197,273 research outputs found

    Travunijana gloeri Grego 2020, sp. nov.

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    Travunijana gloeri sp. nov. urn:lsid:zoobank.org:act: 2A411831-BA56-490B-8754-80A0A0EE048F Figs 9 K–L,15D Type material Holotype BOSNIA AND HERCEGOVINA • 1 shell; Republika Srpska, Bileća Municipality, Berkovići, Vrelo “Vrijeka” in Dabarsko Polje NW of Bjeljani (Fig. 3A); 43°04′28.9″ N, 18°14′20.6″ E; 498 m a.s.l.; 22 Apr. 2019; J. Grego, G. Jakab, M. Olšavský and M. Kováčiková leg.; HNHM-MOLL-104418/1. Paratypes BOSNIA AND HERCEGOVINA • 10 shells; same data; HNHM-MOLL-104419/1, SBMNH 632722 /1, coll. Grego /8, coll. Glöer /1 • 2 shells; same locality as for holotype; D. Angyal, J. Grego, G. Jakab and M. Olšavský leg.; coll. Grego /2. Other material BOSNIA AND HERCEGOVINA • 6 shells; Republika Srpska, Bileća Municipality, Orahovice, Estavela “ Obod ” in Fatničko Polje; leg. 22 Apr. 2019; J. Grego, G. Jakab, M. Olšavský and M. Kováčiková; 43.018543° N, 18.348643° E; coll. Grego /9. Dimensions Holotype: H 3.38 mm; W 1.96 mm; BH 2.23 mm; BW 1.56 mm; AH 1.51 mm; AW 1.29 mm. Paratype: H 3.38 mm; W 1,87 mm; BH 2.31 mm; BW 1.60 mm; AH 1.42 mm; AW 1.24 mm. Etymology Named after my friend and renowned malacologist Peter Glöer from Hetlingen, Germany, who largely contributed to the freshwater gastropod fauna of the Balkans. Description Inflated conical 3.38 mm-high shell with dominant convex body whorl. Light yellowish corneous shell with smooth shiny surface, weak close-set axial ribs and 4.5 convex whorls separated by a weak suture. Protoconch surface smooth, weakly and distantly pitted with larger weak malleation over the pitting. Apex blunt and flat. Umbilicus closed. Elongate asymmetric-oval aperture attached to the body whorl by a weak sulcus. Peristome callous and blunt, not reflexed. Outer lip almost flat at and weakly sinuated at columellar profile. Aperture profile aligned with the body whorl and with the evenly tapering whorls at basal view. Differentiating features The shells of T. gloeri sp. nov. differ from T. ovalis (Fig. 7 I–K) from spring of Bunica River near Hodbina (but not found in Vrelo “Buna” as declared by Kuščer (1933) as its type locality) by its more slender shell shape, closed umbilicus and more flat protoconch. From the type species T. vruljakensis it differs by adapically more protruded and differently shaped aperture, by more coarsely ribbed teleoconch surface and by more smooth protoconch surface. Shell morphometry comparison with the related Travunijana species is presented in the Table 4. Habitat See habitat of P. reischuetzorum sp. nov. Distribution The new species is known only from the type locality.Published as part of Grego, Jozef, 2020, Revision of the stygobiont gastropod genera Plagigeyeria Tomlin, 1930 and Travunijana Grego & Glöer, 2019 (Mollusca; Gastropoda; Moitessieriidae and Hydrobiidae) in Hercegovina and adjacent regions, pp. 1-56 in European Journal of Taxonomy 691 on pages 46-52, DOI: 10.5852/ejt.2020.691, http://zenodo.org/record/396184

    Plagigeyeria reischuetzorum Grego 2020, sp. nov.

