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    Miridiba (Miridiba) bannaensis Gao & Bai & Fang & Yu 2018, new species

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    Miridiba (Miridiba) bannaensis Gao & Fang, new species Type locality. “ China: Yunnan Province: Xishuangbanna City ”. Distribution. China (Yunnan).Published as part of Gao, Chuan-Bu, Bai, Ming, Fang, Hong & Yu, Zhi-Guo, 2018, Four new species of the genus Miridiba Reitter (Coleoptera: Scarabaeidae: Melolonthinae) from China, pp. 1-20 in Zootaxa 4527 (1) on page 13, DOI: 10.11646/zootaxa.4527.1.1, http://zenodo.org/record/261188

    Miridiba (Miridiba) youweii Gao & Bai & Fang & Yu 2018, new species

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    Miridiba (Miridiba) youweii Gao & Fang, new species Type locality. “ China: Yunnan Province: Jingdong Yi Autonomous County ”. Distribution. China (Guizhou, Yunnan, Zhejiang).Published as part of Gao, Chuan-Bu, Bai, Ming, Fang, Hong & Yu, Zhi-Guo, 2018, Four new species of the genus Miridiba Reitter (Coleoptera: Scarabaeidae: Melolonthinae) from China, pp. 1-20 in Zootaxa 4527 (1) on page 18, DOI: 10.11646/zootaxa.4527.1.1, http://zenodo.org/record/261188

    Miridiba (Miridiba) kuatunensis Gao & Bai & Fang & Yu 2018, new species

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    Miridiba (Miridiba) kuatunensis Gao & Fang, new species Type locality. “ China: Fujian Province: Wuyishan Mountain ”. Distribution. China (Fujian, Guangdong, Guangxi, Hainan, Zhejiang).Published as part of Gao, Chuan-Bu, Bai, Ming, Fang, Hong & Yu, Zhi-Guo, 2018, Four new species of the genus Miridiba Reitter (Coleoptera: Scarabaeidae: Melolonthinae) from China, pp. 1-20 in Zootaxa 4527 (1) on page 13, DOI: 10.11646/zootaxa.4527.1.1, http://zenodo.org/record/261188

    Miridiba xingkei Gao & Fang 2018

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    Miridiba xingkei Gao & Fang, 2018 Miridiba xingkei Gao & Fang in Gao et al., 2018: 6, figs 3, 6 (type loc.: Xishuangbanna, Yunnan Province, China). Miridiba xingkei – Gao et al. 2019: 462 (species list; in key). Diagnosis See Gao et al. (2018). Remarks Gao et al. (2018) described Miridiba xingkei based on the male holotype and thirteen paratypes (4 males and 9 females) conserved in IZCAS and NWAFU of China. This species shows features that characterize genital morphotype IX “ Ciliatipennis ”. The parameres have the characteristic dorsal tubular complex (Gao et al. 2018: figs 3g –i) with dorsal and ventral branches (upper and lower parts) (Gao et al. 2018: fig. 3h) forked at distal end (Gao et al. 2018: fig. 3g), showing in lateral view the distinctive concavity at distal end (Gao et al. 2018: fig. 3h). On the other hand, the ventral branches appear reduced but obvious, fused laterally to dorsal tubular complex forming an apparent lateral scar (Gao et al. 2018: figs 3h–i). According to these features, we include M. xingkei in the genital morphotype IX “ Ciliatipennis ”. Distribution China (Yunnan).Published as part of Gao, Chuan-bu & Coca-Abia, María Milagro, 2021, Revision of the genus Miridiba Reitter, 1902 (Coleoptera, Scarabaeidae, Melolonthinae): genital morphotypes and new taxonomic data, pp. 1-94 in European Journal of Taxonomy 749 on page 80, DOI: 10.5852/ejt.2021.749.1355, http://zenodo.org/record/477029

