173,082 research outputs found
Ancyronyx tobada Freitag & Kodada 2017
No full textAncyronyx tobada Freitag & Kodada, 2017 (Figs 2 C; 4A, C, E) Material examined. 3LL (CKB, CFM [FR114]): “ INDONESIA: C Sulawesi, Rd. Km 43 Tentena-Gintu; mountain creek; cloud forest; submerged wood, c. 1680m asl., c. 01°48′39′′ S 120°23′17′′ E 0 4 Jan.1998 leg. Freitag (Sul 14f)”; 6LL (CKB [SEM], NMW, CFM [FR 119]): “ INDONESIA: C Sulawesi, Rd. Tentena-Pendola, 9km E Tentena, secondary forest; submerged wood, c. 860m asl., c. 01°46′31′′ S 120°43′05′′ E 0 3 Jan.1998 leg. Freitag (Sul 10f)”; 4LL (CFM [FR121]): “ INDONESIA: S Sulawesi, Rd. Km 56 Tentena-Gintu; black-water river; submerged wood, c. 1190m asl., c. 01°49′28′′ S 120°21′43′′E 0 4 Jan.1998 leg. Freitag (Sul16f)”. Differential diagnosis (based on presumably sixth instar, HW: 0.32 mm). Generally similar to Ancyronyx schoedli except for the following characters: pronotum yellowish in anterior third, yellowish lateral portions of head extending posteriad along the frontal suture; posterior margins of thoracic and abdominal terga slightly paler, somewhat dark yellow-brown; terminal abdominal segment dark along anterior 0.1, yellow-brown along anterior 0.1–0.6, dark brown along posterior 0.6–0.9 and pale yellow-brown at the tip. Entire ventral side pale yellowish, only abdominal segment IX occasionally slightly darker, but still somewhat pale. Dorsal tubercles moderately elevated (Figs 4 A, C). Head (Figs 2 C, 4A) slightly tapering anteriad, widest at posterior 0.2; antennae overreaching anterior margin of labrum. Abdominal segments gently elevated near entire posterior portion; dorsum with scattered short setae (Figs 4 C, E); lateral protuberances with tubercles almost as large as spiracles; segment IX (Figs 2 C, 4E) very long, ca. 2.4 times as long as wide; basomedian portion glabrous, without tubercles; apex deeply emarginated.Published as part of Freitag, Hendrik & Kodada, Ján, 2017, Larvae of Ancyronyx Erichson, 1847 (Insecta: Coleoptera: Elmidae) from Sulawesi, using DNA sequences for the assignment of the larval stages, pp. 121-130 in Zootaxa 4299 (1) on page 126, DOI: 10.11646/zootaxa.4299.1.6, http://zenodo.org/record/83515
Ancyronyx toraja Freitag & Kodada 2017
No full textAncyronyx toraja Freitag & Kodada, 2017 (Figs 2 E; 5A–D) Material examined. 24LL (CFM): “ INDONESIA: S Sulawesi, 17.5km NNW Rantepao 1510m asl., mount. riv. “Salu Maiting”, pine forest/paddy fields, run; submerged wood, c. 02°49′21′′ S 119° 51′17′′E 31 Dec.1997 leg. Freitag (Sul 4f)”; 6LL (NMW, CKB [SEM], CFM [FR133]): “ INDONESIA: S Sulawesi, 17.5km NNW Rantepao 1570m asl., small mount. river “Salu Malarak”, pine forest, run/riffle; submerged wood, c. 02°48′51′′ S 119° 51′21′′E 31 Dec.1997 leg. Freitag (Sul7f)”. Differential diagnosis (based on presumably fifth and sixth instar, HW: 0.32–0.38 mm). In most characters very similar to Ancyronyx tobada, except for the following: generally dark brown dorsally; paler brown areas limited to lateral portion of head, small areas at anterior and anterolateral margins of pronotum and apex of terminal abdominal segment. Entire ventral side pale. Dorsal tubercles moderately elevated, lateral protuberances distinctly protruding. Head distinctly tapering anteriad, widest at posterior 0.2. Abdominal segments distinctly, protuberantly elevated in posterior portions; posterior abdominal segments I–VIII with four erected setae; segment IX with six of such setae dorsally (Fig. 5 D, these setae are less distinct in the six larval instar); posterolateral protuberances with prominent tubercles and two distinct, moderately long setae (Figs 5 B, C); terminal segment IX (Figs 5 C, D) moderately long, ca. 2.0 times as long as wide; basomedian portion tuberculate; apex distinctly emarginated.Published as part of Freitag, Hendrik & Kodada, Ján, 2017, Larvae of Ancyronyx Erichson, 1847 (Insecta: Coleoptera: Elmidae) from Sulawesi, using DNA sequences for the assignment of the larval stages, pp. 121-130 in Zootaxa 4299 (1) on page 128, DOI: 10.11646/zootaxa.4299.1.6, http://zenodo.org/record/83515
Ancyronyx longiparamerus Freitag & Kodada 2017
No full textAncyronyx longiparamerus Freitag & Kodada, 2017 (Figs 2 D; 4B, D, F) Material examined. 