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    Plagigeyeria reischuetzorum sp. nov. urn:lsid:zoobank.org:act: A15894D4-DED0-4678-938A-FE39F32E0C51 Figs 8A, 9 G–J Type material Holotype BOSNIA AND HERCEGOVINA • 1 shell; Republika Srpska, Bileća Municipality, Berkovići, Vrelo “Vrijeka” in Dabarsko Polje NW of Bjeljani (Fig. 2 B–G); 43.07472222° N, 18.23916667° E; 483 m a.s.l.; Aug. 2009; Alexander Reischütz, Nicole Steiner-Reischütz and Peter L. Reischütz leg.; NHMW 111678. Paratypes BOSNIA AND HERCEGOVINA • 49 shells; same data as for holotype; NHMW 111679 /1, HNHM- MOLL-104165/1, coll. Reischütz /45, coll. Grego /2 • 21 shells; same locality as holotype; Mar. 2010; Alexander Reischütz, Nicole Steiner-Reischütz and Peter L. Reischütz leg.; coll. Reischütz /19, coll. Grego /2 • 3 shells; same locality as holotype; 31 Mar. 2018; J. Grego, D. Angyal, G. Jakaba and M. Olšavský leg.; SBMNH 626404 /1, coll. Grego /2 • 18 shells; same locality as holotype; 22 Apr. 2019; J. Grego, G. Jakab, M. Olšavský and M. Kováčiková leg.; coll. Grego /16. Other material BOSNIA AND HERCEGOVINA • 3 shells; Republika Srpska, Bileća Municipality, Orahovice, Estavela “ Obod ” in Fatničko Polje; 22 Apr. 2019; J. Grego, G. Jakab, M. Olšavský and M. Kováčiková leg.; 43.018543° N, 18.348643° E; 452 m a.s.l.; coll. Grego/3. Dimensions Holotype: H 2.73 mm; W 1.73 mm; BH 1.66 mm; BW 1.19 mm; AH 1.29 mm; AW 1.05 mm. Paratypes: H 2.51 mm; W 1.73 mm; BH 1.66 mm; BW 1.19 mm; AH 1.36 mm; AW 1.19 mm. H 2.64 mm; W 1.68 mm; BH 1.64 mm; BW 1.22 mm; AH 1.27 mm; AW 1.15 mm. H 2.42 mm; W 1.69 mm; BH 1.53 mm; BW 1.19 mm; AH 1.16 mm; AW 1.16 mm. Etymology Named after the Austrian malacologists Alexander and Peter L. Reischütz with Nicole Steiner-Reischütz, who were the first to collect the species in the type locality. Description Shell is narrow-conical, 2.7 mm high with rounded apex and yellowish corneous periostracum, with five convex pagoduline whorls with rather weak suture. Shell surface finely axially ribbed at late whorls of the teleoconch. Faintly cancellate nepionic whorls covered by 25 close-set spiral ribs. Density of spiral ribs apically decreasing at protoconch to 15 and fading out at malleate nucleus. Space among the protoconch ribs is covered by characteristically granulated surface. Umbilicus narrow slit-like and obscured by the strongly reflexed columellar marginal fold. Larger expanding reflexed trumpet-like aperture irregularly oval and attached to body whorl by a parietal callus. Peristome sharp, and strongly outward reflexed along the aperture outline. Outer lip lateral profile and columellar lip profile characteristically sinuous. Expanded aperture conspicuously protruded against shell outline at basal view. Differentiating features The shells of P. reischuetzorum sp. nov. differ from the specimens of P. mostarensis known from the spring of the Bunica River near Hodbina (Fig. 5 A–C) by their more inflated whorls and less sinuated labral and columellar margins, as well by a more prominent and a less declined aperture at the shell lateral outline. Shell morphometry comparison with the related species is presented in the Table 1. Habitat The type specimens were found in sediments washed out from the cave spring “Sušica Pećina” and its side permanent spring Vrelo “Vrijeka” (483 m a.s.l.) situated 100 m eastward from the cave and rising from debris under the road to Fatnica. It is a part of northern Bregava Basin, likely draining the karst waters from Lukavačko Polje (895 m a.s.l.) into Dabarsko Polje (472 m a.s.l.). During the sampling in March 2018 the adjacent parts of Dabarsko Polje were flooded by large spring outlets creating a vast lake. Distribution The new species is so-far known only from the type locality.Published as part of Grego, Jozef, 2020, Revision of the stygobiont gastropod genera Plagigeyeria Tomlin, 1930 and Travunijana Grego & Glöer, 2019 (Mollusca; Gastropoda; Moitessieriidae and Hydrobiidae) in Hercegovina and adjacent regions, pp. 1-56 in European Journal of Taxonomy 691 on pages 20-22, DOI: 10.5852/ejt.2020.691, http://zenodo.org/record/396184

    Plagigeyeria ljutaensis Grego 2020, sp. nov.

