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    Charinus brescoviti Giupponi & Miranda 2016

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    Charinus brescoviti Giupponi & Miranda, 2016 Fig. 27; Table 2 Charinus brescoviti Giupponi & Miranda, 2016: 3–6, figs 1a–e, 2a, 3a, 4a. Diagnosis This species may be separated from other Charinus in Amazonia and northern South America by means of the following combination of characters: tegument brownish-yellow in color; median and lateral eyes well developed; tetra- and pentasternum small, rounded; cushion-like gonopods, lateral projections directed posteriorly and internal seminal receptacles; basal spine on pedipalp tarsus large, two-thirds length of distal spine; ventral row of setae of cleaning organ with 28 setae; leg IV basitibia with two pseudo-articles; leg IV basitibia trichobothrium bt situated in proximal third; leg IV distitibia with 16 trichobothria, basal trichobothria, bf, bc and sbf, equidistant. Etymology Patronym honoring Dr Antônio D. Brescovit (Giupponi & Miranda 2016). Type material Holotype BRAZIL • ♀; Amazonas, Piaçaba, Araçá River; [00°25′17.75″ S, 62°54′30.24″ W]; 18 May 1982; B. Mascarenhas leg.; IBSP 149. Paratypes BRAZIL • 1 ♀; same collection data as for holotype; IBSP 149 • 1 ♀, 1 juv.; same collection data as for holotype; MNRJ 9186. Measurements See Table 2. Distribution Known only from the type locality. Natural history Epigean, found close to a river. Remarks See Giupponi & Miranda (2016) for a comparison and discussion of this species.Published as part of Miranda, Gustavo Silva de, Giupponi, Alessandro P. L., Prendini, Lorenzo & Scharff, Nikolaj, 2021, Systematic revision of the pantropical whip spider family Charinidae Quintero, 1986 (Arachnida, Amblypygi), pp. 1-409 in European Journal of Taxonomy 772 on page 48, DOI: 10.5852/ejt.2021.772.1505, http://zenodo.org/record/553641

    Charinus guto Giupponi & Miranda 2016

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    Charinus guto Giupponi & Miranda, 2016 Fig. 27; Table 2 Charinus guto Giupponi & Miranda, 2016: 13–15, figs 2d, 3d, 4d, 7a–e. Diagnosis Based on Giupponi & Miranda (2016), this species may be separated from other Charinus in Amazonia and northern South America by means of the following combination of characters: tegument light brown; median eyes and median ocular tubercle absent; lateral eyes well-developed, but pale; small, rounded tetra- and pentasternum; cushion-like gonopods with lateral projections directed posteriorly, covering aperture of internal seminal receptacles; two dorsal spines on pedipalp tarsus, small basal spine, one-quarter length of distal spine; tibia of leg I with 21 articles, tarsus I with 37 articles; leg IV basitibia with three pseudo-articles; leg IV basitibia trichobothrium bt situated in proximal third; leg IV distitibia with 14 trichobothria; basal trichobothria, bf, bc and sbf equidistant; sc and sf series each with five trichobothria. Charinus guto resembles C. vulgaris, but possesses more spines on the pedipalp femur and patella. Etymology Patronym honoring José Augusto Pereira Barreiros nicknamed Guto (in memoriam; Giupponi & Miranda 2016). Type material Holotype BRAZIL • ♀; Pará, Belém, Bosque Rodrigues Alves; 01°27′21″ S, 48°30′16″ W; 27 Oct. 2002; R. Pintoda-Rocha leg.; MZSP 48146. Paratypes BRAZIL • 3 ♀♀; same collection data as for holotype; MZSP 48146 • 1 ♀; same collection data as for holotype; MNRJ 9188 • 1 juv. ♀; same locality as for holotype; 8 Nov. 2001; J.A.P. Barreiros leg.; MPEG 59 • 1 ♀; same locality as for holotype; 8 Nov. 2001; J.A.P. Barreiros leg.; MNRJ 9202 • 1 ♀; same locality as for holotype; 8 Apr. 2001; A.B. Bonaldo et al. leg.; MPEG 37 • 1 ♀; same locality as for holotype; 8 Apr. 2001; A.B. Bonaldo et al. leg.; MPEG 38. Measurements See Table 2. Distribution Known only from the type locality. Natural history Found in leaf litter. Synanthropic. Remarks See Giupponi & Miranda (2016) for more details about this species.Published as part of Miranda, Gustavo Silva de, Giupponi, Alessandro P. L., Prendini, Lorenzo & Scharff, Nikolaj, 2021, Systematic revision of the pantropical whip spider family Charinidae Quintero, 1986 (Arachnida, Amblypygi), pp. 1-409 in European Journal of Taxonomy 772 on pages 58-59, DOI: 10.5852/ejt.2021.772.1505, http://zenodo.org/record/553641