2LL (NMW, CFM): “ INDONESIA: C Sulawesi, Rd. Km 43 Tentena-Gintu; mountain creek; cloud forest; submerged wood, c. 1680m asl., c. 01°48′39′′ S 120°23′17′′E 0 4 Jan.1998 leg. Freitag (Sul 14f)”; 2LL (CFM [FR132], CKB [SEM]): “ INDONESIA: S Sulawesi, 17.5km NNW Rantepao 1570m asl., small mount. river “Salu Malarak”, pine forest, run/riffle; submerged wood, c. 02°48′51′′ S 119° 51′21′′E 31 Dec.1997 leg. Freitag (Sul7f)”. Differential diagnosis (based on presumably sixth instar, HW: 0.33 mm). Very similar to Ancyronyx tobada except for the following characters: generally the colour is somewhat paler brown, surface of pronotum yellowish on extensive oblique anterolateral portion (Fig. 2 D); yellowish lateral portions of head extending posteriad; entire posterior half of abdominal segments I and VIII indistinctly paler; terminal abdominal segment dark except of paler brownish apex. Entire ventral side pale. Dorsal tubercles prominent and distinctly elevated (Figs 4 D, F); anterolateral portion of all thoracic and abdominal segments with glabrous signa, most conspicuous at pronotum. Head (Fig. 4 B) distinctly tapering anteriad, widest at posterior 0.1. Abdominal segments with scattered short erected setae (Fig. 4 F); posterolateral protuberances with prominent tubercles, which are distinctly smaller than spiracles; terminal segment IX (Fig. 4 F) moderately short, ca. 1.9 times as long as wide; basomedian portion distinctly and densely tuberculate; apex slightly emarginated.Published as part of Freitag, Hendrik & Kodada, Ján, 2017, Larvae of Ancyronyx Erichson, 1847 (Insecta: Coleoptera: Elmidae) from Sulawesi, using DNA sequences for the assignment of the larval stages, pp. 121-130 in Zootaxa 4299 (1) on pages 127-128, DOI: 10.11646/zootaxa.4299.1.6, http://zenodo.org/record/83515
Ancyronyx henningi Freitag & Kodada 2017
No full textAncyronyx henningi Freitag & Kodada, 2017 (Fig. 2 F) Material examined. 2 LL (NMW, CFM [FR131]): “ INDONESIA: C Sulawesi, Rd. Km 32 Tentena-Gintu; mountain creek, grass-land; submerged wood, c. 1600m asl., c. 01°48′10′′ S 120°28′10′′E 0 4 Jan.1998 leg. Freitag (Sul 13f)”. Differential diagnosis (based on presumably fifth instar, HW: 0.33 mm). In most characters very similar to Ancyronyx tobada except for: dominantly pale brown dorsal side, dark brown areas limited to dorsal portion of head and subapical portion of terminal abdominal segment (Fig. 2 F). Anterior half of pronotum yellowish, remaining thoracic and abdominal segments pale brown with darker posteromedian brown patches (most distinct at pronotum) and darker sublateral areas; abdominal apex yellowish (Fig. 2 F). Entire ventral side more or less pale. Dorsal tubercles moderately elevated, lateral protuberances distinctly protruding. Head slightly tapering anteriad, widest at posterior 0.3. Abdominal segments moderately elevated at posterior portions, with few indistinct scattered short erected setae; posterolateral protuberances with prominent tubercles and two distinct, long setae; terminal segment IX long, ca. 2.3 times as long as wide; basomedian portion almost glabrous, only with few small tubercles; apex distinctly notched.Published as part of Freitag, Hendrik & Kodada, Ján, 2017, Larvae of Ancyronyx Erichson, 1847 (Insecta: Coleoptera: Elmidae) from Sulawesi, using DNA sequences for the assignment of the larval stages, pp. 121-130 in Zootaxa 4299 (1) on page 129, DOI: 10.11646/zootaxa.4299.1.6, http://zenodo.org/record/83515
Hydraena (Hydraenopsis) pseudopalawanensis Freitag & Jach, sp. n.
No full textHydraena (Hydraenopsis) pseudopalawanensis Freitag & Jäch, sp. n. (Figs. 23 a–d) Type locality. Estrella Falls, 7 km north of Narra, Municipality of Narra, 09° 18 'N 118 ° 23 'E, southern Palawan, Philippines. Type material. Holotype ɗ (NMW): “ PHIL.: Palawan 1994 7km N Narra, 2.4. Estrella Falls leg. Zettel (57)”, terminal parts of abdomen and aedeagus glued separately, one maxillary palpus and several parts of legs lacking. Paratypes: 1 ɗ (NMW, mutilated, aedeagus lacking): “ PHIL.: Palawan, P. Princesa SSW Maratarpi Cabayugan R 1 10 °09' 46 ’’N 118 ° 49 ' 29 ’’ E 0 2.6. 2001, leg. Freitag (i 5)”; 1 ɗ (NMW): “ PHIL.: Palawan, P. Princesa S Manturon, Cabayugan R.CR 3 10 °09' 16 ''N 118 ° 52 ' 30 '' E 29.3. 2001, leg. Freitag (5)”; 1 ɗ (NMW): “ PHIL.: Palawan, P. Princesa SE Manturon, Karst spring LS 4 R 10 °09' 29 ''N 118 ° 53 ' 30 ''E 14.VI. 2001, leg. Freitag (i 36) E”; 1 ɗ (NMW): “ PHIL.: Palawan, P. Princesa SE Manturon, Karst spring LS 4 R 10 °09' 29 ''N 118 ° 53 ' 30 ''E 28.IX. 2000, leg. Freitag (i 42) E”; 1 ɗ (NMW): “ PHIL.: Palawan, P. Princesa SE Manturon, Karst spring LS 4 R 10 °09' 29 ''N 118 ° 53 ' 30 ''E 18.III. 2001, leg. Freitag (i 53) E”; 1 ɗ (NMW, aedeagus and terminal part of abdomen lacking): “ PHIL.: Palawan, P. Princesa SE Manturon, Karst spring LS 4 R 10 °09' 29 ''N 118 ° 53 ' 30 ''E 18.III. 2001, leg. Freitag (i 54) E”; 1 ɗ (NMW): “ PHIL.: Palawan, P. Princesa SE Manturon, Karst spring LS 4 R 10 °09' 29 ''N 118 ° 53 ' 30 ''E 18.III. 2001, leg. Freitag (i 55) E”; 1 ɗ (UPLB): “ PHIL.: Palawan, P. Princesa SE Manturon, Karst spring LS 4 R 10 °09' 29 ''N 118 ° 53 ' 30 ''E 18.III. 2001, leg. Freitag (i 56) E”; 1 ɗ (NMW): “ PHIL.: Palawan, P. Princesa SE Manturon, Karst spring LS 4 R 10 °09' 29 ''N 118 ° 53 ' 30 ''E 18.III. 2001, leg. Freitag (i 57) E”; 1 ɗ (NMW): “ PHIL.: Palawan, P. Princesa S Manturon, Cabayugan R.CR 4 10 °09' 28 ''N 118 ° 53 ' 26 ''E 31.VII. 2001, leg. Freitag (i 37)E”; 1 ɗ (NMW): “ PHIL.: Palawan, P. Princesa Cabayugan R.; Nagdayan Creek, Manturon Bridge 10 ° 10 'N 118 ° 53 ' E 27.3. 