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    Plagigeyeria ljutaensis sp. nov. urn:lsid:zoobank.org:act: A8D69E19-07EA-4432-9C5A-7FE2DA89BBC4 Figs 5D, 6A Type material Holotype BOSNIA AND HERCEGOVINA • 1 shell; Bosnian Federation, Herzegovina-Neretva Canton, Konjic Municipality, Ljuta River spring (Fig. 2A); 43.644611° N, 18.029714° E; 383 m a.s.l; 29 Mar. 2018; J. Grego, D. Angyal, G. Jakab and M. Olšavský leg.; HNHM-MOLL-104180. Paratypes BOSNIA AND HERCEGOVINA • 37 shells; same data as for holotype; HNHM-MOLL-104181/1, SBMNH 626401 /6, coll. Grego /19, coll. Glöer /1. Dimensions (holotype) H 2.09 mm; W 1.49 mm; BH 1.16 mm; BW 0.93 mm; AH 0.9 mm; AW 0.78 mm. Etymology Named after the type locality: Ljuta River spring (as called in material citation). Description The elongate-conical shell is 2.1 mm in height, with blunt elevated apex, milky-white colour and five inflated whorls separated by a deep suture. Shell surface finely axially ribbed with weak corrugated spiral ribs forming a cancellate structure. Prominent at the nepionic whorl. Wavy spiral cords continue towards the unribbed apex (18 cords) and to protoconch with strongly malleated nucleus. Umbilicus narrow slit-like. Excentric-oval aperture is attached to the body whorl. Peristome blunt and outward reflexed along the aperture outline. Outer lip lateral profile weakly sinuous, columellar lip is weakly concave. Columellar lip profile weakly concave. Expanded aperture conspicuously protruded against body whorl at frontal view. Differentiating features The new species differs from the geographically close P. plagiostoma species from Vrelo Bosna in Ilidža (Fig. 5A) by its more conical shell with more inflated whorls, blunter apex, more narrow umbilicus and larger aperture. Habitat The shells of the new species were found in the sediments of a large karst spring of Ljuta River (383 m a.s.l.) (right tributary of Neretva River) rising from a small cave in a narrow gorge at the southwestern edge of Bjelašnica Massif (2067 m a.s.l.). The karst conduits are likely draining waters from northern Bjelašnica and from Džepi Polje (830 m a.s.l.). They are captured by a waterworks to supply potable water for Konjic. Distribution Only known from the type locality.Published as part of Grego, Jozef, 2020, Revision of the stygobiont gastropod genera Plagigeyeria Tomlin, 1930 and Travunijana Grego & Glöer, 2019 (Mollusca; Gastropoda; Moitessieriidae and Hydrobiidae) in Hercegovina and adjacent regions, pp. 1-56 in European Journal of Taxonomy 691 on page 13, DOI: 10.5852/ejt.2020.691, http://zenodo.org/record/396184

    Plagigeyeria konjicensis Grego 2020, sp. nov.