    Charinus orientalis Giupponi & Miranda 2016

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    Charinus orientalis Giupponi & Miranda, 2016 Fig. 27; Table 2 Charinus orientalis Giupponi & Miranda, 2016: 18–21, figs 2f, 3f, 4f, 9a–e. Diagnosis Based on Giupponi & Miranda (2016), this species may be separated from other Charinus in Amazonia and northern South America by means of the following combination of characters: tegument pale yellow; cushion-like female gonopod with lateral projections directed posteriorly, covering opening of internal seminal receptacles (atrium); median and lateral eyes present, but median ocular tubercle and lateral eyes markedly reduced, as in C. carajas; median ocular tubercle situated in depression; small, rounded tetra- and pentasternum; weakly sclerotized margin of sternum; pedipalp femur with three dorsal spines; proximal spine of pedipalp tarsus ca one-quarter length of distal spine; tibia of leg I with 21 articles, tarsus I with 37 articles; leg IV basitibia with three pseudo-articles; leg IV basitibia trichobothrium bt situated in proximal third; leg IV distitibia with 16 trichobothria; leg IV distitibia bf, bc and sbf trichobothria equidistant; sc and sf series each with five trichobothria. Etymology Latin adjective meaning ‘eastern’, referring to the Sierra Leste mountain range inhabited by the species (Giupponi & Miranda 2016). Type material Holotype BRAZIL • ♀; Pará, Curionópolis, Projeto Serra Leste, SL-82 cave; 7 Jul. 2010; CARSTE leg.; dry season; MNRJ 9249. Paratypes BRAZIL • 2 ♀♀; Pará, Carajás National Forest, Serra do Leste; 05°58′35.171″ S, 49°37′55.377″ W; SAD’ 69; 17–24 Oct. 2008; Andrade et al. leg.; MZSP 29118 • 1 ♀; same collection data as for preceding; Curionópolis; SL-82 cave; 7 Jul. 2010; CARSTE leg.; Projeto Serra do Leste; dry season; MNRJ 9249 • 3 ♀♀, 2 juv.; same collection data as for preceding; SL-82 cave; 7 Jul. 2010; CARSTE leg.; Projeto Serra Leste; dry season; ISLA 3893 • 1 juv. ♀; same collection data as for preceding; SL-75 CL cave; 05°57′54.473″ S, 49°37′55.023″ W; SAD’ 69; 17–24 Oct. 2008; Andrade et al. leg.; MZSP 29119 • 3 ♀♀, 1 juv. ♀; same collection data as for preceding; SL-82 CL cave; 05°57′29.644″ S, 49°38′13.262″ W; SAD’ 69; 17–24 Oct. 2008; Andrade et al. leg.; MZSP 29124. Measurements See Table 2. Distribution This is the only species from the Carajas region which occurs outside the National Reserve, in a cave near the municipality of Curionópolis. Natural history Troglophilous, found only in iron ore caves. Remarks See Giupponi & Miranda (2016) for more details about this species.Published as part of Miranda, Gustavo Silva de, Giupponi, Alessandro P. L., Prendini, Lorenzo & Scharff, Nikolaj, 2021, Systematic revision of the pantropical whip spider family Charinidae Quintero, 1986 (Arachnida, Amblypygi), pp. 1-409 in European Journal of Taxonomy 772 on pages 60-61, DOI: 10.5852/ejt.2021.772.1505, http://zenodo.org/record/553641

