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Lonchopterites Grimaldi and Cumming
Genus Lonchopterites Grimaldi and Cumming Lonchopterites Grimaldi and Cumming, 1999: 82. Type species: L. prisca Grimaldi and Cumming, 1999 (Lebanese amber). DIAGNOSIS: (slightly revised, in italics): Antenna with aristal base situated terminally (L. prisca Grimaldi and Cumming) or slightly dorsally (L. burmensis, n. sp.); arista 2-articled, basal one minute, ringlike. Head with large pair of reclinate interfrontals; large pair of lateroclinate fronto-orbitals; ocellar setae large, inside (L. prisca) or outside of triangle (L. burmensis, n. sp.). Wing apex faintly pointed; vein C with large spinules; R 4+5 ending at tip of wing; R 2+3 and R 4+5 divergent at base of wing; M 1 and M 2 diverge near middle of wing; crossveins r-m and dm-cu lacking; anal lobe present, but small; anal vein short, incomplete. The genus is defined largely on the basis of features that are symplesiomorphic for the Lonchopteroidea. TYPE SPECIES: L. prisca Grimaldi and Cumming. DIAGNOSIS: (slightly revised, in italics): Antenna with aristal base situated terminally (L. prisca Grimaldi and Cumming) or slightly dorsally (L. burmensis, n. sp.); arista 2-articled, basal one minute, ringlike. Head with large pair of reclinate interfrontals; large pair of lateroclinate fronto-orbitals; ocellar setae large, inside (L. prisca) or outside of triangle (L. burmensis, n. sp.). Wing apex faintly pointed; vein C with large spinules; R 4+5 ending at tip of wing; R 2+3 and R 4+5 divergent at base of wing; M 1 and M 2 diverge near middle of wing; crossveins r-m and dm-cu lacking; anal lobe present, but small; anal vein short, incomplete. The genus is defined largely on the basis of features that are symplesiomorphic for the Lonchopteroidea. TYPE SPECIES: L. prisca Grimaldi and Cumming.Published as part of Grimaldi, David A., 2018, Basal Cyclorrhapha In Amber From The Cretaceous And Tertiary (Insecta: Diptera), And Their Relationships: Brachycera In Cretaceous Amber Part Ix David A. Grimaldi, pp. 1-97 in Bulletin of the American Museum of Natural History 2018 (423) on pages 1-97, DOI: 10.1206/0003-0090-423.1.1, http://zenodo.org/record/461300
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Grimaldi-Bensouda L, Marty M, Pollak M, Cameron D, Riddle M, Charbonnel B, Barnett AH, Boffetta P, Boivin JF, Evans M, Rossignol M, Benichou J, Abenhaim L; ISICA group
Calvopeza Grimaldi 2018, new genus
Calvopeza, new genus DIAGNOSIS: (female only). Macrosetae absent from head, thorax, and abdomen (some fine setae/setulae on legs); fine, minute acrostichals present in 2–3 irregular, short rows. Scutum strongly arched; palp large, long. Vein M 1 arched strongly away from M 2; membrane of wing mostly with microtrichia, except in basal cells; A 1 +CuA 2 apically incomplete, A 2 spectral. Tibiae and tarsi with dark bifid scales; metatarsomeres 1–4 slightly flattened and expanded. TYPE SPECIES: C. divergens, new species. ETYMOLOGY: From calvus (L., “naked, bare”), in reference to the virtual lack of setae on the head and body, and - peza, a common suffix in genera of Platypezidae. The genus name is feminine.Published as part of Grimaldi, David A., 2018, Basal Cyclorrhapha In Amber From The Cretaceous And Tertiary (Insecta: Diptera), And Their Relationships: Brachycera In Cretaceous Amber Part Ix David A. Grimaldi, pp. 1-97 in Bulletin of the American Museum of Natural History 2018 (423) on pages 1-97, DOI: 10.1206/0003-0090-423.1.1, http://zenodo.org/record/461300
Narcissomyia Grimaldi 2016, new genus