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    Plagigeyeria konjicensis sp. nov. urn:lsid:zoobank.org:act: 12169801-D468-422F-A065-0263523FB0B5 Figs 5E, 6B Type material Holotype BOSNIA AND HERCEGOVINA • 1 shell; Bosnian Federation, Herzegovina-Neretva Canton, Konjic Municipality, spring at left bank of Ljuta River 300 m west of Ljuta River spring (Fig. 2C); 43.644130° N, 18.020711° E; 345 m a.s.l.; 29 Mar. 2018; J. Grego, D. Angyal, G. Jakab, M. Olšavský leg.; HNHM- MOLL-104174. Paratypes BOSNIA AND HERCEGOVINA • 21 shells; same data as for holotype; HNHM-MOLL-104175/1, SBMNH 626249 /2, coll. Grego /17, coll. Glöer /1. Dimensions (holotype) H 2.09 mm; W 1.28 mm; BH 1.10 mm; BW 0.87 mm; AH 0.84 mm; AW 0.72 mm. Etymology Name derived from Konjic, the closest city to the type locality. Description Slender elongated, 2.1 mm-high shell slightly tapering towards the body whorl; milky-white colour with five convex whorls separated by a deep suture. Shell surface finely and densely axially ribbed. Very faint spiral cords are very close set, almost invisible at the teleoconch, and more prominent at nepionic whorl with 21 close-set spiral cords. Apex blunt and flat, protoconch with very faint spiral cords smoothly fading out towards the smooth, very weakly malleated nucleus. Umbilicus open. Almost round-shaped aperture weakly touching the body whorl in upper columellar part. Peristome blunt, margins reflexed along the aperture outline. Outer lip lateral profile strongly forward protruded and weakly sinuous as well as very weak sinuation present at the almost flat columellar lip profile. Expanded round aperture conspicuously protruding against the body whorl at basal view. Differentiating features The shells of P. konjicensis sp. nov. differ from P. plagiostoma (Fig 5A) and from P. ljutaensis sp. nov. by having a significantly more elongated sub-cylindrical shell with a more blunt apex, a slightly sinuated columellar margin, a less prominent body whorl, by different protoconch structure and by a more rounded aperture. Habitat The empty shells were found in a smaller karst spring rising at the left bank of Ljuta River (345 m a.s.l.) among larger stone debris under the road to Ljuta village; the small rivulet from the spring joining the Ljuta River after 60 m, near the ruins of a water mill. It is likely draining the southernmost tip of Bjelašnica massif, south of the village Vrdolje (between Spiljani, Dubočani, Blace and Vrdolje). Distribution Only known from the type locality.Published as part of Grego, Jozef, 2020, Revision of the stygobiont gastropod genera Plagigeyeria Tomlin, 1930 and Travunijana Grego & Glöer, 2019 (Mollusca; Gastropoda; Moitessieriidae and Hydrobiidae) in Hercegovina and adjacent regions, pp. 1-56 in European Journal of Taxonomy 691 on page 14, DOI: 10.5852/ejt.2020.691, http://zenodo.org/record/396184

    Plagigeyeria pseudocostellina Grego 2020, sp. nov.

