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    Chrysotus laesus

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    Chrysotus laesus (Wiedemann, 1817) Material examined JAPAN: 29 ƋƋ, Honshu, Aichi Prefecture, Nagoya, Nagakute-cho Park, 23–30 May 1999, leg. V. Fursov. Distribution Western Europe, Armenia, Georgia, Japan and Russia (Karelia, Saint Petersburg, Pskov, Yaroslavl, Moscow, Ryasan, Bashkiria, Tatarstan, Kursk, Lipetsk, Voronezh, Krasnodar, Adygea, North Caucasus, Orenburg, Omsk, Tomsk, Altai, Krasnojarsk, Irkutsk, Buryatia, Yakutia, Amur).Published as part of Negrobov, Oleg P., Kumazawa, Tatsunori, Tago, Toshihiro & Fursov, Viktor N., 2016, Species of the genus Chrysotus Meigen, 1824 (Diptera: Dolichopodidae) from Japan, with descriptions of two new species, pp. 1-15 in European Journal of Taxonomy 197 on page 4, DOI: 10.5852/ejt.2016.197, http://zenodo.org/record/383796

    Fig. 1 in Species of the genus Chrysotus Meigen, 1824 (Diptera: Dolichopodidae) from Japan, with descriptions of two new species

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    Fig. 1. Chrysotus masunagai Negrobov, Kumazawa & Tago sp. nov., Ƌ, paratype, coll. 28 Apr. 2012. A. Hypopygium, lateral view. B. Apex of surstylus, lateral view. C. Antenna. D. Apex of phallus, lateral view. E. Cerci, lateral view.Published as part of Negrobov, Oleg P., Kumazawa, Tatsunori, Tago, Toshihiro & Fursov, Viktor N., 2016, Species of the genus Chrysotus Meigen, 1824 (Diptera: Dolichopodidae) from Japan, with descriptions of two new species, pp. 1-15 in European Journal of Taxonomy 197 on page 5, DOI: 10.5852/ejt.2016.197, http://zenodo.org/record/383796

    Chrysotus Meigen 1824

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    Genus Chrysotus Meigen, 1824 We found the following six species in the collection of Mr. T. Tago and of the Schmalhausen Institute of Zoology of National Academy of Sciences Ukraine, including a species new to Japan and two species new to science.Published as part of Negrobov, Oleg P., Kumazawa, Tatsunori, Tago, Toshihiro & Fursov, Viktor N., 2016, Species of the genus Chrysotus Meigen, 1824 (Diptera: Dolichopodidae) from Japan, with descriptions of two new species, pp. 1-15 in European Journal of Taxonomy 197 on page 3, DOI: 10.5852/ejt.2016.197, http://zenodo.org/record/383796

    Fig. 5. Male habitus, lateral view. A in Species of the genus Chrysotus Meigen, 1824 (Diptera: Dolichopodidae) from Japan, with descriptions of two new species

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    Fig. 5. Male habitus, lateral view. A. Chrysotus masunagai Negrobov, Kumazawa & Tago sp. nov., Ƌ, paratype, coll. 15 May 2010. B. Chrysotus saigusai Negrobov, Kumazawa & Tago sp. nov., Ƌ, paratype, coll. 8 Jul. 2012.Published as part of Negrobov, Oleg P., Kumazawa, Tatsunori, Tago, Toshihiro & Fursov, Viktor N., 2016, Species of the genus Chrysotus Meigen, 1824 (Diptera: Dolichopodidae) from Japan, with descriptions of two new species, pp. 1-15 in European Journal of Taxonomy 197 on page 11, DOI: 10.5852/ejt.2016.197, http://zenodo.org/record/383796

    Fig. 4. A–B in Species of the genus Chrysotus Meigen, 1824 (Diptera: Dolichopodidae) from Japan, with descriptions of two new species

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    Fig. 4. A–B. Chrysotus tagoi Negrobov, Maslova & Fursov, 2015, Ƌ, paratype, Japan (Honshu, Ibaraki Pref., Tsukuba), coll. 16. Jul. 1997 by V. Fursov. A. Hypopygium, lateral view. B. Apex of phallus, lateral view. –– C–D. Chrysotus parilis Parent, 1926, Russia (Primorje). C. Hypopygium, lateral view. D. Apex of phallus, lateral view. –– E–F. Chrysotus cilipes Meigen, 1824, Russia (Primorje). E. Hypopygium, lateral view. F. Apex of phallus, lateral view.Published as part of Negrobov, Oleg P., Kumazawa, Tatsunori, Tago, Toshihiro & Fursov, Viktor N., 2016, Species of the genus Chrysotus Meigen, 1824 (Diptera: Dolichopodidae) from Japan, with descriptions of two new species, pp. 1-15 in European Journal of Taxonomy 197 on page 10, DOI: 10.5852/ejt.2016.197, http://zenodo.org/record/383796