    Miridiba youweii Gao & Fang 2018

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    Miridiba youweii Gao & Fang, 2018 Miridiba youweii Gao & Fang in Gao et al., 2018: 9, figs 4, 6 (type loc.: Jingdong, Yunnan, China). Miridiba youweii – Gao et al. 2019: 462 (species list; in key). Diagnosis See Gao et al. (2018). Remarks Gao et al. (2018) described Miridiba youweii based on the male holotype and 110 paratypes (71 males and 39 females) conserved in IZCAS. Antenna 9-segmented, pronotal surface densely punctate and parameres with two dorsal and two ventral branches (Gao et al. 2018: fig. 4) allow us to include M. youweii in morphotype I “ Trichophora ”. Distribution China (Guizhou, Yunnan, Zhejiang) Species included in Morphotype II “Gressitti” The group is composed of two species, which are characterized by antenna 10-segmented, pronotal surface scattered punctate (distance between punctures greater than diameter of puncture), and male and female genitalia as specified above.Published as part of Gao, Chuan-bu & Coca-Abia, María Milagro, 2021, Revision of the genus Miridiba Reitter, 1902 (Coleoptera, Scarabaeidae, Melolonthinae): genital morphotypes and new taxonomic data, pp. 1-94 in European Journal of Taxonomy 749 on page 47, DOI: 10.5852/ejt.2021.749.1355, http://zenodo.org/record/477029