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    <i>Plagigeyeria pseudocostellina</i> sp. nov. <p>urn:lsid:zoobank.org:act: 09A7B3DE-5B03-46E1-A0F9-A07BFEDBCA4B</p> <p>Figs 7 G–H, 8 B</p> Type material <p> <b>Holotype</b></p> <p>BOSNIA AND HERCEGOVINA • 1 shell; Bosnian Federation, Mostar Municipality, Blagaj, left bank springs of Buna River 400 m from main Buna River Spring, Vrelo “Buna”; 43.256090° N, 17.900073° E; 36 m a.s.l.; 30 Mar. 2018; J. Grego, D. Angyal, G. Jakab and M. Olšavský leg.; HNHM-MOLL-104164.</p> <p> <b>Paratypes</b></p> <p>BOSNIA AND HERCEGOVINA • 15 shells; same data as for holotype; SBMNH 626414 /10, coll. Grego /9 • 26 shells; Bosnian Federation, Mostar Municipality, Blagaj, spring at left bank of Buna River, under road, 500 m from main Buna River Spring; 43.256090° N, 17.900073° E; 36 m a.s.l.; 30 Mar. 2018; J. Grego, D. Angyal, G. Jakab and M. Olšavský leg.; HNHM MOLL 104177 /1, coll. Grego /24, coll. Glöer /1 • 5 shells; Bosnian Federation, Mostar Municipality, Blagaj, spring at left bank of Buna River, under road close to bridge towards mosque, 600 m from main Buna River Spring; 43.256518° N, 17.897381° E; 36 m a.s.l.; 30 Mar. 2018; J. Grego, D. Angyal, G. Jakab and M. Olšavský leg.; coll. Grego /5.</p> Dimensions <p>Holotype: H 2.34 mm; W 1.30 mm; BH 1.28 mm; BW 1.00 mm; AH 0.97 mm; AW 0.95 mm.</p> <p>Paratype: H 2.10 mm; W 1.55 mm; BH 1.34 mm; BW 1.03 mm; AH 1.00 mm; AW 0.86 mm.</p> Etymology <p> Named after its resemblance to <i>Costellina turrita</i> (Kuščer, 1933), known from a small spring named Izvor in the valley of the Jadro near Split, Croatia.</p> Description <p>Elongate-pyramidal shell 2.3 mm high, with rounded blunt apex; milky-yellowish corneous translucent colour with five convex whorls separated by a deep suture. The shell surface is densely covered by lamellar ribs, the ribbing is absent at the nepionic whorl, where it is replaced by 24 densely set spiral ribs cross-sected by fine axial growth lines forming a cancellate structure. Spiral ribs fading down adapically, in pre-nepionic whorl forming faint transversal intercostal structures. Wavy spiral ribs continue towards the nucleus with intercostal granulation. Umbilicus open. Aperture elongate oval to rounded, expanded, shortly attached to the body whorl. Peristome blunt, and outward reflexed along the aperture outline. Outer lip lateral profile straight and flat as well as flat along the columellar lip. Aperture slightly, sometimes conspicuously protruded against the teleoconch and body whorl profile at basal view.</p> Differentiating features <p> The new species differs from <i>P. mostarensis</i> (Fig. 5 A–C) mainly by its straight labral and columellar margins and more open umbilicus. While possessing a similar shell morphology to <i>Costellina turrita</i> (Kuščer, 1933) (Fig. 5F) from the family Moitessieriidae, it differs by having a more open umbilicus and a more oval and more open aperture. Shell morphometry comparison with the related species is presented in the Table 1.</p> Habitat <p> The type specimens were found in sandy sediments washed out from the four small side springs at the right bank of Bunica River about 300–600 m from the main spring of Buna River (36 m a.s.l.). The position of sandy sediments and condition of the fragile shells suggest better adaptation of the new species to an interstitial habitat than we experienced in any other species of <i>Plagigeyeria</i>, which, in contrast, are more specialised to open phreatic cave habitats. The small springs likely draining the low karst plateau situated at the southern foot of the mountain range west of Blagaj without interconnection to the main karst conduit of the Buna.</p> Distribution <p> The new species is so-far known only from the four small springs of the type locality at the left bank of river Buna in Blagaj, while a local Buna form of <i>P. mostarensis</i> (Fig. 5 D–E) can be found in the sediment of the main spring of Buna River. The typical morphotype of <i>P. mostarensis</i> is found in the nearby spring of river Bunica near Hodbina (Fig. 5 A–C) and we suppose this is the type locality, which was erroneously confused with the Buna spring by Kuščer (1933).</p> Remarks <p> The shell morphology similarity to the moitessieriid <i>Paladilhiopsis (Costellina) turrita</i> could suggest an interlink towards <i>Plagigeyeria mostarensis</i> and other members of the genus <i>Plagigeyeria</i>, and its possible affiliation to the family Moitessieriidae.</p>Published as part of <i>Grego, Jozef, 2020, Revision of the stygobiont gastropod genera Plagigeyeria Tomlin, 1930 and Travunijana Grego & Glöer, 2019 (Mollusca; Gastropoda; Moitessieriidae and Hydrobiidae) in Hercegovina and adjacent regions, pp. 1-56 in European Journal of Taxonomy 691</i> on pages 18-20, DOI: 10.5852/ejt.2020.691, <a href="http://zenodo.org/record/3961840">http://zenodo.org/record/3961840</a&gt

    Travunijana klemmi Grego 2020, comb. nov.