2001, leg. Freitag (1)M”; 1 ɗ (WPU, aedeagus lacking): “ PHIL.: Palawan, P. Princesa Cabayugan R.; Nagdayan Creek, Manturon Bridge 10 ° 10 'N 118 ° 53 ' E 27.3. 2001, leg. Freitag (9)M”; 3 ɗɗ (NMW): “ PHIL.: Palawan, P.Princesa; Luzviminda, forest creek, small fall, 9km SSW PPC; sandy banks, pool, c. 50m asl 9 ° 39 'N 118 ° 38 'E, 14.12.1994,leg.Freitag(17 b)M”; 6 ɗɗ (3 exs. NMW; 1 ex. IRML, mutilated; 1 ex. UPLB; 1 ex. WPU): “ PHIL.: Palawan, P.Princesa; Luzviminda, forest creek, small fall, 9km SSW PPC; side pools w/ leaves, c. 50m asl 9 ° 39 'N 18 ° 38 'E, 14.12.1994,leg.Freitag(17 c)M”; 1 ɗ (NMW, aedeagus lacking): “ Philippines, Palawan centr. Ulangoan river, 18 km NE San Rafael 6.XII. 1995, J. Kodada & B. Rigová lgt.”. Additional material. 1 Ψ (NMW): “ PHIL.: Palawan 1994 7km N Narra, 5.4. Estrella Falls leg. Zettel (59)”. This female was collected at the type locality of Hydraena pseudopalawanensis. Although it is most likely that this female belongs to H. pseudopalawanensis, we refrain from including it in the type series (for explanation, see below). Diagnosis. Body 1.15–1.20 mm long. Colour and habitus as in Hydraena palawanensis (Fig. 10). Aedeagus (Figs. 23 a–c) agrees in several principal characters with that of H. palawanensis, but differs in numerous other conspicuous characters: e.g., apical half of main piece more strongly curved (ventral/dorsal view), right margin of main piece without conspicuous emargination near insertion of paramere; presence of one dorsal seta; left paramere moderately long, slender, apically with five moderately long setae; right paramere ventromedially with a longitudinal row of ca. four moderately long setae. We were not able to distinguish females of H. pseudopalawanensis and H. palawanensis reliably. Since both species occur in the same habitat, we were not able to identify all females from central Palawan (deposited in the NMW) properly: 1 Ψ: “ PHIL.: Palawan, P. Princesa S Manturon, Cabayugan R.CR 3 10 °09' 16 ''N 118 ° 52 ' 30 '' E 29.3. 2001 leg. Freitag (4)”; 1 Ψ: “ PHIL.: Palawan, P. Princesa S Manturon, Cabayugan R.CR 4 10 °09' 28 ''N 118 ° 53 ' 26 ''E 31.VII. 2001, leg. Freitag (i 38)E”; 1 Ψ: “ PHIL.: Palawan, P. Princesa S Manturon, Cabayugan R.CR 4 10 °09' 28 ''N 118 ° 53 ' 26 ''E 31.VII. 2001, leg. Freitag (i 39)E”; 1 Ψ: “ PHIL.: Palawan, P. Princesa S Manturon, Cabayugan R.CR 4 10 °09' 28 ''N 118 ° 53 ' 26 ''E 18.III. 2001, leg. Freitag (i 40)E”; 1 Ψ (mutilated): “ PHIL.: Palawan, P. Princesa SE Manturon, Cabayugan R. 10 °09.486'N 118 ° 53.493 ' E 3.3. 2001, leg. Freitag (i 4)”; 1 Ψ: “ PHIL.: Palawan, P. Princesa SE Manturon, Karst spring LS 4 R 10 °09' 29 ''N 118 ° 53 ' 30 ''E 18.III. 2001, leg. Freitag (i 58) E”; 1 Ψ: “ PHIL.: Palawan, P. Princesa SE Manturon, Karst spring LS 4 R 10 °09' 29 ''N 118 ° 53 ' 30 ''E 18.III. 2001, leg. Freitag (i 59) E”; 1 Ψ (mutilated): “ PHIL.: Palawan, P. Princesa Cabayugan R.; Nagdayan Creek, Manturon Bridge 10 ° 10 'N 118 ° 53 ' E 27.3. 2001, leg. Freitag (6)M”; 1 Ψ: “ PHIL.: Palawan, P. Princesa Cabayugan R.; Nagdayan Creek, Manturon Bridge 10 ° 10 'N 118 ° 53 ' E 27.3. 2001, leg. Freitag (10)M”; 5 ΨΨ: “ PHIL.: Palawan, P.Princesa; Luzviminda, forest creek, small fall, 9km SSW PPC; sandy banks, pool, c. 50m asl 9 ° 39 'N 118 ° 38 'E, 14.12.1994,leg.Freitag(17 b)M”; 9 ΨΨ: “ PHIL.: Palawan, P.Princesa; Luzviminda, forest creek, small fall, 9km SSW PPC; side pools w/ leaves, c. 50m asl 9 ° 39 'N 18 ° 38 'E, 14.12.1994,leg.Freitag(17 c)M”. Distribution (Fig. 31). Hydraena pseudopalawanensis is obviously confined to the island of Palawan, where it is not uncommon. Ecology. Hydraena pseudopalawanensis resembles H. palawanensis even in ecological terms. The habitats include rather undisturbed (e.g. CR 1) and to some extent anthropogenically influenced rural areas (e.g. CR 3; Freitag 2005), suggesting a rather high ecological potency. Besides numerous specimens caught in emergence traps, some individuals were manually collected together with H. palawanensis in the same habitat (detritus interstitial). Etymology. The species is named in reference to its close relationships with Hydraena palawanensis.Published as part of Freitag, Hendrik & Jäch, Manfred A., 2007, Revision of the species of Hydraena Kugelann (Coleoptera: Hydraenidae) from Palawan and Busuanga, with descriptions of eleven new species, and redescription of Hydraena (Hydraenopsis) scabra d'Orchymont, 1925, pp. 1-44 in Zootaxa 1431 on pages 29-31, DOI: 10.5281/zenodo.17579
Ancyronyx skalei Freitag & Kodada 2017