    Chrysotus parilis Parent 1926

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    Chrysotus parilis Parent, 1926 Material examined JAPAN: Saitama Prefecture, Misato City, Edogawa river bank, 9 Aug. 2008 (3 ƋƋ, 2 ♀♀), 13 Oct. 2008 (1 Ƌ), 15 May 2010 (2 ƋƋ), leg. T. Tago; 1 Ƌ, Honshu, Ibaraki Prefecture, Tsukuba, Yawara, rice fields, 16 Jul. 1997, leg. V. Fursov; 1 Ƌ, Honshu Aichi Prefecture, Nagoya, Nagakute-cho Park, 23–30 May 1999, leg. V. Fursov. Distribution China, Japan. The species originally described from China is here recorded for the first time from Japan.Published as part of Negrobov, Oleg P., Kumazawa, Tatsunori, Tago, Toshihiro & Fursov, Viktor N., 2016, Species of the genus Chrysotus Meigen, 1824 (Diptera: Dolichopodidae) from Japan, with descriptions of two new species, pp. 1-15 in European Journal of Taxonomy 197 on page 4, DOI: 10.5852/ejt.2016.197, http://zenodo.org/record/383796

    Chrysotus nudisetus Negrobov & Maslova 1995

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    Chrysotus nudisetus Negrobov & Maslova, 1995 Chrysotus sp. 3t – Tago 2010: 44. Material examined JAPAN: 1 Ƌ, Saitama Prefecture, Misato City, Edogawa river bank, 9 Aug. 2008, leg. T. Tago; 1 Ƌ, Chiba Prefecture, Nagareyama City, Ichinoya, 30 Sep. 2008, leg. T. Tago; 1 Ƌ, Saitama Prefecture, Hidaka City, Yokote, 12 Jul. 2009, leg. T. Tago; 1 Ƌ, Saitama Prefecture, Ootaki, Chichibu City, Mameyakibashi, 7 Aug. 2010, leg. T. Tago. Distribution Japan and Russia: Sakhalin, Primorye, Khabarovsk, Magadan, Chukotka. The species was described from Japan and the Far East of Russia by Negrobov & Maslova (1995).Published as part of Negrobov, Oleg P., Kumazawa, Tatsunori, Tago, Toshihiro & Fursov, Viktor N., 2016, Species of the genus Chrysotus Meigen, 1824 (Diptera: Dolichopodidae) from Japan, with descriptions of two new species, pp. 1-15 in European Journal of Taxonomy 197 on page 4, DOI: 10.5852/ejt.2016.197, http://zenodo.org/record/383796

    Chrysotus cilipes Meigen 1824

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    Chrysotus cilipes Meigen, 1824 Material examined JAPAN: 9 ƋƋ, Saitama Prefecture, Misato City, Edogawa river bank, 9 Aug. 2008, 15 May 2010, leg. T. Tago. Distribution Western Europe, Abchazia, Afghanistan, Armenia, Azerbaijan, China, Japan, Kazakhstan, Korea, Mongolia, Turkey, Ukraine and Russia (Leningrad, Novgorod, Pskov, Moscow, Voronezh, Adygea, Kabardino-Balkaria, Krasnodar, Rostov, Tomsk, Altai, Krasnoyarsk, Baikal, Transbaikalia, Yakutia, Amur, Maritime Territory). There are a few records of the species in Japan (Negrobov et al. 2008; Masunaga 2014).Published as part of Negrobov, Oleg P., Kumazawa, Tatsunori, Tago, Toshihiro & Fursov, Viktor N., 2016, Species of the genus Chrysotus Meigen, 1824 (Diptera: Dolichopodidae) from Japan, with descriptions of two new species, pp. 1-15 in European Journal of Taxonomy 197 on page 3, DOI: 10.5852/ejt.2016.197, http://zenodo.org/record/383796

    Figure 3 in Description and biology of two new egg parasitoid species (Hymenoptera: Trichogrammatidae) reared from eggs of Heliconiini butterflies (Lepidoptera: Nymphalidae: Heliconiinae) in Panama

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    Figure 3. Biology of Trichogramma chagres sp. nov. strain L21: (a–c) female adult(s) parasitizing egg(s) of Mamestra brassicae; (d) freshly laid wasp egg; (e) newly hatched larva; (f–h) mature larva: (f,g) two mature larvae and chorion of consumed host egg in direct (f) and reflected (g) light, (h) habitus of mature larva with pulsing mid gut full of host egg yolk.Published as part of Woelke, Jozef B., Fursov, Viktor N., Gumovsky, Alex V., Rijk, Marjolein de, Estrada, Catalina, Verbaarschot, Patrick, Huigens, Martinus E. & Fatouros, Nina E., 2019, Description and biology of two new egg parasitoid species (Hymenoptera: Trichogrammatidae) reared from eggs of Heliconiini butterflies (Lepidoptera: Nymphalidae: Heliconiinae) in Panama, pp. 639-657 in Journal of Natural History 53 (11) on page 646, DOI: 10.1080/00222933.2019.1606360, http://zenodo.org/record/367547