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    Travunijana klemmi (Schütt, 1961) comb. nov. Figs 13B, 16 A–B Plagigeyeria klemmi Schütt, 1961: 133–134, fig. 2. Plagigeyeria klemmi – Schütt 1972: 115, 119 pl. 6 figs 5–6. — Willmann & Pieper 1978: 126. — Bole & Velkovrh 1986: 202. — Bodon, Manganelli & Giusti 1996: 33, fig. 13. — Bank 2013: Fauna Europaea v. 2.6. — Bank & Neubert 2017: 25. Diagnosis T. klemmi has regularly narrow-conical shells (2.2 mm) with blunt apex, deep suture and widely spaced coarse ribs on shell surface, rounded callous irregular-oval shaped aperture and closed umbilicus. Lateral labral profile straight and columellar labrum only weakly sinuated. Protoconch surface overall covered by regular and deep pitting. Distribution T. klemmi is known from Vrelo “Stenjevac” and Vrelo “Baja” in Vrgorsko Polje (20 m a.s.l.) near Vrgorac and from Vrelo “Kutac” and other springs around Baćinska Lakes (0.8 m a.s.l.) near Ploče. The aquifer also receives its water from Rastok Polje (72 m a.s.l.). Remark s Travunijana klemmi represents the northernmost species of the genus. Its shell morphology has some features of the genus Belgrandia (e.g., Belgrandia torifera Schütt, 1961) known from the region, however its shell is more conical and lacks the typical callous varix near the labrum.Published as part of Grego, Jozef, 2020, Revision of the stygobiont gastropod genera Plagigeyeria Tomlin, 1930 and Travunijana Grego & Glöer, 2019 (Mollusca; Gastropoda; Moitessieriidae and Hydrobiidae) in Hercegovina and adjacent regions, pp. 1-56 in European Journal of Taxonomy 691 on page 44, DOI: 10.5852/ejt.2020.691, http://zenodo.org/record/396184

    Figure 4 from: Grego J, Pešić V (2021) First record of stygobiotic gastropod genus Travunijana Grego & Glöer, 2019 (Mollusca, Hydrobiidae) from Montenegro. Subterranean Biology 38: 65-76. https://doi.org/10.3897/subtbiol.38.64762

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    Figure 4 M protoconch SEM images of TravunijanaA, BTravunijana djokovici sp. nov., paratype, SBMNH 632721) CT. vruljakensis Grego and Glöer, paratype SBMNH 625961 DT. gloeri Grego, 2020, paratype SBMNH 632722. Scale bars: 1 mm. (SEM SBMNH Vanessa Delnavaz)

    Plagigeyeria erossi Grego 2020, sp. nov.