No full textAncyronyx skalei Freitag & Kodada, 2017 (Fig. 2 B) Material examined. 3LL (NMW, CKB, CFM [FR126]): “ INDONESIA: Sulawesi, T. Toraja, Salu Toriu River, 810m asl., c. 02°56′38′′ S 119°52′24′′E 0 2 Jan.1998 leg. Freitag (Sul 8 Af)”. Differential diagnosis (based on presumably fifth instar, HW: 0.32 mm). Very similar to the previous species, except for the following: anterior quarter of pronotum and posterior quarter of terminal abdominal segment yellowish to pale brown (Fig. 2 B); lateral portions of the head, entire first abdominal segment and posterior half/entire eight abdominal segment pale; small transverse pale spots might also be present at medioposterior margin of pro-, meso-, and metanotum. Dorsal tubercles slightly elevated. Head more tapering anteriad, widest at posterior 0.2; antennae rather small, approximately reaching anterior margin of labrum. Abdominal segment IX moderately long, ca. 1.9 times as long as wide; basomedian portion slightly tuberculate; apex distinctly emarginated.Published as part of Freitag, Hendrik & Kodada, Ján, 2017, Larvae of Ancyronyx Erichson, 1847 (Insecta: Coleoptera: Elmidae) from Sulawesi, using DNA sequences for the assignment of the larval stages, pp. 121-130 in Zootaxa 4299 (1) on page 126, DOI: 10.11646/zootaxa.4299.1.6, http://zenodo.org/record/83515
Hydraena (Hydraenopsis) kodadai Freitag & Jach, sp. n.
No full textHydraena (Hydraenopsis) kodadai Freitag & Jäch, sp. n. (Figs. 9, 21a–h) Type locality. Small stream in primary forest, ca. 100 m a.s.l., tributary of Cabayugan River, St. Paul Subterranean River National Park, Cabayugan district, 40 km NNE of Puerto Princesa City, central Palawan, Philippines. Type material. Holotype ɗ (NMW): “ Philippines, PALAWAN centr. smal stream in primary forest ca. 100 m, Cabayugan env., 2. 12. 1995, Ján Kodada lgt.”, terminal parts of abdomen and aedeagus glued separately. Paratypes: 2 ΨΨ (1 ex. NMW, 1 ex. UPLB): same locality data as holotype; 1 ɗ (NMW), 4 ΨΨ (NMW): “ Philippines, PALAWAN centr., Ulangoan river, 18 km NE San Rafael 6.XII. 1995, J. Kodada & B. Rigová lgt.”; 2 ΨΨ (NMW, 1 ex. mutilated): “ PHIL.: Palawan, P. Princesa Sabang, Waterfall (upper, 30m asl) 10 ° 13 'N 118 ° 53 ' E 26.3. 1995, leg. Freitag (19 a)”; 1 ɗ (UPLB): “ PHIL.: Palawan, P. Princesa Panaguman R. 10 ° 15.150 'N 118 ° 58.051 ' E 17.5. 2001, leg. Freitag (i 25)”; 1 ɗ (NMW, mutilated), 1 Ψ (NMW): “ PHIL.: Palawan, P. Princesa Panaguman R. 10 ° 15.150 'N 118 ° 58.051 ' E 17.5. 2001, leg. Freitag (e 18)”; 1 ɗ (NMW): “ PHIL.: Palawan, P. Princesa Panaguman R. 10 ° 15.150 'N 118 ° 58.051 ' E 17.5. 2001, leg. Freitag (e 26)”; 1 ɗ (WPU): “ PHIL.: Palawan, P. Princesa Panaguman R. 10 ° 15.150 'N 118 ° 58.051 ' E 17.5. 2001, leg. Freitag (e 29)”; 1 ɗ (NMW): “ PHIL.: Palawan, P. Princesa Panaguman R.PR1, 1.Drift 10 ° 15 '09''N 118 ° 58 '03''E 17.V. 2001, leg. Freitag (a 48)”; 1 Ψ (NMW): “ PHIL.: Palawan, P. Princesa Panaguman R.PR1, 1.Drift 10 ° 15 '09''N 118 ° 58 '03''E 17.V. 2001, leg. Freitag (a 49)”; 1 ɗ (NMW): “ PHIL.: Palawan, P. Princesa SSW Maratarpi Cabayugan R 1 10 °09' 46 ’’N 118 ° 49 ' 29 ’’ E 11.12. 2000, leg. Freitag (a 7)”; 1 Ψ (WPU, mutilated): “ PHIL.: Palawan, P. Princesa SSW Martarpi Cabayugan R. 1 10 °09' 46 ''N 118 ° 49 ' 29 '' E 11.12. 2000, leg. Freitag (9)”; 1 ɗ, 1 Ψ (NMW): “ PHIL.: Palawan, P. Princesa SSW Maratarpi Cabayugan R 1 10 °09' 46 ’’N 118 ° 49 ' 29 ’’ E 17.6. 2001, leg. Freitag (e 10)”; 1 ɗ (IRML; aedeagus mutilated): “ PHIL.: Palawan, P. Princesa SSW Martarpi Cabayugan R.1, 10°09' 46 ''N 118 ° 49 ' 29 ''E 1.VII. 2001, leg. Freitag (a 46)E”; 1 Ψ (NMW): “ PHIL.: Palawan, Taytay; 1 / 3 way highway to Lake Manguao, Manguao Stream trib., prim.forest c. 35m asl, 10 ° 47 'N 119 ° 31 ' E 29.4. 1995, leg. Freitag (13 b)M”; 1 Ψ (IRML) “ PHIL.: Palawan, P. Princesa SSW Maratarpi Cabayugan R 1 10 °09' 46 ’’N 118 ° 49 ' 29 ’’ E 16.11. 2000, leg. Freitag (e 12)”. Description. Body 1.30–1.45 mm long. Habitus as in Fig. 9. Entire body reddish brown, sometimes goldbrown; frons slightly darker brown, legs and maxillary palpi slightly paler. Pronotum wider than long, broadest at about posterior 0.3, entire pronotum moderately densely punctate; punctures small and shallowly (posteriorly moderately deeply) impressed, interstices glabrous; foveae hardly perceptible, or absent; lateral margins anteriorly convergent, distinctly sinuously convergent to rectangular posterior angles, forming an almost rectangular pronoto-elytral angle; lateral rim denticulate; anterior margins concave, distinctly broader than straight posterior margin; lateral hypomeron approximately as broad as profemur, inner margin (hypomeral carina) slightly biconvex; mesal hypomeron narrow, imperceptible. Elytra elongately oval, apex with tiny excision next to sutural keel, with ca. 12 longitudinal, not very regular, unimpressed rows of punctures (seven between suture and shoulder); punctures small and shallowly (basally moderately deeply) impressed, sparsely arranged; interstices and intervals flat, glabrous; lateral gutter of elytra widely explanate, not reaching elytral apex; elytral rim posteriorly finely denticulate; pseudepipleuron anteriorly about as broad as metafemur, reaching