    Miridiba (Miridiba) xingkei Gao & Bai & Fang & Yu 2018, new species

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    Miridiba (Miridiba) xingkei Gao & Fang, new species (Figs. 3, 6) Type locality. CHINA: Yunnan Province: Xishuangbanna City. Type material examined. Holotype, male, labelled: “ Menga, Xishuangbanna, Yunnan / 1050–1180 m / Chinese Academy of Sciences [typeset, Chinese] // 1958. V. 13 / Collector Pu Fuji [typeset, Chinese] // Holotrichia (P.) ciliatipennis Moser ♂ / Designed by You-wei Zhang [handwritten, Chinese] // HOLOTYPE ♂ / Miridiba xingkei sp. nov. / Chuanbu Gao & Hong Fang / det. 2018 [typeset, red label]”, deposited in IZCAS; paratype, female: “Jingdong, Yunnan 1200 m / 1957. IV. 29 / A. Monchadsky [typeset, Chinese and Russian] // PARATYPE ♀ / Miridiba xingkei sp. nov. / Chuanbu Gao & Hong Fang / det. 2018 [typeset, yellow label]”, deposited in IZCAS; paratype, male: “Jingdong 1170 m / 1956. V. 26 / Zagulyaev lamp [typeset, Chinese and Russian] // PARATYPE ♂ / Miridiba xingkei sp. nov. / Chuanbu Gao & Hong Fang / det. 2018 [typeset, yellow label]”, deposited in IZCAS; paratype, female: “Xiaomengyang, Yunnan 850 m / 1957. V. 3 / Qiuzhen Liang [typeset, Chinese and Russian] // Holotrichia (P.) ciliatipennis Moser ♀ / Identified by Youwei Zhang 1981. VI [handwritten, Chinese] // PARATYPE ♀ / Miridiba xingkei sp. nov. / Chuanbu Gao & Hong Fang / det. 2018 [typeset, yellow label]”, deposited in IZCAS; paratype, male: “Yun Jinghong, Xishuangbanna, Yunnan / 650 m / Chinese Academic of Sciences [typeset, Chinese] // 1959-?-5 / Collector Xuezhong Zhang [typeset] // H. ciliatipennis Moser / genitalia / 62.3.19 [handwritten] // Holotrichia ciliatipennis Moser / Identified by Youwei Zhang [handwritten, Chinese] // IOZ(E)1967842 [typeset] // PARATYPE ♂ / Miridiba xingkei sp. nov. / Chuanbu Gao & Hong Fang / det. 2018 [typeset, yellow label]”, deposited in IZCAS; paratype, male: “Yun Jinghong, Xishuangbanna, Yunnan / 650 m / Chinese Academic of Sciences [typeset, Chinese] // 1958-V-11 / Collector Xuewu Meng [typeset] // Holotrichia ciliatipennis Moser / Identified by Youwei Zhang [handwritten, Chinese] // IOZ(E)1967836 [typeset] // PARATYPE ♂ / Miridiba xingkei sp. nov. / Chuanbu Gao & Hong Fang / det. 2018 [typeset, yellow label]”, deposited in IZCAS; paratype, female: “Yunpenghun, Xishuangbanna, Yunnan / 1200- 1400 m / Chinese Academic of Sciences [typeset, Chinese] // 1958-V-11 / Collector Xuewu Meng [typeset] // PARATYPE ♀ / Miridiba xingkei sp. nov. / Chuanbu Gao & Hong Fang / det. 2018 [typeset, yellow label]”, deposited in IZCAS; paratypes, 3 females: “Jingdong, Yunnan 1200 m / 1957. IV. 25 / A. Monchadsky [typeset, Chinese and Russian] // PARATYPE ♀ / Miridiba xingkei sp. nov. / Chuanbu Gao & Hong Fang / det. 2018 [typeset, yellow label]”, deposited in IZCAS; paratype, female: “Simao, Yunnan 1300 m / 1957. V. 9 / Fuji Pu [typeset, Chinese and Russian] // PARATYPE / Miridiba xingkei sp. nov. / Chuanbu Gao & Hong Fang / det. 2018 [typeset, yellow label]”, deposited in IZCAS; paratype, male: “Jingdong 1170 m / 1956.?. 1 / Zagulyaev lamp [typeset, Chinese and Russian] // PARATYPE ♂ / Miridiba xingkei sp. nov. / Chuanbu Gao & Hong Fang / det. 2018 [typeset, yellow label]”, deposited in IZCAS; paratype, female: “ Yunnan Xishuangbanna Menghai / 1982-?- 16 / Wang Sumei, Zhou Jingruo [handwritten, Chinese] // C586 [handwritten] // PARATYPE ♀ / Miridiba xingkei sp. nov. / Chuanbu Gao & Hong Fang / det. 2018 [typeset, yellow label]”, deposited in NWAUF; paratypes, 2 females: “ Yunnan, Xishuangbanna, Menglong / Miridiba xingkei sp. nov. / Chuanbu Gao & Hong Fang / det. 2018 [typeset, yellow label]”, deposited in NWAUF Description of holotype (male). Body length 22.8 mm; width across humeri 9.8 mm, oval-elongate, strongly convex, dorsal surface glabrous. Colour: head, pronotum, scutellum and legs dark reddish brown; antennae, elytra and abdomen brown (Figs. 3a, b). Head: Frons densely punctate and glabrous; clypeus arcuate shaped, apex weakly emarginate medially; anterior margin moderately reflexed; fronto-clypeal suture wave-shaped; frontal carina distinctly raised and nearly straight; clypeus shorter than frons, length ratio between clypeus and frons before carina 0.60; distance between eyes nearly four times wider than eye (Fig. 3c). Antenna with 9 antennomeres; club with 3 antennomeres, slightly longer than antennomeres 2-6 combined (Fig. 3d). Thorax: Dorsal surface of pronotum densely punctate and glabrous, widest at basal 2/5; anterior margin smooth and flanged; lateral margin smooth and broadly reflexed; posterior margin smooth; anterolateral angles obtuse, posterior angles obtuse and round. Prosternal process basiconic shaped. Scutellum triangular, dorsal surface densely punctate and glabrous, 1.58 times wider than long. Dorsal surface of elytra glabrous, sutural costae developed; epipleuron with long, dense cilia (Figs. 3a, b). Legs: Protibia tridentate, denticle 2 sharp; dorsal carina forking to denticle 2; apical spur of protibia sharp, apical spur reaching basal one half of protarsomere 1. Metafemora moderately covered with long soft hairs near anterior margin and rough setae near posterior margin (Fig. 3b); outside of metatibia with 1 completed carina at distal 2/5, carina arc-shaped; dorsolateral margin with 2 thorns; dorsomedial margin with 5 pubescent thorns small to large; metatibia with two apical spurs of different sizes, the big spur acinaciform, 1.72 times longer than the small spur (Fig. 3e); metatarsomere 1 equal in length to metatarsomere 2. Abdomen: Lateral sides of abdominal sternites densely punctate and setiferous, setae short; posterior sternites 5 and 6 covered with long soft hairs. Pygidium: Fan-shaped pygidium densely punctate and setiferous, 1.37 times wider than long, apical pygidium at a right angle; middle area weakly convex. Male genitalia. Phallobase: Phallobase 1.2 times longer than parameres; dorsal surface with a depressed longitudinal sulcus at middle; anterior 1/5 of phallobase weakly constricted. Parameres: Parameres with two pairs of branches: two thick long dorsal branches and two short ventral branches; each dorsal branch with a thick protuberance on dorsum; tip of each dorsal branch with a notch in the ventral side; ventral branch with a big coneshaped hook ventrally before the apex (Figs. 3 g–i). Endophallus: Dorsal endophallus with temones tapered; medial endophallus with small hooks; distal end of endophallus with a round seminal vesicle (Fig. 3j). Female. Body slightly longer and broader than in male. Antenna with 9 antennomeres; club with 3 antennomeres, shorter than antennomeres 2–6 combined. Metafemora and apex of metatibia broader than in male. The big spur of apical metatibia broader and blunter than male, 1.3 times longer than the smaller spur (Fig. 3f). Variability. All the paratypes slightly differ in size (total body length 21.7–25.6 mm, width across humeri 9.5– 11.6 mm). Diagnosis. Miridiba (M.) xingkei new species and Miridiba (M.) ciliatipennis (Moser, 1913) are similar in the elytral epipleuron bearing dense, long cilia. The two species can be separated by following characters: the presence or absence of the midline on the posterior of the pronotum (absent in M. (M.) xingkei new species; present in M. (M.) ciliatipennis), the form of setae near the posterior margin of the metafemora (few large setae in M. (M.) xingkei new species, dense small setae in M. (M.) ciliatipennis), the shape of the spurs of the apical metatibia (arcshaped in M. (M.) xingkei new species, straight in M. (M.) ciliatipennis), the form of the parameres (each dorsal branch with a developed protuberance in M. (M.) xingkei new species, the dorsal branches without protuberances in M. (M.) ciliatipennis). Distribution. China (Yunnan: Puer, Xishuangbanna) (Fig. 6). Etymology. This new species is dedicated to Dr Xingke Yang, a respected researcher, who has devoted himself into taxonomy and systematics study of Chinese Coleoptera for more than thirty years. He will retire from the Institute of Zoology, Chinese Academy of Sciences in 2018. Chinese name. 杨脊鳃金龟 Remarks. Chang (1964) listed M. (M.) ciliatipennis from China. However, we examined all specimens in IZCAS identified as M. (M.) ciliatipennis by Chang himself and they all belong to the new species M. (M.) xingkei. Based on this, we have excluded M. (M.) ciliatipennis from the Chinese fauna.Published as part of Gao, Chuan-Bu, Bai, Ming, Fang, Hong & Yu, Zhi-Guo, 2018, Four new species of the genus Miridiba Reitter (Coleoptera: Scarabaeidae: Melolonthinae) from China, pp. 1-20 in Zootaxa 4527 (1) on pages 6-9, DOI: 10.11646/zootaxa.4527.1.1, http://zenodo.org/record/261188