    Charinus troglobius Baptista & Giupponi 2002

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    Charinus troglobius Baptista & Giupponi, 2002 Fig. 43; Table 4 Charinus troglobius Baptista & Giupponi, 2002: 106–110, figs 1–14. Charinus troglobius – Baptista & Giupponi 2003: 79–80. — Trajano & Bichuette 2010: 12. — Miranda & Giupponi 2011: 66, fig. 13. — Vasconcelos et al. 2013: 496, fig. 12. — Monte et al. 2015: 5. — Torres-Contreras et al. 2015: 145–146. — Vasconcelos & Ferreira 2016: 185. — Miranda et al. 2016c: 19, 29. Diagnosis Based in part on Baptista & Giupponi (2002), this species may be separated from other Charinus in eastern South America by means of the following combination of characters: corners of anterior carapace margin produced into truncated processes; median eyes and median ocular tubercle absent; lateral eyes reduced to small whitish spot on each side of carapace, without clearly defined lens; sternal platelets narrow, convex and rounded, tritosternum small, slightly larger than tetrasternum; pedipalps articulating transversally with cephalothorax, but not horizontally; pedipalp spines longer than in other species of genus; pedipalp femur with two dorsal spines and two ventral spines; pedipalp tarsus with two long spines, second spine longer than first; proventral corners of walking legs spiniform; leg IV basitibia + distitibia with 17 trichobothria (1 + 16); leg IV basitarsus much longer than distitarsus. This is the most markedly troglomorphic Brazilian species of Charinus, in which the eyes are completely absent and the leg articles and pedipalp spines extremely elongated. Etymology Latin adjective referring to the troglobitic habitus of the species (Baptista & Giupponi 2002). Type material Holotype BRAZIL • ♂; Bahia, Carinhanha, Serra do Ramalho, Gruna do Zé Bastos; 14°16′53.92″S, 43°46′24.29″W; 28 Jun. 2001; R.L.C. Baptista and A.P.L. Giupponi leg.; MNRJ 9067. Paratypes BRAZIL • 1 ♂; same collection data as for holotype; MNRJ 9069 • 1 ♀; same collection data as for holotype; MNRJ 9078 • 1 ♂; same collection data as for holotype; MNRJ 9081 • 2 ♀♀, 9 ♂♂; same collection data as for holotype; MNRJ 9068 • 1 ♀, 1 ♂; same collection data as for holotype; IES ex MNRJ 9079 [not examined] • 1 ♂; same collection data as for holotype; MZSP ex MNRJ 9080 [not examined] • 1 ♀; same collection data as for holotype; MNRJ 9082 • 1 ♀, 1 ♂; same collection data as for holotype; IES 3.2972 and IES 3.2973 [not examined]. Additional material BRAZIL • 1 juv.; Bahia, Serra do Ramalho, Lapa do Boqueirão; 29 Jun. 2001; R.L.C. Baptista and A.P.L. Giupponi leg.; MNRJ 9034. Supplementary description CARAPACE. Margin of carapace ventral to lateral eyes with rounded projection; six anterior setae; frontal process triangular, with oval apex, not visible in dorsal view. Small granules densely scattered between ocular triads and among sulci. Median eyes and median ocular tubercle absent; pair of setae in place of median ocular tubercle; lateral eyes reduced to three small, whitish lenses, with seta posterior to lateral ocular triad. STERNUM. Tritosternum short, with four long setae and several smaller setae posteriorly; other sternal platelets narrow and concave, with pair of setae anteriorly on plaque and some smaller setae posteriorly; pentasternum with four setae anteriorly and seta on membranous region. OPISTHOSOMA. Ventral sacs and ventral sac cover absent. GENITALIA. Female genital operculum with prominent setae posteromedially and some smaller setae near margin; gonopod sucker-like and barrel shaped. Male gonopod with base of fistula and lateral lobe sclerotized; lateral lobe 2 fimbriate. CHELICERAE. Small tooth projecting from retrolateral surface of basal segment, opposite to bifid tooth; retrolateral surface of claw with continous row of setae, basally to medially; claw with ten teeth; transverse row of setae on prolateral surface of basal segment absent, with longitudinal row of dense setae, instead; bifid tooth on basal segment with dorsal cusp larger than ventral cusp. PEDIPALPS. Coxal dorsal carina with six prominent setae encircled by round carina and three setae on margin. Femur with two or three (dextral pedipalp of male paratype) dorsal spines and two ventral spines; dorsal spines situated in distal third; four prominent setiferous tubercles between dorsal spine 1 and proximal margin; setiferous tubercle between ventral spine 1 and proximal margin. Patella with three dorsal spines in primary series; prominent setiferous tubercle distal to spine I, one-third length of spine I, and prominent setiferous tubercle between spines I and II, slightly shorter than first setiferous tubercle; two ventral spines in primary series; extra spine between spine I and distal margin. Tibia with ventral spine distally and three prominent setiferous tubercles proximal to ventral spine; twelve or thirteen setae between spine and distal margin. Tarsus with two dorsal spines, proximal spine one-third length of distal spine; cleaning organ with 29–31 setae in ventral row. LEGS. Tibia of leg I with 23 articles (up to 47 on regenerated legs); tarsus I with 40–42 articles (up to 62 on regenerated legs); first tarsal article shorter than second and subsequent articles. Leg IV basitibia with four pseudo-articles, with sclerotized, denticulate margin projecting from apex of articles; trichobothrium bt situated in proximal third of pseudo-article; distitibia trichobothrium bc situated closer to sbf than to bf, sc and sf series each with five trichobothria. Measurements See Table 4. Distribution Known from Carinhanha in the Brazilian state of Bahia. Natural history According to Baptista & Giupponi (2002), all specimens were collected on muddy walls near water bodies, in the dark zone of caves. A troglomorphic harvestman, Giupponia chagasi Pérez & Kury, 2002, was also found in the cave. Other taxa observed in the cave include spiders (Ctenidae Keyserling, 1877: Ctenus sp.), millipedes (Chelodesmidae Cook, 1895), crickets (Phalangopsidae Blanchard, 1845: possibly Endecous sp.), cockroaches (Epilampridae Brunner von Wattenwyl, 1865) and beetles (Cholevidae Kirby, 1837).Published as part of Miranda, Gustavo Silva de, Giupponi, Alessandro P. L., Prendini, Lorenzo & Scharff, Nikolaj, 2021, Systematic revision of the pantropical whip spider family Charinidae Quintero, 1986 (Arachnida, Amblypygi), pp. 1-409 in European Journal of Taxonomy 772 on pages 135-137, DOI: 10.5852/ejt.2021.772.1505, http://zenodo.org/record/553641