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    Plagigeyeria erossi sp. nov. urn:lsid:zoobank.org:act: 405BAA2B-296D-4B18-9D62-343C35A6963E Figs 9 A–C, 10A Type material Holotype BOSNIA AND HERCEGOVINA • 1 shell; Bosnian Federation, Herzegovina-Neretva Canton, Donja Jablanica, Komadinovo Vrelo, 2 km SW of, and under the road from Mostar to Jablanica, side outlet of main spring at left bank of Neretva (Fig. 2D); 43.617298° N, 17.741081° E; 180 m a.s.l.; 30 Jul. 2016; J. Grego and Z. P. Erőss leg.; HNHM-MOLL-104166. Paratypes BOSNIA AND HERCEGOVINA • 3 shells; same data as for holotype; HNHM-MOLL-104167/1, coll. Grego /1, coll. Erőss /1. • 1 shell; same locality as holotype; 29 Mar. 2018; Jozef Grego leg; coll. Grego /1. Dimensions Holotype: H 2.10 mm; W 1.13 mm; BH 1.25 mm; BW 0.90 mm; AH 0.87 mm; AW 0.72 mm. Paratypes: H 2.01 mm; W 1.28 mm; BH 1.12 mm; BW 0.93 mm; AH 0.88 mm; AW 0.77 mm. H 1.79 mm; W 1.09 mm; BH 1.12 mm; BW 0.88 mm; AH 0.79 mm; AW 0.70 mm. Etymology Named after my friend Zoltán Péter Erőss, president of the Hungarian Malacological Society, Budapest, who participated in the field trip and brought my attention to the locality of the new species. Description Solid, narrow, conical shell, 2.1 mm in height, with blunt apex, milky-white colour and four convex whorls separated by a deep suture. Shell surface very finely axially ribbed especially at the late teleoconch and ribs cross-sected by irregular wavy faint spiral structures. Nepionic whorl covered by 18 close-set spiral ribs, continuing apically into fine irregular granulose cord-like structure at protoconch and smoothly malleate nucleus. Umbilicus open, partly obscured by the strongly reflexed columellar margin. Larger expanding reflexed aperture oval, attached to the body whorl in its upper columellar part. Peristome blunt, reflexed along aperture outline. Outer lip lateral profile characteristically sinuous adapically as well as having a characteristic sinuation which is present at the columellar lip. Expanded aperture stepwise protruded against body whorl at frontal view. Differentiating features Compared with the geographically closest Plagigeyeria mostarensis Kuščer, 1933 (Hercegovina, Blagaj) (Fig. 7 A–E), the new species differs by its smaller cylindrical and less conical shell shape, blunter rounded apex and less prominent protoconch, more open umbilicus, as well as by a proportionally smaller and more oval aperture. The new species has a characteristic sinuation at outer and inner lips and a less prominent aperture within the shell peripheral outline. The shell morphology of the new species is closely related to Plagigeyeria plagiostoma (A. J. Wagner, 1914) from the spring of the river Bosna near Ilidža, which differs by its more slender shell, cylindrical shape, more open umbilicus, different sculpture of the protoconch, proportionally smaller aperture and a weaker marginal sinuation. Shell morphometry comparison with the related species is presented in the Table 2. Habitat The type locality is a large karst spring rising at the right bank of Neretva River among large stone blocks under highway (Sarajevo-Mostar) and draining the limestone massif of Mt Prenj (2115 m a.s.l.) in Čvrsnica Massif (highest peak Pločno, 2228 m a.s.l.). Due to a strong current in the spring zone, there was not enough fine sediment in the main spring zone, but empty shells had been found in two smaller side springs at the left side of the main outlet. This species most likely inhabits an undiscovered karst conduit upstream the spring inside the Čvrsnica Massif. Distribution Only known from the type locality.Published as part of Grego, Jozef, 2020, Revision of the stygobiont gastropod genera Plagigeyeria Tomlin, 1930 and Travunijana Grego & Glöer, 2019 (Mollusca; Gastropoda; Moitessieriidae and Hydrobiidae) in Hercegovina and adjacent regions, pp. 1-56 in European Journal of Taxonomy 691 on pages 25-26, DOI: 10.5852/ejt.2020.691, http://zenodo.org/record/396184

    Plagigeyeria angyaldorkae Grego 2020, sp. nov.

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    Plagigeyeria angyaldorkae sp. nov. urn:lsid:zoobank.org:act: 6BC5A082-0F63-4116-92B3-3BD320D5D1FA Fig. 12E Type material Holotype BOSNIA AND HERCEGOVINA • West Herzegovina Canton, Tihaljina District, Lukende, Jakšenica Village, “Vrelo Jakšenica” (Fig. 3F); 43.322897° N, 17.373026° E; 173 m a.s.l.; 2 Apr. 2018; J. Grego, D. Angyal, G. Jakab and M. Olšavský leg.; HNHM-MOLL-104163. Dimensions Holotype: H 3.07 mm; W 1.86 mm; BH 1.45 mm; BW 1.31 mm; AH 1.21 mm; AW 1.21 mm. Etymology Named after my friend Dorottya Angyal from the Budapest Natural History Museum, who substantially helped with the samplings during our field trip. Description Solid, elongate-conical shell, 3.1 mm-high, with 5.5 regularly tapering narrow inflated whorls, deep suture and blunt rounded apex, milky-white colour and fine regular sinuated axial ribs. Late protoconch faintly irregularly spirally ribbed. Umbilicus open, obscured by the outward reflexed flaring columellar margin. Expanding reflexed aperture with irregular elongate-oval shape and a slightly depressed and inward-bent labral margin. The upper labral margin is elongate-flaring. Labral lateral profile protruded and bent backward at the body whorl, columellar margin profile sinuated. Expanded upper tip of aperture is conspicuously protruded, flaring against the frontal shell profile. Differentiating features P. angyaldorkae sp. nov. differs from all other known species of the region by its elongate-conical shell shape with 6 whorls, by more convex whorls and proportionally smaller body whorls and by a characteristically elongated labrally depressed aperture with a slightly depressed labral margin. From Lanzaia ephantota (Megerle von Mühlfeld, 1824) it differs by larger more elongate shell wit finer, more close-set ribbs and more depressed aperture. Shell morphometry comparison with the related species is presented in the Table 3. Habitat The empty shell of this subterranean species was washed out by a middle-sized karst spring (173 m a.s.l.) rising from a 1.5 m-broad limestone crack among houses in the village. The strong stream flows under the main road, depositing the sand at a pond just before the old watermill ruins. The spring is likely getting its water from sinkholes north of village Drinovci (245 m a.s.l.) in Imotsko Polje. Distribution Only known from the type localityPublished as part of Grego, Jozef, 2020, Revision of the stygobiont gastropod genera Plagigeyeria Tomlin, 1930 and Travunijana Grego & Glöer, 2019 (Mollusca; Gastropoda; Moitessieriidae and Hydrobiidae) in Hercegovina and adjacent regions, pp. 1-56 in European Journal of Taxonomy 691 on pages 33-36, DOI: 10.5852/ejt.2020.691, http://zenodo.org/record/396184