posterior 0.2, with indistinct foveae and one inconspicuous row of punctures; epipleuron narrow, short, inconspicuous. Mesoventrite with well developed longitudinal ridges; meso- and metaventrite, pubescent; abdominal ventrites sparsely pubescent; mesoventral intercoxal process moderately long, very wide; metaventral disc deeply impressed; metaventral plaques distinct, widely separated (wider than inflexed lateral portion of elytra), narrow, glabrous; intercoxal sternite large, approximately as broad as metafemora and as long as broad. Femora with inner margin straight to concave, outer margin conspicuously convex. Aedeagus (Figs. 21 a–c) slender, gently evenly curved ventrad and dextrad, with one dorsal subapical seta; dorso-ventrally flattened apically, appearing almost symmetric (ventral/dorsal view); phallobase asymmetric. Distal lobe not clearly delimited from main piece, median portion weakly sclerotized. Parameres articulately connected with main piece, moderately long; each paramere with a row of about 10 moderately long setae; left paramere inserted at about basal 0.50, right paramere at basal 0.55 of aedeagus. Gonocoxite (Fig. 21 e) subsemioval; apical area of ventral plate almost as long as basal area; subapical tufts curved; apical margin of basal area with median projection; condyles very small; dorsal plate very simple, slightly surpassing outer plate basally, without cavea. Spermatheca (Figs. 21 g–f). Proximal portion extremely large, strongly curved; distal portion discoidal. Secondary sexual characters. Male usually slightly larger than female. Meso- and metatibia in male slightly curved. Male profemur with (rather inconspicuous) ventro-basal ledge. Male terminal sternite (Fig. 21 d) oblongly subhexagonal, apical margin rather straight, base broad, distinctly produced into lateral appendages; spiculum gastrale about twice as long as terminal sternite, almost straight. Male tergite X distinctly excised apically. Female tergite X (Fig. 21 h) subsemicircular, with rather few long discal trichoid setae; subapical fringe with moderately long trichoid setae; hyaline apical margin entire. Differential Diagnosis. Hydraena kodadai resembles H. zetteli in size, colour and general habitus. Externally, H. kodadai can be distinguished from the latter by its narrower explanate elytral gutter, which ends near apical 0.2 of elytra. Furthermore, H. kodadai can easily be recognized by its posteriorly distinctly convergent pronotum, number of elytral striae (usually 12) and its broad, subsymmetric, bilobed and dorso-ventrally flattened aedeagus, and by its extremely large spermatheca. Distribution (Fig. 31). Hydraena kodadai is known from four small stream systems in central Palawan. Ecology. All known macro-habitats are rather undisturbed streams (first to fourth order) surrounded by forest, however, with a remarkable variation in physicochemical conditions. PR 1: acidic pH (6.32–7.02), low electrical conductivity (84 µS/cm on average), CR 1: alkaline water (pH 8.07–8.45), high conductivity (435 µS/cm on average) due to ultra-basic subsoils (Freitag 2004). Besides numerous specimens trapped in drift nets, some were collected repeatedly in gravel substrates under a small fall and in moderately fast flowing sections. Obviously, this species prefers (or at least tolerates) higher velocity. However, it was collected several times by emergence traps positioned over a pool section as well. Etymology. Named for Dr. Ján Kodada (Comenius University, Bratislava, Slovakia), excellent water beetle specialist, who considerably contributed to the knowledge of the fauna of Palawan.Published as part of Freitag, Hendrik & Jäch, Manfred A., 2007, Revision of the species of Hydraena Kugelann (Coleoptera: Hydraenidae) from Palawan and Busuanga, with descriptions of eleven new species, and redescription of Hydraena (Hydraenopsis) scabra d'Orchymont, 1925, pp. 1-44 in Zootaxa 1431 on pages 24-26, DOI: 10.5281/zenodo.17579
Ancyronyx schoedli Freitag & Kodada 2017
No full textAncyronyx schoedli Freitag & Kodada, 2017 (Figs 2 A; 3A–F) Material examined. 4LL (CKB [SEM], CFM [FR128, FR129]): “ INDONESIA: Sulawesi, T. Toraja, Salu Toriu River, 810m asl., c. 02°56′38′′ S 119°52′24′′E 0 2 Jan.1998 leg.Freitag (Sul8Af)”; 1L (NMW): “S - SULAWESI 1992 Malino-Manipi 700m (31) leg. Jäch 1.V. ”. Larval diagnosis (based on presumably fifth instar, HW: 0.30 mm). Body elongate, nearly semicircular in cross section, dorsally vaulted, ventrally almost flat. Posterolateral margins of abdominal segments I–VII moderately produced (Fig. 3 C). Colour (Fig. 2 A) dorsally dominantly dark brown except for pale brown to yellowish lateral portion of head, apex of the terminal abdominal segment (up to posterior 0.15) and the entire first and eight abdominal segments; clypeus and labrum reddish brown; legs, antennae, remaining mouthparts, entire ventral side including operculum