    Charinus bichuetteae Giupponi & Miranda 2016

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    Charinus bichuetteae Giupponi & Miranda, 2016 Fig. 27; Table 2 Charinus bichuetteae Giupponi & Miranda, 2016: 23–25, figs 2h, 3h, 11a–f. Diagnosis Based on Giupponi & Miranda (2016), this species may be separated from other Charinus in Amazonia and northern South America by means of the following combination of characters: tegument pale yellow in color; median eyes and median ocular tubercle absent; lateral eyes well developed; small, rounded, markedly sclerotized tetra- and pentasternum; cushion-like female gonopod with lateral projections directed posteriorly, covering opening of internal seminal receptacles (atrium); pedipalp femur with two dorsal spines and three ventral spines; pedipalp tarsus with two spines, proximal spine small, ca one quarter length of distal spine; tibia of leg I with 21 articles, tarsus I with 37 articles; leg IV basitibia with two pseudo-articles; trichobothrium bt situated in proximal third of distitibia; leg IV distitibia with 16 trichobothria; trichobothrium bc situated closer to sbf than to bf. Charinus bichuetteae is one of ten species of Charinus in Amazonia and northern South America in which the basitibia of leg IV consists of two pseudo-articles; the others are C. bonaldoi, C. brescoviti, C. bromeliaea, C. carvalhoi sp. nov., C. camachoi, C. pardillalensis, C. platnicki, C. quinteroi and C. ricardoi. Charinus bichuetteae differs from C. bromeliaea and C. carvalhoi sp. nov. in the cushionlike gonopods, which are sucker-like in C. bromeliaea and in C. carvalhoi sp. nov. The absence of median eyes and ocular tubercle differentiates C. bichuetteae from C. brescoviti and C. bromeliaea. Seven of these species lack median eyes and median ocular tubercle, and can be differentiated from C. bichuetteae by the number of teeth on the cheliceral claw and the number of spines on the pedipalp femur. Etymology Patronym honoring Dr Maria Elina Bichuette (Giupponi & Miranda 2016). Type material Holotype BRAZIL • ♂; Pará, Vitória do Xingu Municipality, AL 44, Gruta do China; 9 Jul. 2009; D. Pedroso, Pellegatti-Franco, M.E. Bichuette and T.L.C. Scatolini leg.; MNRJ 9204. Paratypes BRAZIL • 1 ♂; same collection data as for holotype; MNRJ 9173 • 1 ♀, Pará, Vitória do Xingu Municipality, AL 19, Paratizão; 10 Jul. 2009; D. Pedroso, F. Pellegatti-Franco, M.E. Bichuette and T.L.C. Scatolini leg.; MNRJ 9174 • 1 ♀, 1 juv. ♀; Altamira Municipality, AL 05, Gruta do Sismógrafo; 05°52′44″ S, 53°39′29″ W; 7 Jul. 2009; D. Pedroso, F. Pellegatti-Franco, M.E. Bichuette and T.L.C. Scatolini leg.; MNRJ 9172 • 3 ♀♀, 1 ♂; Altamira Municipality, B0103 - Caverna Sugiro-Roncador; 2 Jan. 2011; B0026 - river bank, under stone; 3 Apr. 2011; B0049 - Pedra da Cachoeira Cave; 3 Apr. 2011; M.E. Bichuette, J.E. Gallão, D.M. von Schimonsky and D.R. Pedroso leg.; B0026: riverbank, under stone; B0049: Pedra da Cachoeira Cave; MNRJ 9311. Measurements See Table 2. Distribution Known from the type localites. Natural history Specimens were found on cave walls. Remarks See Giupponi & Miranda (2016) for more details concerning the taxonomy of C. bichuetteae.Published as part of Miranda, Gustavo Silva de, Giupponi, Alessandro P. L., Prendini, Lorenzo & Scharff, Nikolaj, 2021, Systematic revision of the pantropical whip spider family Charinidae Quintero, 1986 (Arachnida, Amblypygi), pp. 1-409 in European Journal of Taxonomy 772 on pages 41-42, DOI: 10.5852/ejt.2021.772.1505, http://zenodo.org/record/553641

    La sicurezza urbana tra legislatore e Corte costituzionale

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    Il contributo analizza le conseguenze derivanti dall'introduzione, nel nostro ordinamento, del concetto ambivalente di "sicurezza urbana", approfondendo in particolare i profili problematici in merito agli interventi in materia dei diversi livelli di governo (statale, regionale e locale) e la connessa giurisprudenza costituzionale