    Chrysotus

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    Key to the males of Japanese Chrysotus 1. Femora mostly dark-green or brown ………………………………………………………………2 – Femora mostly yellow ………………………………………………………………………………5 2. Postocular setae black (Fig. 3 A–C) ………………… Ch. nudisetus Negrobov & Maslova, 1995 – Postocular setae pale ………………………………………………………………………………3 3. Face wide, more than 2 times wider than the length between ocellar bristles. Thorax and abdomen with purple tingle. Postpedicel large, approximately 2 times wider than long (Fig. 3 D–E) ………… ……………………………………………………..………….…… Ch. laesus (Wiedemann, 1817) – Face narrow, narrower than the length between ocellar bristles. Thorax and abdomen green. Postpedicel small, approximately as long as wide ……………………………………………………4 4. Postpedicel triangular apically. Surstylus approximately 1.5 times longer than cercus. Apicoventral epandrial lobe well developed, broad oval, without excavation …………………… ……………………………………………… Ch. saigusаi Negrobov, Kumazawa & Tago sp. nov. – Postpedicel oval apically. Surstylus approximately equal to cercus. Apicoventral epandrial lobe not developed, with excavation at apex (Fig. 3 F–G) …………………………………………………… ……………………………………………… Ch. kumazawai Negrobov, Maslova & Fursov, 2015 5. Fore coxae with black hairs …………………………………………………………………………6 – Fore coxae with white hairs …………………………………………………………………………7 6. Postpedicel longer than wide. Dorsal part of fore femora, middle femora in the middle part and apical third of hind femora dark. Phallus with oval process at apex on the left side ……… ……………………………………………… Ch. masunagai Negrobov, Kumazawa, Tago sp. nov. – Postpedicel small, approximately as long as wide. Hind femora with dark spot at apex, the rest of femora yellow. Phallus with claw-shaped process at apex on the left side (Fig. 4 A–B) ………… ……………………………………………………… Ch. tagoi Negrobov, Maslova & Fursov, 2015 7. Hind tibia without long erect hairs. Hind tibia at apex without group of ventral hairs. Postpedicel large, triangular at apex (Fig. 4 C–D) ……………………………………… Ch. parilis Parent, 1926 – Hind tibiae and hind tarsi with long erect hairs, longer than tibia width. Hind tibia at apex with group of ventral hairs. Postpedicel small, oval apically (Fig. 4 E–F) ……… Ch. cilipes Meigen, 1824 Cladistic analyses According to Capellari (2013), the absence of a small projection in the basal part of the epandrium represents the plesiomorphic character state. The combination of this character state with such character states as pale palpus, dark fore coxae, yellow hind tibia with short erect anteroventral hairs and short erect hairs on the first segment of hind tarsus separate the group of Ch. saigusai and Ch. tagoi from all other Japanese Chrysotus (Fig. 6). The species Ch. saigusai can be separated from Ch. tagoi by having anteroventral hairs on the entire length of find femora and mostly dark fore femora. The presence of the epandrial projection is the typical character states of the species Ch. cilipes, Ch. parilis and Ch. masunagai sp. nov., they are, however, included in the group with Ch. saigusai sp. nov. and Ch. tagoi on the basis of the characters of face width, position of arista, cooler of postocular bristles, fore coxa, hind trochanter and morphology of CuA 1. Concerning the distinctive characters of the group including Ch. cilipes, Ch. parilis and Ch. masunagai sp. nov., in addition to the above mentioned, the following character states can be ascribed: yellow middle femora and dark basal part of hind femur. The morphological similarity between Ch. cilipes and Ch. parilis is sufficiently significant, while the relationship with Ch. masunagai sp. nov. is unstable. The species Ch. laesus, Ch. nudisetus and Ch. kumazawai compose the group with the least stable internal connections. Their association is based on the similarity of color of fore, middle and hind femora and the absence of the anteroventral hairs on entire length of hind femur.Published as part of Negrobov, Oleg P., Kumazawa, Tatsunori, Tago, Toshihiro & Fursov, Viktor N., 2016, Species of the genus Chrysotus Meigen, 1824 (Diptera: Dolichopodidae) from Japan, with descriptions of two new species, pp. 1-15 in European Journal of Taxonomy 197 on pages 9-12, DOI: 10.5852/ejt.2016.197, http://zenodo.org/record/383796
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