and except abdominal segment IX yellowish to pale brown. The larva is very similar in the external characters to that of A. minerva (see Freitag & Balke 2011), except for the following characters: lateral abdominal projections (Figs 3 C, D) with two large robust long setae (referred to as “trichoid teeth” in Freitag & Balke 2011), one robust short seta and one long thin trichoid seta. Dorsal tubercles slightly elevated (Figs 3 A–D). Head (Fig. 3 A) widest at posterior 0.3, sides arcuate in posterior half; frontal suture V-shaped, conspicuous in wet specimens; dorsolateral portion with three pairs of conspicuous setae: first pair consists of short setae situated mediofrontally, second pair of moderately long setae inserted near the frontoclypeal suture and third pair of long setae originate near frontal suture behind the eyes; lateral portion with about 4 pairs of moderately long setae. Antennae moderately long, overreaching anterior margin of labrum; setae of scapus trichoid. Labrum with subapical fringe of ramose setae and few trichoid setae laterally. Maxillary palpus ca. as long as stipes wide. Mentum subrectangular, slightly narrower along posterior 0.1–0.4; pair of trichoid setae moderately long, not reaching anterior margin, inserted sublaterally at anterior 0.35 of mentum. Pro- meso- and metathorax subtrapezoidal, slightly narrowed anteriorly, broader than long; with slightly produced lateral rim; lateral rim with few scattered long trichoid setae; dorsosagittal carinae small, narrow and present at anterior half of mesonotum; surface of meso- and metanotum with microtubercles; pronotum with ca. six pairs of rather inconspicuous, irregularly shaped and partly fused signa (flat, subglabrous areas) in posterior half (Figs 3 A, B); remaining areas of pronotum including anterolateral grooves with setiferous tubercles. Ventral sclerites of thorax granulose. Abdomen (Figs 2 A, 3C–F) without dorsosagittal carina, with distinct mesal suture except for segment IX (Figs 3 C, E); lateral rim with few scattered long, trichoid setae; squamose setae at posterior rim of segments large; ventral sclerites of segment I with rather indistinct sagittal ridge in anterior half; segment IX (Fig. 3 E) moderately short, ca. 1.6 times as long as wide; basomedian portion tuberculate, apical portion rugose; apex slightly emarginated; operculum ca. 1.7 times as long as wide.Published as part of Freitag, Hendrik & Kodada, Ján, 2017, Larvae of Ancyronyx Erichson, 1847 (Insecta: Coleoptera: Elmidae) from Sulawesi, using DNA sequences for the assignment of the larval stages, pp. 121-130 in Zootaxa 4299 (1) on pages 124-126, DOI: 10.11646/zootaxa.4299.1.6, http://zenodo.org/record/83515
Hydraena (Hydraenopsis) nielshaggei Freitag & Jach, sp. n.
No full textHydraena (Hydraenopsis) nielshaggei Freitag & Jäch, sp. n. (Figs. 5, 17a–h) Type locality. Cabayugan River, near Nagsatayan Creek, south of Martarpi; 10 °09.78'N 118 ° 50.61 'E; St. Paul Subterranean River National Park, Cabayugan district, Puerto Princesa City, central Palawan, Philippines. Type material. Holotype ɗ (NMW): “ PHIL.: Palawan, P. Princesa S Maratarpi, Cabayugan R. 10 °09.780'N 118 ° 15.610 ' E 16.5. 2001, leg. Freitag (a 21)”, abdomen and aedeagus glued separately, some parts of legs lacking. Paratypes: 1 Ψ (NMW, terminal part of abdomen lacking): same locality data as holotype; 1 ɗ (NMW): “ PHIL.: Palawan, P. Princesa S Maratarpi, Cabayugan R. 10 °09.780'N 118 ° 15.610 ' E 16.5. 2001, leg. Freitag (a 2)”; 1 ɗ (WPU, aedeagus lacking): “ PHIL.: Palawan, P. Princesa S Maratarpi, Cabayugan R. 10 °09.780'N 118 ° 15.610 ' E 16.5. 2001, leg. Freitag (a 19)”; 2 ɗɗ (NMW, 1 ex. mutilated): “ PHIL.: Palawan, P. Princesa S Maratarpi, Cabayugan R. 10 °09.780'N 118 ° 15.610 ' E 16.5. 2001, leg. Freitag (a 20)”; 1 ɗ (NMW): “ PHIL.: Palawan, P. Princesa S Maratarpi, Cabayugan R. 10 °09.780'N 118 ° 15.610 ' E 16.5. 2001, leg. Freitag (b 8)”; 2 ΨΨ (UPLB; mutilated): “ PHIL.: Palawan, P. Princesa S Maratarpi, Cabayugan R. 10 °09.780'N 118 ° 15.610 ' E 16.5. 2001, leg. Freitag (a 30)”; 1 Ψ (WPU, mutilated): “ PHIL.: Palawan, P. Princesa S Maratarpi, Cabayugan R. 10 °09.780'N 118 ° 15.610 ' E 16.5. 2001, leg. Freitag (d 1)”; 1 ɗ (NMW): “ PHIL.: Palawan, P. Princesa SSW Martarpi Cabayugan R 1 10 °09' 46 ''N 118 ° 49 ' 29 '' E 16.5. 2000, leg. Freitag (a 24)”; 1 ɗ (NMW): “ PHIL.: Palawan, P. Princesa SSW Martarpi Cabayugan R. 1 10 °09' 46 ''N 118 ° 49 ' 29 '' E 16.5. 2001, leg. Freitag (8)”; 1 ɗ (NMW): “ PHIL.: Palawan, P. Princesa SSW Martarpi Cabayugan R.1, 10°09' 46 ''N 118 ° 49 ' 29 ''E 11.XII. 2000, leg. Freitag (i 44)E”; 1 ex. (WPU, terminal part of abdomen lacking): “ PHIL.: Palawan, P.Princesa; mountain stream 2km SE Laptay/Napsan, semi-prim.forest, 9 ° 43 'N 118 ° 28 ' E 29.3. 