<i>Narcissomyia</i>, new genus <p> DIAGNOSIS: Antenna with 8 flagellomeres, tapered apicad, most longer than wide; eyes bare, holoptic in male; palp 2-segmented; no scutellar spines; tibial spurs 0-1-0, mesotibial spurs long, slender, apex slightly hooked; Abdominal tergites 2–6 each with deeply impressed transverse sulcus; thorax deep in lateral view; wing slender W/L 0.34, Rs branching from R slightly basal to level of CuA 2 (vs. apical to this), cell d long (L/ W 3.2 vs. ≤ 2.5 in almost all other stratiomyids); CuA 2 strongly arched; R 4 long, forking from R 4+5 at level of m 1 -m 2 crossvein. Male: gonostylus with 3 apical spines, gonocoxa with strongly curved basal-medial spine. Female terminalia not telescoping, cerci slender.</p> <p> TYPE SPECIES: <i>Narcissomyia bella</i>, new species.</p> <p> ETYMOLOGY: After the beautiful mythological youth of Greek mythology, Narcissus, in reference to the exquisite preservation of the holotype male, plus <i>myia</i>, Greek for “fly.”</p>Published as part of <i>Grimaldi, David A., 2016, Diverse Orthorrhaphan Flies (Insecta: Diptera: Brachycera) In Amber From The Cretaceous Of Myanmar: Brachycera In Cretaceous Amber, Part Vii David A. Grimaldi, pp. 1-132 in Bulletin of the American Museum of Natural History 2016 (408)</i> on page 27, DOI: 10.1206/0003-0090-408.1.1, <a href="http://zenodo.org/record/5380942">http://zenodo.org/record/5380942</a>
Lebanopeza Grimaldi 2018, new genus
Lebanopeza, new genus DIAGNOSIS: (male only). Eyes holoptic, dorsal facets larger; small prescutellum present; acrostichals in short uniserial row; wing with microtrichia over entire surface; C ends at apex of R 4+5 (vs. M 1); crossvein dm-cu (cell dm) absent, CuP curved; M entirely spectral, branches of M 1 -M 2 fork virtually symmetrical; dark bifid scales lacking on legs; metatarsomeres not expanded; male terminalia lateroflexed to right. Except for the lack of crossvein dm-cu and the faint M vein, all these features appear plesiomorphic for the Platypezidae. TYPE SPECIES: L. azari, new species. ETYMOLOGY: From Lebanon, the country of origin, and - peza (foot), a common suffix for generic names in the Platypezidae. COMMENTS: Lebanopeza appears to be an extinct stem group to Microsania + Melanderomyia (fig. 27). It lacks at least four features that define these two extant genera (chars. 3, 16, 17, 18; table 2), but shares with them the loss of crossvein dm-cu (cell dm), a distinctly shortened Sc vein and somewhat shortened R 1, and reduction in the medial veins. In Melanderomyia M 2 is lost; in Microsania the base of M 1 is lost; in Lebanopeza the entire stem and fork of M 1 -M 2 is extremely faint and unsclerotized. The wing of Lebanopeza is similar to Mauritulus sospes Mostovski, preserved as a compression (along with portions of the body) in shale from the Early Cretaceous of Eurasia (Mostovski, 1995a). Both genera have a slender wing; membrane densely covered with microtrichia; R 1 straight, relatively short; C definitely (Lebanopeza) or apparently (Mauritulus) ending at the apex of R 4+5; M 1 -M 2 fork nearly symmetrical; and crossvein dm-cu (cell dm) absent. Lebanopeza differs from Mauritulus by having a shorter Sc cell (0.3× wing length, vs. 0.45×), cell cup with apical stem of CuP+A 1 (in Mauritulus these veins join virtually at the wing margin), and in having no metatarsomeres expanded (in Mauritulus the basitarsomere is slightly expanded).Published as part of Grimaldi, David A., 2018, Basal Cyclorrhapha In Amber From The Cretaceous And Tertiary (Insecta: Diptera), And Their Relationships: Brachycera In Cretaceous Amber Part Ix David A. Grimaldi, pp. 1-97 in Bulletin of the American Museum of Natural History 2018 (423) on pages 1-97, DOI: 10.1206/0003-0090-423.1.1, http://zenodo.org/record/461300
Stenomicra brunnea Grimaldi 2009, new species