    Plagigeyeria vriosticaensis Grego 2020, sp. nov.

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    Plagigeyeria vriosticaensis sp. nov. urn:lsid:zoobank.org:act: BE0E3ADC-7F3F-4866-9BCD-2769746A2A81 Fig. 12 C–D Type material Holotype BOSNIA AND HERCEGOVINA • 1 shell; West Herzegovina Canton, Vitina, Vrelo “Vrioštica” (Fig. 3E); 43.237595° N, 17.485963° E; 98 m a.s.l.; 2 Apr. 2018; J. Grego, D. Angyal, G. Jakab and M. Olšavský leg.; HNHM-MOLL-104182. Paratypes BOSNIA AND HERCEGOVINA • 4 shells; same data as for holotype; SBMNH 626400 /2, coll. Grego /2. Dimensions Holotype: H 2.82 mm; W 1.76 mm; BH 1.66 mm; BW 1.31 mm; AH 1.21 mm; AW 1.17 mm. Paratype: H 3.00 mm; W 1.45 mm; BH 1.72 mm; BW 1.28 mm; AH 1.24 mm; AW 1.28 mm. Etymology Named after the type locality: spring of Vrioštica River. Description Elongate oval shell with 4.5 convex whorls and deep suture, 2.8 mm high, with blunt rounded apex, milky-white colour with rusty incrustations. Shell surface shiny and finely axially ribbed at the teleoconch. Nepionic whorl faintly spirally ribbed. Umbilicus narrow, slit-like. Aperture rhomboid-oval with outward reflex, posteriorly and anteriorly more expanded. Peristome blunt, and reflexed along the aperture outline. Outer lip lateral profile straight and sinuous posteriorly. Sinuation present at columellar marginal profile. Aperture protruded against teleoconch outline from frontal view. Differentiating features The species is closely related to P. angyaldorkae sp. nov., from which it differs by having a smaller number of whorls and a more prominent body whorl and, by a more narrow umbilicus and different shape of aperture. Conchologically, it represents an intermediate shell morphology between P. angyaldorkae sp. nov. and P. jakabi sp. nov., from which it can be distinguished by its more robust shell shape and different shape of the aperture with much broader posterior channel and by a smaller umbilicus. From sympatric P. ozimeci sp. nov. it differs by a larger shell with more whorls, a less conical shell shape and by a different shape of the aperture. Shell morphometry comparison with the related species is presented in the Table 3. Habitat See the habitat of P. ozimeci sp. nov. Distribution The species is only known from the type locality.Published as part of Grego, Jozef, 2020, Revision of the stygobiont gastropod genera Plagigeyeria Tomlin, 1930 and Travunijana Grego & Glöer, 2019 (Mollusca; Gastropoda; Moitessieriidae and Hydrobiidae) in Hercegovina and adjacent regions, pp. 1-56 in European Journal of Taxonomy 691 on page 36, DOI: 10.5852/ejt.2020.691, http://zenodo.org/record/396184
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