    Miridiba bannaensis Gao & Fang 2018

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    <i>Miridiba bannaensis</i> Gao & Fang, 2018 <p> <i>Miridiba bannaensis</i> Gao & Fang in Gao <i>et al</i>., 2018: 2, figs 1, 6 (type loc.: Xishuangbanna, Yunnan, China).</p> <p> <i>Miridiba bannaensis</i> – Gao <i>et al</i>. 2019: 461 (species list; in key).</p> Diagnosis <p> See Gao <i>et al</i>. (2018).</p> Remarks <p> Gao <i>et al</i>. (2018) described <i>Miridiba bannaensis</i> based on one male holotype and three paratypes (one male and two females) preserved in IZCAS. The antenna 9-segmented, pronotal surface densely punctate and parameres with two dorsal and two ventral branches (Gao <i>et al</i>. 2018: fig. 1) allow us to include <i>M. bannaensis</i> in genital morphotype I “ <i>Trichophora</i> ”.</p> Distribution <p>China (Yunnan).</p>Published as part of <i>Gao, Chuan-bu & Coca-Abia, María Milagro, 2021, Revision of the genus Miridiba Reitter, 1902 (Coleoptera, Scarabaeidae, Melolonthinae): genital morphotypes and new taxonomic data, pp. 1-94 in European Journal of Taxonomy 749</i> on page 19, DOI: 10.5852/ejt.2021.749.1355, <a href="http://zenodo.org/record/4770293">http://zenodo.org/record/4770293</a&gt