Stenomicra brunnea, new species Figures 4, 7 DIAGNOSIS: Frons whitish, pollinose; scutum and scutellum dark brown, scutum with yellow longitudinal stripe that is narrower than distance between dorsocentrals; pleura without brown infuscation. All abdominal tergites brown; wings hyaline. Scutellum narrow. Anterior fronto-orbital and postvertical setae minute and light, barely detectable. Arista with 5 dorsal, 2 ventral branches. Besides coloration, also distinguished from similar species by male terminalia: Surstylus relatively short, apical third curved upward, apex acute; with dense setulae over much of surstylus. Aedeagus with straight trunk, apex brushy and with narrow tip. TYPES: Holotype, male (dissected, no. 57), FIJI: Taveuni: Cakaudrove Prov., 5.6 km SE Tavuki Vlg., Devo Peak, 1187 m, 30.VI– 14.VIII. 2004, Malaise 1, Schlinger, M. Tokota’a, 16.843 ° S, 179.966 ° W, FBA 50904. ThL 5 1.22 mm; L r-m / L between r-m and mcu 5 0.53. In BPBM. Specimen is missing the scutellar and most dorsocentral setae. In addition, there are 3 females that match the male holotype in color and chaetotaxy, also from Cakaudrove Prov.: 3.2 km NW Lavena Vlg., Mt. Koronibuabua, 217 m, 24. II –11. III.04, Malaise 3, Schlinger and Tokota’a, 16.855 ° S, 179.89 ° W, FBA127415, 153258, 168376. ETYMOLOGY: From the Latin, brunneus, for brown, in reference to the color of the scutum and tergites.Published as part of Grimaldi, David A., 2009, The Asteioinea of Fiji (Insecta: Diptera: Periscelididae, Asteiidae, Xenasteiidae), pp. 1-60 in American Museum Novitates 3671 on page 5, DOI: 10.1206/685.1, http://zenodo.org/record/535802
Stenomicra xoutha Grimaldi 2009, new species
Stenomicra xoutha, new species Figures 4, 11 DIAGNOSIS: Frons whitish, pollinose; scutum and scutellum very light brown/dark ochre, lighter (yellow) between dorsocentrals; tergites I–IV light brown. Dorsal portion of pleura light brown. Pair of anterior frontoorbital setae minute, very light and barely detectable. Arista with 4 dorsal and 2 dorsal branches. Wing hyaline. Best distinguished from similar Stenomicra by male terminalia: cerci without pair of long, thick setae on ventroapical margins; aedeagus curved, apex brushlike; surstylus curved, apex acute, base broad and with dense, long setulae on inner margin. TYPES: Holotype, male (dissected, no. 56), FIJI: Viti Levu: Naitasiri Prov., Nakobalevu Mt., FJ-4, 22.IX–9.X.02, 178 ° 259E, 18 ° 039S, rainforest, M. Irwin, E. Schlinger, M. Tokota’a, Malaise trap 340 m., FBA 013231. ThL 5 1.32 mm; wing W/L 5 0.40; L r-m / L between r-m and m-cu L r-m / L between r-m and m-cu 5 0.33. In BPBM. In addition, there is a series of seven females that are tentatively being assigned to S. xoutha, on the basis of coloration, minute frontoorbitals, and aristal branches. Four females are from the type locality (Nakobalevu, 2 collected at 340 m, other 2 not specified: nos. FBA013232, 013233, 210864, 210865). A further two females are also from Viti Levu, Naitasiri Prov.: 1 female 3.2 km E Navsi Vlg., Veilaselase Track, 1020 m, 19.IV–14.V.2004, Malaise 2, Schlinger, Tokota’a, 17.624 ° S, 178.009 ° E, FBA 155656. 1 female: Naitasiri Prov., Eteni, Navai, Malaise trap, 6.VI–15.VII.2003, FJ-11B, 700 m, E. Schlinger, M. Irwin, M. Tokota’a, 177 ° 599E, 17 ° 379S, FBA013857. Lastly, there are two females from two other islands that are similar to the above females. 1 female: Vanua Levu, Bua Prov., Batiqere Range, 6 km NW Kilaka Village, 98 m, 15– 24.VI.2004, Malaise 5, Schlinger and Tokota’a. 16.807 ° S, 178.991 ° E, FBA161196. 1 female: Taveuni, Cakadrove Prov. Devo Peak Radio Tower, Malaise in rainforest, 10.X–17.X.02, FJ-8, M. Irwin, E. Schlinger, M. Tokota’a, 179 ° 589E, 16 ° 519S, 1200 m, FBA018304. ETYMOLOGY: From the Greek, xouthos, meaning yellowish brown, in reference to the color of the scutum and scutellum.Published as part of Grimaldi, David A., 2009, The Asteioinea of Fiji (Insecta: Diptera: Periscelididae, Asteiidae, Xenasteiidae), pp. 1-60 in American Museum Novitates 3671 on page 17, DOI: 10.1206/685.1, http://zenodo.org/record/535802
Stenomicra castanea Grimaldi 2009, new species
