, Mexico, Dos Amates, Municipality Catemaco (18 ° 29.35 ’ N, 95 °03.58’ W), 120 m, 15 November 1987, from Paxillus leachi MacLeay, P. Hunter coll. Paratypes. 2 Ƥ (CNAC 006598, CNAC 006599), 2 33 (CNAC 006600, CNAC 006601), Veracruz, Mexico, Coyame Bastonal, Municipality Catemaco (18 ° 23.25 ’ N, 94 ° 56.36 ’ W), 14 November 1987, from Heliscus tropicus, P. Hunter coll.; 1 3 (CNAC 006602), Veracruz, Mexico, Coyame Bastonal, Municipality Catemaco (18 ° 23.25 ’ N, 94 ° 56.36 ’ W), 830–970 m, 14 November 1987, from Ve t u r i u s marilucae Boucher, P. Hunter coll. Holotype, two paratype females and three paratype males deposited in the CNAC. Female (n = 4). Body oval. Idiosoma, L = 864 (813–902), W = 634 (614–691) (Fig. 12 A). Ve n tr al idiosoma. Hyaline hood extending to level of coxa I, seta a 1 moderately long and simple, L = 22 (19–25). Tritosternum base wider than long. Tetartosternum shield (Tst), L = 34 (34–40), anterior W= 115 (105–115); shagreened, anterior margin with two lines (Fig. 13 E), tetartosternal notch like a small cavity in centre of margin. Seta st 1 near shield margin, medium and slightly pilose (Fig. 13 F), L = 25 (19–29), lyriform pore stp 1 near posterior margin. Sternal shield (StS) L = 71 (68–71), shagreened. Pore stp 2 near centre of shield, seta st 2 below and behind it, a simple microseta. Sete st 3, L = 7.9 0.9 (7.4–9.3) and st 4, L = 7 (7–9) near posterior margin of the shield, both small and simple (Fig. 12 A). Sternogynial shield (SgS), L = 53 (53– 56), W = 146 (146–171) triangular, anterior margin slightly pointed at apex, surface shagreened, stp 3 near anterior margin (Fig. 13 G). Sternogynial apodeme long and thin, extending posteriorly to level of latigynial and mesogynial condyles. Latigynial shields, L = 127 (124–130), W = 71 (71– 74) each with an oval pore and three setae, medial margins of shield medium and straight, surface shagreened; latigynial apodeme thin. Mesogynial shield, L = 74 (68–81), W = 99 (93–102), triangular, pointed at apex and slightly wider posteriorly, surface shagreened. Mesogynial condyles behind level of shield apex. Ventral shield, L = 202 (202–223), posterior W = 461 (448–512), shield triangular, truncated posteriorly, surface shagreened, with 12 pairs of small setae (Fig. 12 A). One pore underneath posterior margin of coxa IV. Metapodal-peritremalexopodal shield with a pair of pores and two setae, reticulated. Anal shield, L = 140 (118–146); anterior W = 397 (384–429), with five pairs of simple setae plus longer antero anal seta, aa, L = 19 (19–25), three pairs of pores, two near anterior margin, the other below the anus (Fig. 12 A). Gnathosoma. Capitular setae (sc) short and serrated, 9 (9–12). Hypostomal setae (Fig. 13 D), hyp 1 long and simple, L = 53 (53–56), longest. Seta hyp 2 moderate and barbed, L = 34 (28–34). Seta hyp 3 short and simple, L = 12. Chelicera with barbed seta, movable digit with five teeth, fixed digit with six teeth. Palp setae simple, except al 1 (av 1) and al 2 (av 2) on trochanter branched and serrate respectively. Femur setae av 1, pl 1 and pd 1, serrated, pd 2 pilose. Genu setae pl 1, pl 2, pd 1, pd 2 and av 1 serrated. Anterior margin of palp trochanter with blunt process (b) and denticulated process (s) (Fig. 13 H). Legs. Most leg setae simple, except: Leg I: trochanter I al 1 and pv 1 serrated; femur I pv 1 and pv 3 serrated; genu I pl 1 and pv 1 slightly serrated; tibia I ad 1, ad 2, ad 3, pd 1 and pd 3 serrated. Leg II: trochanter II pv 2 serrated; femur II av 1 and pv 1 slightly serrated; genu II av 1 and pv 1 slightly serrated. Leg III: trochanter pd 1 and pd 2 pilose; femur av 1 slightly serrated; genu av 1, pl 1 and pv 1 barbed; tarsus ad 3 serated. Leg IV: trochanter pv 1 pilose; femur av 1 serrated; genu pv 1 