    Charinus eleonorae Baptista & Giupponi 2003

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    Charinus eleonorae Baptista & Giupponi, 2003 Fig. 43; Table 4 Charinus eleonorae Baptista & Giupponi, 2003: 80–84, figs 1–14. Charinus eleonorae – Trajano & Bichuette 2010: 12. — Miranda & Giupponi 2011: 66, fig. 13. — Vasconcelos et al. 2013: 496, fig. 12. — Torres-Contreras et al. 2015: 148. — Monte et al. 2015: 6, figs 1, 4. — Vasconcelos & Ferreira 2016: 185. — Miranda et al. 2016c: 19, 29. Diagnosis This species may be separated from other Charinus in eastern South America by means of the following combination of characters: anterior margin of carapace with corners weakly developed into small roundish knobs; median ocular tubercle indistinct; lateral eyes present, but reduced, with flattened lenses; female gonopods small, slightly longer than wide, sucker-like, with rounded opening; cheliceral claw with nine or twelve teeth; pedipalp femur with three dorsal spines and three ventral spines; pedipalp patella with four dorsal spines and three ventral spines; pedipalp tarsus with two long spines similar in length; tibia of leg I with 23 articles, tarsus I with 40 or 41 articles; leg IV basitibia with four pseudo-articles. Etymology Patronym honoring Dr Eleonora Trajano (Baptista & Giupponi 2003). Type material Holotype BRAZIL • ♂; Minas Gerais, Itacarambi, Gruta Olhos d’Água; 15°06′47″ S, 44°10′11″ W; 26 Jun. 2001; R.L.C. Baptista and A.P.L. Giupponi leg.; MNRJ 9070. Paratypes BRAZIL • 6 ♀♀, 7 ♂♂, 2 juv. ♂♂, 2 juv. ♀♀, 2 protonymphs; same collection data as for holotype; MNRJ 9033 • 1 ♂; same collection data as for holotype; MZSP [not examined] • 1 ♂, same collection data as for holotype; CZACC [now IES; L.F. de Armas, pers. com.; not examined] • 1 ♀; same locality as holotype; 10 Jul. 1993; E. Trajano leg.; MNRJ 9024 • 1 ♀, 1 juv.; same locality as for holotype; 30 Mar.–2 Apr. 1994; P. Gnaspini and E. Trajano leg.; MNRJ 9023. Supplementary description CARAPACE.Ten anterior setae; frontal process triangular, not visible in dorsal view. Small granules densely scattered between ocular triads and among sulci. Median eyes reduced, median ocular tubercle absent; seta near remnants of median eyes; lateral eyes reduced, pale, seta posterior to lateral ocular triad. STERNUM. Tritosternum projected anteriorly, with typical setation; other sternal platelets narrow, concave, with pair of setae anteriorly and some smaller setae posteriorly; pentasternum with twelve setae anteriorly and three setae near membranous region. OPISTHOSOMA. Ventral sacs and ventral sac cover absent. GENITALIA. Female genital operculum with prominent setae posteromedially and some smaller setae near margin; posterior margin of genital operculum with pair of shallow lateral sulci; gonopod sucker-like, with short stem and wide opening of genital atrium; gonopods unsclerotized basally. Male gonopod with base of fistula and lateral lobe sclerotized; lateral lobe 2 fimbriate. CHELICERAE. Small tooth projecting slightly from retrolateral surface of basal segment, opposite to bifid tooth; retrolateral surface of claw with two patches of setae, one situated basally, the other medially; claw with 9–12 teeth; transverse row of around ten setae on prolateral surface of basal segment; bifid tooth on basal segment with dorsal cusp larger than ventral cusp. PEDIPALPS. Coxal dorsal carina with three prominent setae encircled by round carina and three setae on margin. Femur with three dorsal spines and four ventral spines; small spine between dorsal spines 2 and 3; two prominent setiferous tubercles between dorsal spine 1 and proximal margin; long spine between ventral spine 1 and proximal margin; small spine between spines 1 and 2 and 2 and 3. Patella with three dorsal spines in primary series; prominent setiferous tubercle distal to spine I, one-third length of spine I; three ventral spines in primary series; long spine between spine I and distal margin. Tibia with ventral spine distally and five setae between spine and distal margin. Tarsus with two dorsal spines, proximal spine half length of distal spine; cleaning organ with 29–30 setae in ventral row. LEGS. Tibia of leg I with 23 articles; tarsus I with 41 articles; first tarsal article shorter than subsequent articles. Leg IV basitibia with four pseudo-articles, without sclerotized, denticulate border at apex of articles; trichobothrium bt situated in proximal third of pseudo-article; distitibia trichobothrium bc situated closer to sbf than to bf, sc and sf series each with six trichobothria. Measurements See Table 4. Distribution Known only from the type localities. Natural history All specimens were collected in the dark zone of the cave; none were found in the dry, upper galleries, where the phrynichid, Trichodamon princeps Mello-Leitão, 1935, was abundant. Charinus eleonorae may be found inside small galleries or holes in the cave walls. Females carry up to 15 eggs and five embryos (Baptista & Giupponi 2003). The cave is experiencing intense human disturbance including the installation of a water pump and construction of a water dam (L.S. Carvalho, pers. com.).Published as part of Miranda, Gustavo Silva de, Giupponi, Alessandro P. L., Prendini, Lorenzo & Scharff, Nikolaj, 2021, Systematic revision of the pantropical whip spider family Charinidae Quintero, 1986 (Arachnida, Amblypygi), pp. 1-409 in European Journal of Taxonomy 772 on pages 108-109, DOI: 10.5852/ejt.2021.772.1505, http://zenodo.org/record/553641