1995, leg. Freitag (23)M”. Description. Body 1.05–1.15 mm long. Habitus as in Fig. 5. Elytra and rectangular spot – if present – in the middle of pronotal disc dark brown, otherwise pronotal disc pale grey-brown; frons darker, in most specimens reddish-brown; pronotal margins and legs much paler. Pronotum wider than long, broadest at posterior 0.3–0.5; entire pronotum densely punctate; punctures moderately large and moderately deeply impressed, interstices glabrous; foveae shallowly impressed and hardly perceptible; anterior margin slightly concave; anterior angles rounded; lateral rim denticulate; lateral margins anteriorly convergent, posteriorly sinuously convergent (inflexed) forming a distinct pronoto-elytral angle; posterior margin straight to slightly bisinuate; lateral portions moderately deflexed; lateral hypomeron approximately as broad as profemora, mesal hypomeron narrow, inconspicuous. Elytra elongately oval, conjointly rounded apically; with ca. 15 longitudinal, quite irregular, not impressed rows of punctures (10 between suture and shoulder); punctures small and shallowly impressed; interstices and intervals almost flat, glabrous; inflexed lateral portion of elytra moderately explanate, not reaching elytral apex, distinctly narrower than metafemora; rim inconspicuously denticulate posteriorly; epipleuron inconspicuous; pseudepipleuron distinct, reaching ca. posterior 0.25, glabrous, with an indistinct row of punctures. Mesoventrite with well developed longitudinal ridges; mesoventral intercoxal process short, almost as broad as pseudepipleuron; metaventral disc deeply impressed; metaventral plaques distinct, widely separated (distance wider than inflexed lateral portion of elytra), very narrow, glabrous; intercoxal sternite very large, approximately as broad as metafemur and as long as broad, sub-triangular; metaventrite and abdominal sternites moderately densely pubescent. Femora with inner margins straight to concave, outer margins conspicuously convex. Aedeagus (Figs. 17 a–c) rather simple, elongate and slender. Main piece rather straight, slightly curved to right side (ventral/dorsal view), without setae, with five micropores; phallobase asymmetric. Distal lobe small, with few short amorphic appendages, not clearly delimited from main piece. Parameres very short, almost completely reduced, not articulately connected with main piece; left paramere inserted at about basal 0.55 of aedeagus, with ca. eight moderately long setae (two dorsal apical, one ventral apical, five ventral lateral, arranged in a longitudinal row); right paramere inserted at about basal 0.6 of aedeagus, with ca. eight moderately long setae (five dorsal, three ventral). Gonocoxite (Fig. 17 e) subtriangular, rather short; lateral margins strongly convex subbasally, slightly emarginate medially; apical area large; margin between apical and basal area well developed, sinuous; subapical tuft setae curved; basal area without distinct pubescence; condyles prominent; dorsal plate simple, slightly surpassing ventral plate, without distinct cavea. Spermatheca (Figs. 17 f–g) very similar to that of Hydraena jojoorculloi. Secondary sexual characters. Male profemur with tiny denticle near base of ventral face. Metaventral plaques narrower in male; meso- and metatibia in male slightly curved. Male terminal sternite and spiculum gastrale (Fig. 17 d) very similar to Hydraena jojoorculloi, but micropores at mediolateral portion of terminal sternite less conspicuous, and spiculum gastrale comparatively longer. Male tergite X distinctly excised apically. Female tergite X (Fig. 17 h) subtrapezoidal; anterior and posterior corners widely rounded; lateral margins evenly rounded; disc sparsely covered with setae; subapical fringe of trichoid setae widely interrupted medially; hyaline apical margin distinctly notched medially. Differential diagnosis. General habitus and colour resemble Hydraena jojoorculloi, from which it can be distinguished by its slightly smaller size, the slender bent aedeagus with characteristic short amorphic appendages, and by the subapical fringe of trichoid setae of female tergite X. Distribution (Fig. 30). This species is known from the Cabayugan River and the Ulangoan River, central Palawan. It is most likely endemic to Palawan. Ecology. The occurrence in headwaters of rather undisturbed, forest-covered areas suggests an affinity to pristine habitats. Most specimens were collected in drift traps. Two specimens were caught in an emergence trap. Etymology. Named for Niels Hagge (Hamburg), zoologist, for his support during field work in Palawan.Published as part of Freitag, Hendrik & Jäch, Manfred A., 2007, Revision of the species of Hydraena Kugelann (Coleoptera: Hydraenidae) from Palawan and Busuanga, with descriptions of eleven new species, and redescription of Hydraena (Hydraenopsis) scabra d'Orchymont, 1925, pp. 1-44 in Zootaxa 1431 on pages 15-17, DOI: 10.5281/zenodo.17579
Hydraena (Hydraenopsis) jojoorculloi Freitag & Jach, sp. n.