    Miridiba lamellata Gao & Fang 2019

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    <i>Miridiba lamellata</i> Gao & Fang, 2019 <p> <i>Miridiba</i> (<i>Pledina</i>) <i>lamellata</i> Gao & Fang in Gao <i>et al.</i>, 2019: 459, figs 1–18 (type loc.: Gui Yang city, Guizhou Province, China).</p> Diagnosis <p> See Gao <i>et al</i>. (2019).</p> Remarks <p> Gao <i>et al</i>. (2019) described <i>Miridiba</i> (<i>Pledina</i>) <i>lamellata</i> based on the male holotype only, deposited in the collection of Shenyang Agricultural University of China. This species was included in the subgenus <i>Pledina</i> based on the antenna 10-segmented (Gao <i>et al</i>. 2019), without specifying the features of <i>M. lamellata</i> shared with <i>M. sinensis</i> (type species of <i>Pledina</i>), which would justify considering this new species in the subgenus. We have studied herein <i>M. sinensis</i> that, together with <i>M. pseudosinensis</i>, <i>M. axanensis</i> and <i>M. quasisinensis</i>, constitute the morphotype V “ <i>Sinensis</i> ”, candidate to be subgenus <i>Pledina</i> (see Discussion and Conclusions), and we have not found enough similarities to consider <i>M. lamellata</i> belonging to morphotype V “ <i>Sinensis</i> ”. The antenna 10-segmented, the only character considered by Gao <i>et al</i>. (2019) to classify this new species in the subgenus <i>Pledina</i>, is shared by many other <i>Miridiba</i> species, those included herein in morphotypes II to VIII, and even by many other species of <i>Holotrichia</i>. Therefore, this character is not enough to consider <i>M. lamellata</i> belonging to <i>Pledina</i>. In addition to that, this new species can be separated from <i>M. sinensis</i> by the branches of parameres and setae on mentum and pygidium (Gao <i>et al</i>. 2019: figs 9, 14–17), both species are so different that <i>M. lamellata</i> cannot be considered belonging to <i>Pledina</i>. Likewise, the mentum and labrum of <i>M. lamellata</i> differ from those of <i>Miridiba</i>. Unlike <i>Miridiba</i>, the new species has the mentum with sparse pubescence on the anterior part, and the labrum more strongly depressed at its ventral part than at dorsal one (Gao <i>et al</i>. 2019: figs 5, 9). Therefore, the second author of this paper has doubts about inclusion of <i>M. lamellata</i> in the genus <i>Miridiba</i>. However, taking into account that the first author of this paper is one of the authors of the species and the second author has not studied it, <i>M. lamellata</i> is considered a species incertae sedis of <i>Miridiba</i>.</p> Distribution <p>China (Guizhou).</p>Published as part of <i>Gao, Chuan-bu & Coca-Abia, María Milagro, 2021, Revision of the genus Miridiba Reitter, 1902 (Coleoptera, Scarabaeidae, Melolonthinae): genital morphotypes and new taxonomic data, pp. 1-94 in European Journal of Taxonomy 749</i> on pages 82-83, DOI: 10.5852/ejt.2021.749.1355, <a href="http://zenodo.org/record/4770293">http://zenodo.org/record/4770293</a&gt

    Yongzhou xiao ji: 3 juan / Gao Zhengchen. Xiangshan jiu lao hui shi / [Bai Juyi ji]. Gu yang yi xiang ji / Wen Tong.

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    陶弼. 方叔淵遺稿 / 方瀾. 高氏三宴詩 :Tao Bi. Fang Shuyuan yi gao / Fang Lan. Gao shi san yan shi

    Figs. 19–22. Miridiba sinensis. 19 in Miridiba (Pledina) Lamellata Gao and Fang (Coleoptera: Scarabaeidae), a New Species from Guizhou, China, and a Key to the Chinese Species in the Genus

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    Figs. 19–22. Miridiba sinensis. 19) Mentum; 20) Pygidium; 21) Parameres, dorsal view; 22) Parameres, lateral view. Scale bar = 1 mm.Published as part of Gao, Chuan-Bu, Li, Yan & Fang, Hong, 2019, Miridiba (Pledina) Lamellata Gao and Fang (Coleoptera: Scarabaeidae), a New Species from Guizhou, China, and a Key to the Chinese Species in the Genus, pp. 458-464 in The Coleopterists Bulletin 73 (2) on page 462, DOI: 10.1649/0010-065X-73.2.458, http://zenodo.org/record/537614
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