Stenomicra castanea, new species Figures 4, 8 DIAGNOSIS: Frons white; scutum and scutellum evenly light brown, without central longitudinal light stripe; light brown infuscation on dorsal portion of pleura. Wing hyaline. Abdominal tergites I–IV light brown. Besides the scutum coloration, best distinguished on basis of male terminalia: surstyli straight, not curved, with blunt apex, without microtrichia and few setulae; trunk of aedeagus straight, apex brushy; syntergosternite VII ventrally with median lobe on posterior margin. TYPE: Holotype, male (dissected, no. 59), Viti Levu: Naitasiri Prov., 4.8 km N Veisari Stlmt, log rd to Waivudawa, 300 m, 12.XII– 3.I.03. Malaise 1. Schlinger, Tokota’a, 18.075 ° S, 178.362 ° E, FBA 178234. ThL 5 1.20 mm; L rm / L between r-m and m-cu 5 0.50. Type is in good condition, with most of head and thoracic setae intact. In BPBM. ETYMOLOGY: From the Latin castaneus, for the color of chestnuts, in reference to the color of the scutum and scutellum.Published as part of Grimaldi, David A., 2009, The Asteioinea of Fiji (Insecta: Diptera: Periscelididae, Asteiidae, Xenasteiidae), pp. 1-60 in American Museum Novitates 3671 on page 5, DOI: 10.1206/685.1, http://zenodo.org/record/535802
Chandleromyia Grimaldi 2018, new genus
Chandleromyia, new genus DIAGNOSIS: (male only). Eyes extensively holoptic; gena bare, occiput with very fine, scattered short setae; clypeus a broad triangle; palp minute; arista bare; acrostichals entirely absent, only 1 pair of (short, prescutellar) dorsocentrals; 1 short pair of scutellars; 4–5 notopleurals. Wing entirely covered with dense microtrichia; M not forked (M 2 lost), dm-cu close to wing margin; CuP slightly curved. Tibiae and tarsi with bifid scales. Metatarsomeres 1–4 flattened, expanded; tarsomere 2 shortest, with blunt spine; tarsomere 3 with clavate seta. Abdomen short, broad; male tergite 7 narrow, tubular. TYPE SPECIES: C. anomala, n. sp. ETYMOLOGY: For Peter J. Chandler, in recognition of his work on global Platypezidae, and on Diptera in general. The genus name is feminine. COMMENTS: This genus is extremely similar phenetically, and apparently close phylogenetically, to Lindneromyia neomedialis, n. sp., in Dominican amber, described below. This is based on the squared oral margin, minute palps; face, gena, arista bare; complete lack of acrostichals, one (short, prescutellar) pair of dorsocentral setae; one short pair of scutellars; no supraalars, one short postalar; loss of vein M 2; r-m crossvein near level of cell Sc apex; and thick, stiff, spinelike setae on posterior margin of tergite 6 lacking. Chandleromyia differs from L. neomedialis based on the setose occiput (completely bare in Lindneromyia); 4–5 notopleurals (vs. 2); wing membrane entirely with dense microtrichia (vs. partially covered with minute, faint ones); cell dm long, with dm-cu close to wing margin (vs. cell short, vein near middle of wing); vein M 1 deflexed (vs. almost straight); vein CuP curved (vs. straight); spine and clavate seta on metarsomeres 2 and 3, respectively (vs. without); male tergite 7 narrow, tubular (vs. short, much broader than long, inserted into emargination of tergite 6). Though separated by approximately 80 million years, the two flies appear closely related. There is no question about the provenance of the two specimens; AMNH DR14-35 was acquired in the 1990s in the Dominican Republic by me; AMNH Bu-KL30-27 was acquired from Myanmar. With a derived platypezine in the mid-Cretaceous, this greatly extends the age of crown-group Platypezidae (fig. 27).Published as part of Grimaldi, David A., 2018, Basal Cyclorrhapha In Amber From The Cretaceous And Tertiary (Insecta: Diptera), And Their Relationships: Brachycera In Cretaceous Amber Part Ix David A. Grimaldi, pp. 1-97 in Bulletin of the American Museum of Natural History 2018 (423) on pages 1-97, DOI: 10.1206/0003-0090-423.1.1, http://zenodo.org/record/461300
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