    Charinus mysticus Giupponi & Kury 2002

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    Charinus mysticus Giupponi & Kury, 2002 Fig. 43; Table 4 Charinus mysticus Giupponi & Kury, 2002: 1–4, figs 1–10. Charinus mysticus – Baptista & Giupponi 2003: 80. — Miranda & Giupponi 2011: 66, fig. 13. — Vasconcelos et al. 2013: 496, fig. 12. — Vasconcelos & Ferreira 2016: 185. — Miranda et al. 2016c: 19, 29. Diagnosis This species may be separated from other Charinus in eastern South America by means of the following combination of characters: retrolateral surface of basal segment of chelicera with small blunt tooth; pedipalp tarsus with three spines; leg IV distitibia sc and sf series each with six trichobothria. Etymology Latin adjective, translated from the name of the type locality, which means ‘charmed’ (Giupponi & Kury 2002). Type material Holotype BRAZIL • ♀; Bahia, Cave in Encantados, Gentil do Ouro, 16 km from Santo Inácio, road to Gameleira; 11°25′58.94″ S, 42°30′20.94″ W; MNRJ 9022. Paratype BRAZIL • 1 ♀; same collection data as for holotype; MNRJ 9022. Additional material examined BRAZIL • 1 ♂; Bahia, Central; 11°08′07.62″ S, 42°06′41.43″ W; MNRJ 9289 • 1 ♂; Bahia, Central, Gruta Rolling Stones; 7 Jul. 2007; M.E. Bichuette leg.; MNRJ 9291 • 1 ♂; same collection data as for preceding; MNRJ 9290. Supplementary description CARAPACE. Frontal process large, subtriangular, visible in dorsal view; distinct curved carina between ocular triads and margin of carapace. Median eyes and median ocular tubercle present; median ocular tubercle shallow, slightly higher than carapace surface, with pair of small setae. Lateral eyes well developed, pale, small seta posterior to each lateral ocular triad. STERNUM. Tritosternum projected anteriorly into large, blunt tubercle, surpassing base of pedipalp coxae, and rounded posteriorly. Medial platelet (tetrasternum) and third platelet (pentasternum) forming single convex platelet, with pair of large setae anteriorly, and several small setae posteriorly; metasternum with two anterior setae at border between membranous and sclerotized region, and three setae in longitudinal row, from anterior to posterior; four to six setae posteriorly. OPISTHOSOMA. Ventral sacs and ventral sac cover absent. GENITALIA. Female genital operculum posterior margin slightly convex, with several setae along margin and on surface. Gonopods unsclerotized basally; gonopods sucker-like with small invagination anteriorly. CHELICERAE. Small tooth in retrolateral row of basal segment. Prolateral surface with transverse row of around six small setae, ventrally to dorsally. Four setae, retrolateral, retromedial, prolateral and promedial, on dorsodistal margin, near membranous region of claw. Claw with eleven denticles and row of setae on retrolateral surface from base to near apex (dorsal side). PEDIPALPS. Coxal dorsal carina with two or three small setae and four setae on anterior margin of round carina. Femur with three distinct setiferous tubercles proximal to spine 1; primary series with five dorsal spines, and smaller spine between spines 2 and 3; small spine, similar in length to spine 4 (suggesting it was duplicated), present between spines 3 and 4 in holotype female (MNRJ 9022), as on sinistral pedipalp of C. diamantinus sp. nov.; four or five ventral spines, small spine between spines 1 and 2, and 2 and 3; large ventral spine proximal to spine 1. Patella with five dorsal spines; large setiferous tubercle distal to spine I, about one-third length of spine I; four ventral spines decreasing in size; distinct setiferous tubercle between spines II and III; two or three setiferous tubercles between spine I and distal margin. Tibia with two dorsal spines, proximal spine two-thirds length of distal spine; ventral spine situated in distal half of tibia; three setiferous tubercles in row near base of spine; six or seven long setae between spine and distal margin. Tarsus with three dorsal spines; ventral row of cleaning brush with 26 or 27 setae. LEGS. Tibia of leg I with 23 articles; tarsus I with 41 articles. Leg IV basitibia with four pseudoarticles; trichobothrium bt situated in proximal third; distal apex of basitibial pseudo-articles with dark, denticulate projection. Leg IV distitibia trichobothrium bc situated closer to sbf than to bf; sc and sf each with six trichobothria; tarsus with distinct white annulus distally on first article. Measurements See Table 4. Distribution Known from two caves in the western part of the Brazilian state of Bahia. Natural history Holotype and paratypes were collected 8 m from the entrance of the cave. The behavioral repertoire of the species was studied by Tagliatela et al. (2019).Published as part of Miranda, Gustavo Silva de, Giupponi, Alessandro P. L., Prendini, Lorenzo & Scharff, Nikolaj, 2021, Systematic revision of the pantropical whip spider family Charinidae Quintero, 1986 (Arachnida, Amblypygi), pp. 1-409 in European Journal of Taxonomy 772 on pages 120-122, DOI: 10.5852/ejt.2021.772.1505, http://zenodo.org/record/553641