No full textHydraena (Hydraenopsis) jojoorculloi Freitag & Jäch, sp. n. (Figs. 4, 16a–h) Type locality. Manguao Stream (Lake Manguao tributary), Municipality of Taytay, northern Palawan, Philippines. Type material. Holotype ɗ (NMW): “ PHIL.: Palawan, Taytay; Lake Manguao tributary, Alipuran Stream, semi-prim. forest c. 40m asl, 10 ° 45 'N 119 ° 32 ' E 29.4. 1995, leg. Freitag (15)M”, terminal parts of abdomen and aedeagus glued separately. Paratypes: 2 ΨΨ (NMW): same locality data as holotype; 1 ɗ (NMW, aedeagus lacking), 2 ΨΨ (NMW, mutilated): “ PHIL.: Palawan, Taytay; 1 / 2 way high-way to Lake Manguao, Manguao Stream trib., prim.forest c. 30m asl, 10 ° 47 'N 119 ° 31 'E 03.4.1995,leg. Freitag(13 a)M”; 2 ΨΨ (NMW, 1 ex. mutilated): “ PHIL.: Palawan, Taytay; 1 / 3 way high-way to Lake Manguao, Manguao Stream trib., prim.forest c. 35m asl, 10 ° 47 'N 119 ° 31 ' E 29.4. 1995, leg. Freitag (13 b)M”; 1 Ψ (NMW): “ PHIL.: Palawan, P.Princesa; S. Vincente Taranaban Riv; 0.5km N of Highway, mountain riv. resid. pool, 10m asl 10 °01'N 119 °01'E, 30.4. 1995 leg. Freitag (16)M”; 1 Ψ (NMW): “ PHIL.: Palawan, P.Princesa; Junction to Napsan, 8km SW PPC; 20m S Binuan Bridge, sec.veget. c. 20m asl 9 ° 43 'N 118 ° 40 'E, 28.3. 1995 leg. Freitag (22)M”; 1 Ψ (UPLB): “ PHIL.: Palawan, P. Princesa Panaguman R.PR1, 1.Drift 10 ° 15 '09''N 118 ° 58 '03''E 17.V. 2001, leg. Freitag (47)”; 1 Ψ (NMW): “ Philippines, Palawan centr. Ulangoan river, 18 km NE San Rafael, 6. XII. 1995, J. Kodada & B. Rigová lgt.”. Description. Body approximately 1.25 mm long. Habitus as in Fig. 4. Elytra brown; frons dark brown; pronotum, legs and maxillary palpi distinctly paler yellowish brown to gold-brown. Pronotum wider than long, broadest at anterior 0.3; entire pronotum densely punctate; punctures small and moderately deeply impressed, interstices glabrous; anterior margin slightly concave; anterior angles rounded; lateral rim denticulate; lateral margins anteriorly convergent, posteriorly sinuously convergent; pronoto-elytral angle obtuse; posterior margin slightly biconcave; lateral hypomeron posteriorly slightly broader than profemur, mesal hypomeron narrow, inconspicuous. Elytra elongately oval, not conjointly rounded apically; each elytron with a tiny excision next to sutural keel, with ca. 16 longitudinal, more or less regular rows of punctures (approximately nine between suture and shoulder); punctures small and moderately deeply (apically shallowly) impressed, interstices and intervals convex to microreticulate; lateral portion moderately explanate, not reaching elytral apex, anteriorly and posteriorly denticulate; inflexed lateral portion (pseudepipleuron) anteriorly slightly broader than metafemur, reaching posterior 0.2, row of punctures inconspicuous; epipleuron narrow, short. Mesoventrite with well developed longitudinal ridges; mesoventral intercoxal process moderately broad, slightly narrower than pseudepipleuron; metaventral disc distinctly impressed; metaventral plaques inconspicuous, pubescent; intercoxal sternite moderately large, approximately as broad as metafemur and as long as broad; metaventrite and ventrites 1–4 moderately pubescent. Femora with inner margin straight to concave, outer margin conspicuously convex; tibiae with inner margin straight, outer one slightly convex. Aedeagus (Figs. 16 a–c) elongate and slender; main piece rather straight in ventral view, slightly sinuous in lateral view; without setae (three micropores present); phallobase asymmetric. Distal lobe intricately shaped, not clearly delimited from main piece; ejaculatory duct distally ending in a tapered stout flagellum, which is dorsally embedded in a less sclerotized ovoid structure. Parameres firmly fused to main piece; left paramere short, moderately wide, inserted at about basal 0.50 of aedeagus, latero-apically with ca. seven moderately long setae; right paramere inserted at about basal 0.55 of aedeagus, longer, wide, apically attenuately rounded, subapically with two parallel rows of ca. four moderately long setae. Gonocoxite (Fig. 16 e) subsemicircular, more or less evenly rounded; lateral margin only slightly convex subbasally; setae of subapical tufts straight; basal area of ventral plate with very few small scattered setae; condyles small and inconspicuous; dorsal plate asymmetric, simple, not surpassing outer plate. Spermatheca as in Figs. 16 f–g; proximal portion distally globular, proximately tubular, bent; distal portion wide and tubular, slightly curved. Secondary sexual characters. Male profemur with tiny denticle near base of ventral face (dissection of leg recommended). Male terminal sternite elongately subpentagonal, apical margin widely rounded; base slender, very slightly produced laterad; spiculum gastrale almost twice as long as terminal sternite, slightly bent (Fig. 16 d). Male tergite X apically distinctly excised. Female tergite X (Fig. 16 h) subsemicircular; lateral margins evenly rounded; disc sparsely covered with setae; subapical fringe of vermiform setae widely interrupted medially, laterally with short trichoid setae; hyaline apical margin notched medially. Differential diagnosis. This species can easily be distinguished from Hydraena scabra by the smaller size, by the conspicuously yellowish pronotum, and by its smaller and only moderately deeply impressed elytral and pronotal punctures. Furthermore, it can easily be recognized by its aedeagus (main piece rather straight, distal lobe ovoid with flagellum-like structure), the shape of the gonocoxite, and the presence of vermiform setae on female tergite X. Distribution (Fig. 30). Known only from three localities in northern and central Palawan. Ecology. This species occurs in small and medium-sized creeks draining hilly lowlands with forest cover. The species was collected manually in shallow, slowly flowing, unshaded reaches with sand, gravel, FPOM and CPOM. Etymology. Named for Jojo Orcullo (Puerto Princesa City), Palawan artist, for his support during the field work in Palawan and his personal loyalty to the first author.Published as part of Freitag, Hendrik & Jäch, Manfred A., 2007, Revision of the species of Hydraena Kugelann (Coleoptera: Hydraenidae) from Palawan and Busuanga, with descriptions of eleven new species, and redescription of Hydraena (Hydraenopsis) scabra d'Orchymont, 1925, pp. 1-44 in Zootaxa 1431 on pages 12-15, DOI: 10.5281/zenodo.17579
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