serrated. Seta pv 1 is large and slightly serrated, L = 22 (22–25) (Fig. 15 A). Male (n = 3) Body oval similar, to female. Idiosoma, L = 845–890, W = 614–685 (Fig. 12 B). Ve n tr al idiosoma. Tetartosternum, L = 34–37, anterior W = 109–118, shagreened and reticulated, with four horizontal lines, anterior margin reduced medially but without a notch (Fig. 15 B), pore stp 1 near posterior margin. Sternal seta st 1, L = 6–17, simple (Fig. 15 C). Sternal shield shagreened and reticulated, L = 130–139. Setae, st 2, st 3 and st 4, are simple microsetae; with two pores, one near anterior margin of shield, one close to genital opening (Fig. 15 D). Genital opening oval, L = 53–59, W = 71–81. Ventral shield, L = 301– 316, posterior W = 461–512, shagreened, with 38 setae and two pores, one at level of anterior margin of coxa IV and other below coxa IV. Anal shield shagreened, L = 130–140, anterior W = 384–416, with five pairs of simple setae plus longer antero anal seta, L = 25–28, and two pores, one near anterior margin and one below the anus. Metapodal-peritremal-exopodal shield reticulated, with a pair of pores and two setae (Fig. 12 B). Gnathosoma. Hypostomal seta hyp 1 long and simple, L = 58–59, hyp 2 barbed and intermediate in length, L = 34–40, hyp 3 simple and shorter, L = 7–12. Etymology. This name is from Latin rectus (= straight) and marginata (= margin) referring to the shape of the anterior and posterior margins of the tetartosternal shield. Remarks. This species can be distinguished from all others because the tetartosternal shield is rectangular; the tetartosternal notch is very small in the female and absent in the male. The female has a large genital shield. The sternoventral suture is near the anterior margin of the anal shield. Seta st 1 is long and serrated in the female, and medium length and simple in the male. In the female sternal seta st 2 is a microseta and st 3 and st 4 are short and simple; while in male all sternal setae are microsetae. Seta al 2 (av 2) is large and slightly serrated on both sexes. The male has four horizontal lines on the tetartosternal shield, the sternal shield is reticulated and shagreened, and the ventral shield is shagreened, with 19 pairs of setae. Klinckowstroemia rectimarginata is similar to K. schusteri and K. brevimarginata because they have st 1 and hyp 2 serrated and barbed respectively. In the new species the medial margins of the latigynial shields are straight and reduced (16–22), which is similar to K. brevimarginata; seta st 1 are of medium length (19–29) and serrate but slightly longer in K. rectimarginata than K. brevimarginata. In both species the other sternal setae are simple microsetae, while in K. schusteri the setae are bigger, st 2 and st 3 are simple and st 4 is slightly serrated. The sternal, tetartosternal, ventral and genital shields are shagreened; these are similar in K. brevimarginata and K. rectimarginata, while in K. schusteri they are reticulated. There are four setae on the latigynial shield in K. schusteri, four to six in K. brevimarginata and three in K. rectimarginata. Males present more differences; the tetartosternal shield is reticulated and its anterior margin reduced, and the tetartosternal notch is slightly evident. The tetartosternal, sternal and ventral shields are reticulated and shagreened. The specimens were found in decaying trunks in the same locality from Veracruz, but on three different hosts.Published as part of Villegas-Guzman, Gabriel A., Pérez, Tila M. & Reyes-Castillo, Pedro, 2009, New species of the genus Klinckowstroemia Baker & Wharton from Mexico (Acari: Mesostigmata: Trigynaspida: Klinckowstroemiidae), pp. 1-46 in Zootaxa 2248 on pages 25-26, DOI: 10.5281/zenodo.19071
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