    Charinus carajas Giupponi & Miranda 2016

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    <i>Charinus carajas</i> Giupponi & Miranda, 2016 <p>Figs 8E–H, 27; Table 2</p> <p> <i>Charinus carajas</i> Giupponi & Miranda, 2016: 15–18, figs 2e, 3e, 4e, g, 8a–f.</p> Diagnosis <p> Based on Giupponi & Miranda (2016), this species may be separated from other <i>Charinus</i> in Amazonia and northern South America by means of the following combination of characters: median and lateral eyes present; median ocular tubercle and lateral eyes markedly reduced; small, rounded tetra- and pentasternum; reduced tritosternum, slightly surpassing base of pedipalp coxae; cushion-like female gonopod with lateral projections directed posteriorly, covering openings of internal seminal receptacles (atrium); male gonopods with long, curved, wrinkled medial lobes (Fig. 8E–F); lateral lobes fimbriate (Fig. 8E–F, H); dorsal lobe surpassing length of other lobes and with elevated scales (Fig. 8E, G); marked secondary sexual dimorphism evident, males with longer pedipalps, ca twice as long as female pedipalps; pedipalp femur with four dorsal spines; two dorsal spines on pedipalp tarsus, small basal spine, one-quarter length of distal spine; tibia of leg I with 23 articles, tarsus I with 42 articles; leg IV basitibia with three pseudo-articles; leg IV basitibia trichobothrium <i>bt</i> situated in proximal third; leg IV distitibia <i>sc</i> and <i>sf</i> series each with five trichobothria; leg IV distitibia trichobothrium <i>bf</i> equidistant between <i>bc</i> and <i>sbf</i>; tegument pale yellow.</p> <p>This is among the few Amazonian species in which the median eyes and median ocular tubercle are present.</p> Etymology <p> Noun in apposition, taken from the Caraja mountain range where the species was collected, and from the indigenous people known as the <i>karajas</i> (Giupponi & Miranda 2016).</p> Type material <p> <b>Holotype</b> BRAZIL • 1 ♀; Pará, Carajás National Forest, Gruta N4E-14 AF; 06°02′2.096″ S, 50°10′03.741″ W; SAD’ 69; 7–12 Oct. 2008; Andrade <i>et al.</i> leg.; MZSP 29136.</p> <p> <b>Paratypes</b> BRAZIL • 1 juv.; same locality as for holotype, Gruta N4E-10; 20 Oct.–1 Nov. 2006; Andrade <i>et al</i>. leg.; MZSP28287 • 1juv.;same locality as for holotype, Gruta N4E-21; 20 Oct.–1Nov.2006; Andrade <i>et al</i>. leg.; MZSP28288 • 2juv.;same locality as for holotype, Gruta N4E-22; 20 Oct.–1Nov.2006; Andrade <i>et al</i>. leg.; MZSP 28289 • 1 ♂; same locality as for holotype, Gruta N4E-22; 06°02′2.096″ S, 50°10′03.741″ W; SAD’ 69; 7–12 Oct. 2008; MZSP 29126 • 2 juv.; same locality as for holotype, Gruta N4E-33; 8–12 Feb. 2007; Andrade <i>et al</i>. leg.; MZSP 28290 • 1 ♂; same locality as for holotype, Gruta N4E-95; 7–12 Oct. 2008; Andrade <i>et al</i>. leg.; MZSP 29127 • 2 juv.; same locality as for holotype, Gruta N4WS-15; 20 Oct.</p> <p> 2006; Andrade <i>et al.</i> leg.; MZSP 28292 • 2 ♀♀, 1 ♂, 2 juv.; same locality as for holotype, Gruta N4WS8- CL; 06°05′22.183″ S, 50°11′40.996″ W; SAD’ 69; 7–12 Oct. 2008; MNRJ 9270 ex MZSP 29132 • 1 juv.; same locality as for holotype, Gruta N5E-03 (new -07); 22 Mar.–3 Apr. 2005; Andrade and Arnoni leg.; MZSP 28281 • 3 juv.; same locality as for holotype, Gruta N5E-04 (new -06); 22 Mar.–3 Apr. 2005; Andrade and Arnoni leg.; MZSP 28282 • 1 juv.; same locality as for holotype, Gruta N5E-05 (new -04); 22 Mar.–3 Apr. 2005; Andrade and Arnoni leg.; MZSP 28283 • 1 juv.; same locality as for holotype, Gruta N5S; 3 May 2005; Andrade and Arnoni leg.; MZSP 28286 • 1 juv.; same locality as for holotype, Gruta N5S-01; 3–13 May 2005; Andrade and Arnoni leg.; MZSP 28285 • 1 ♀; same locality as for holotype; MII GEM 1722 EU; ISLA 3892 • 1 ♀, 1 ♂; same locality as for holotype; MII GEM 1755EU; ISLA 3890 • 2 ♀♀, 1 ♂; Parauapebas, Projeto N5S, Morro I, Cav. GEM, 1782; 28 Feb. 2011; CARSTE leg.; dry season; ISLA 3897.</p> <p> <b>Additional material</b> (examined)</p> <p> BRAZIL • 1 ♂; same locality as for holotype, Gruta GEM 1564 CL Tarzan; 06°19′04.284″ S, 50°05′48.261″ W; SAD’ 69; 17–24 Oct. 2008; Andrade <i>et al</i>. leg.; MZSP 29120 • 1 juv.; same locality as for holotype, Gruta GEM 1590 CL Tarzan; 17–24 Oct. 2008; Andrade <i>et al</i>. leg.; MZSP 29123 • 1 ♂, 3 juv.; same locality as for holotype, Gruta GEM15170, AF Tarzan; 17– 24 Oct. 2008; Andrade <i>et al</i>. leg.; MZSP 29121 • 1 juv.; same locality as for holotype, Gruta N1- 15, Mangangá, 28 Sep.–3 Oct. 2007; Andrade <i>et al</i>. leg.; MZSP 29115 • 1 juv.; same locality as for holotype, Três Mosqueteiras, Gruta N1-25; 28 Sep.–3 Oct. 2007; Andrade <i>et al</i>. leg.; MZSP 29113 • 1 ♀; same locality as for holotype; Mangangá, Gruta N1-15; 28 Sep.–3 Oct. 2007; Andrade <i>et al</i>. leg.; MZSP 29116 • 1 ♀; Gruta N1-64 Amailton; 28 Sep.–3 Oct. 2007; Andrade <i>et al</i>. leg.; MZSP 29114 • 1 ♀, 3 juv; same locality as for holotype, Gruta N4E-61; 8–12 Feb. 2007; Andrade <i>et al</i>. leg.; MZSP 28291 • 1 juv.; same locality as for holotype, Gruta N4E-08 CL; 06°02′20.419″ S, 50°09′36.064″ W; SAD’ 69; 7–12 Oct. 2008; Andrade <i>et al</i>. leg.; MZSP 29134 • 1 juv.; same locality as for holotype, Gruta N4E-10 AF; 06°02′20.162″ S, 50°09′38.277″ W; SAD’ 69; 7–12 Oct. 2008; Andrade <i>et al</i>. leg.; MZSP 29129 • 1 juv.; same locality as for holotype, Gruta N4E-22 AF; 06°02′02.096″ S, 50°10′03.741″ W; SAD’ 69; 7–12 Oct. 2008; MZSP 29133 • 1 ♂; same locality as for holotype, Gruta N4E-26 CL; 06°02′14.273″ S, 50°10′02.194″ W; SAD’ 69; 7–12 Oct. 2008; Andrade <i>et al</i>. leg.; MZSP 29128 • 1 ♂; same locality as for holotype, Gruta N4E-33 AF; 06°02′26.607″ S, 50°09′36.64″ W; SAD’ 69; 7–12 Oct. 2008; Andrade <i>et al</i>. leg.; MZSP 29125 • 1 ♂; same locality as for holotype, Gruta N4E-61 AF; 06°02′20.754″ S, 50°10′02.965″ W; SAD’ 69; 7–21 Oct. 2008; Andrade <i>et al</i>. leg.; MZSP 29135 • 1 juv.; same locality as for holotype, Gruta N4WS8 AF; 06°05′22.183″ S, 50°11′40.996″ W; SAD’ 69; 7–12 Oct. 2008; Andrade <i>et al</i>. leg.; MZSP 29137 • 1 ♀; same locality as for holotype, Gruta N5E-11 (new -21); 22 Mar.–3 Apr. 2005; Andrade and Arnoni leg.; MZSP 28284 • 1 juv.; same locality as for holotype, Gruta N5S21 AF; 06°05′15.383″ S, 50°07′33.51″ W; SAD’ 69; 7–12 Oct. 2008; Andrade <i>et al</i>. leg.; MZSP 29130 • 1 ♂; same locality as for holotype, Gruta N5S8 CL; 06°05′51.597″ S, 50°07′56.646″ W; SAD’ 69; 7–12 Oct. 2008; Andrade <i>et al</i>. leg.; MZSP 21131.</p> Measurements <p>See Table 2.</p> Distribution <p>Known from several iron caves in the northern part of Serra de Carajas, Pará, Brazil.</p> Natural history <p>Troglophilous, only known from inside iron ore caves.</p> Remarks <p>See Giupponi & Miranda (2016) for more details about this species.</p>Published as part of <i>Miranda, Gustavo Silva de, Giupponi, Alessandro P. L., Prendini, Lorenzo & Scharff, Nikolaj, 2021, Systematic revision of the pantropical whip spider family Charinidae Quintero, 1986 (Arachnida, Amblypygi), pp. 1-409 in European Journal of Taxonomy 772</i> on pages 51-53, DOI: 10.5852/ejt.2021.772.1505, <a href="http://zenodo.org/record/5536410">http://zenodo.org/record/5536410</a&gt
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